Contributions to the Knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China : Part 22. Genus Quedius STEPHENS, 1829. Subgenus Microsaurus DEJEAN, 1833. Section 12

15 Pages·2002·15.2 MB·English

by  Smetana, A

Eh・t・,a,To(k、o,30(1):137-151,June30,2002
ContributionstotheKnowledgeoftheQuediina
(Coleoptera,Staphylinidae,Staphylinini)ofChina
Part 22. GenusQuediusSTEPHENs,1829.
SubgenusMicrosaurusDEJEAN,1833.Section12
AlesSMETANA
AgricultureandAgri-FoodCanada,ResearchBranch,
CentralExperimentalFarm,K.W.NeatbyBldg.,
Ottawa,OntarioKIAOC6,Canada
Abstract Taxonomicandfaunisticdataofthespeciesof thegenusQuedius,sub-
genusMic,・osaurus,fromthePeople'sRepublicofChinaareprovided.Fourspeciesarede-
scribedasnew: g. ,-aan(Shaanxi),Q tz、_ (Shaanxi),Q h1,lenn(Shaanxi),andQ. ''eng
(Hubej).Theentiretergite10ofthefemalegenitalsegmentofQgiley1isillustratedfo「
thefirsttime.CommentsconcerningthetaxonomyofQantennalisarcpresented.QuediuS
a11tenna/isisrecordedforthefirst time fromthePeople'sRepublicofChina,basedon
specimenstakeninShaanxi.QuediusinquietusisrecordedforthefirsttimefromShaanxi,
Qbeesotu fromHubei,QdeciusfromHubei,Q.1/olzsc/1u/ufromShaanxi,andQp'ze-
、、,alski1fromSichuan.
Thisisthetwenty-secondofaseriesofpapersdealingwiththeQuediinaofthe
people'sRepublicofChina.Itpresentsthedescriptionsoffurthernewspeciesofthe
subgenusMjc1-osaurusDEJEAN,1833,mostofthemcollectedasrecentlyaslastSum-
mer,duringajointfieldtripoftheauthor,MichaelSCHOLKEandDavidWRASE(both
fromBerljn)toshaanxiandHubei,withmostofthefieldworkdoneinQinlin9Shan
andDabashan(seealso“Contributions”21and23thatarepublishedintheSame
issueoftheElytra).AdditionaldistributionalandtaxonomicinformationiSP「eSented
fornumerousspeciesofthesubgenusMicrosaurus
ued加s(MI'crosaurus)加 Metus(CHAMPION)
Ve11eiusinquietusCHAMPION,1925,107.
Quedius1,lquletus:SMETANA,1997C,l29.
Newreco,・ds. [shaanxi]:Qinlingshan, 12kmSWXunyangba,1900-2250m,
14_18 62000, c Holzschuh,5(5,5 intheNaturhistorischesMuseum,Wien,Aus-
triaandintheSMETANAcollection,Ottawa,Canada.
This isthefirst recordofthisspecies fromShaanxi. ItWasP「eVi-
comments.
138 AlesSMETANA
ouslyknownfromSichuanandYunnan(SMETANA,2001b,194)
Quedius(Microsaurus)beesonlCAMERON
Quedlttsbee.soniCAMERON,1932,285.
New records. [Shaanxi]:borderShaanxi/Sichuan,DabaShan,pass20kmSSE
Zhenping,1700-1800m, 31°44'N109°35'E, 9.VII 2001,A.Smetana [C96b], Ie.
[Hubei]:DabaShan,mtn rangeNEMuyuping,creekvalley4kmNMuyuping,1,700
m, 21.VII 2001, A. Smetana [CII6], I 1 ;DabaShan,passEofMt. DaShen-
nongjia,12kmNWMuyuping,3°130'Ni,l°e21'E,1950m,A.Smetana[CII7],1e,
allspecimensintheSMETANAcollection,Ottawa.
Comments. ThespecimenwithcodeC96bwastakenbysiftingapileofdecay-
ingweedsattheedgeofacornfield;thespecimenswithcodeCII6weretakenbys量l一
ingdebrisunder thebranchesofanfreshly felledtree;thespecimencodedCII7was
takenbysiftingdeepleaflitterunderbroadleavedevergreentrees.
Thisisthe firstrecordofthisspecies fromHubei. Itwaspreviously known from
Fujian,Guangxi,ShaanxiandSichuan(SMETANA,2001b, l94).
Quedius(Microsaurus)antennalisCAMERoN
QuediusantennalisCAMERoM,1932,285.
Quediusantennalis:SMETANA,1988,201.
No、,v reco1・ds. [Shaanxi]: DabaShan SE pass 20km NW Zhenping 1680m
31°59'N109°22'E11.VII2001A.Smetana[C100],1e, intheSMETANAcollection,
Ottawa;samedataasabovebut leg.M.Schiilke[C01-10A]. IntheScHOLKEcollec-
tion,Berlin.
Comments. Thisisthefirst recordofthisspecies frommainlandChina. 0nly
onespecimen(holotype)ofthisspecies,takenintheDaljeelingareaintheHimalaya,
waspreviouslyknown(SMETANA,1988,202).
QuedlusantennalisisthethirdChinesespeciesofthebeeson1group(SMETANA,
2001b,207).It isverydistinctiveanditcannotbeconfusedwithanyotherspeciesof
thegroup,occurringinmainlandChina.Inadditiontothequitedistinctiveaedoeagus
(Seefigs.17-20inSMETANA,1988,394-395),itdiffersfromQaccoSMETANA,1996a
bythelargerandrobusterbody formwithmorevoluminous,widerpronotum,andby
themarkedlycoarserandmoreasperatepunctationoftheelytrawiththeinterspaces
lackingappreciablemicrosculpture. It differs fromQ beeson1, whichoccurs in the
samearea, by themarkedly largerand robusterbody formwithmore voluminous,
widerpronotumwithlateralportionsmoreexplanate, thelongerantennaewhichbe_
Comeslightlypalertowardapex(theyareentirelyblackinQbeeson1),bytheposterior
Punctureofthesublateral rowsonthepronotumsituatedbeforethelevelofthelarge
lateralpuncture(inQbeesonithispunctureissituatedbehindtheleve1ofthelargelat_
era1puncture)andbytheevenlypunctatefirstvisibleabdominaltergite(inQ beeson1
ContributionstotheQuediinaofChina22 139
thetergitebearsasmallimpunctateareainthemiddle).
Intheholotypeofthisspecies,thelargelateralpunctureonthepronotumissitu-
atedquiteclosetothelateralmargin(seeSMETANA,1988,201 fordetails)andIhave
usedthischaracter toseparateQ antennalis fromtheonlyknown femalespecimen
(holotype)ofQ blrmanusCAMERON,1932(SMETANA,1988,186,187). InthetwoChi-
nesespecimensofQantennalisthelargelateralpunctureonthepronotumissituated
away fromthelateralmargin(asusual),makingthepositionofthispunctureuseless
fordistinguishingthese twospecies. Thissupportsmysuggestion (SMETANA,1988,
203)thatQ blrmanusmaybeconspecificwithQantennalis.
Quedius(Microsaurus)deciusSMETANA
QuediusdeciusSMETANA,l996a,12.
New records. [Hubei]:DabaShan,passEofMt.DaShennongjia,12kmNW
Muyuping3°130'Nile°21'E,2050m,19.VII2001,A.Smetana[CII2],15(,inthe
SMETANAcollection, Ottawa. [Sichuan]: GonggaShan, 29°41'N 101°58'E, 2800m,
14.-19.VI.1999,Siniaev&Plutenko,1 intheScHOLKEcollection.
,
comments. ThespecimencodedCII2wastakenbysifting fairly freshmush-
roomsgrowingonafallentree.
ThisisthefirstrecordofthisspeciesfromHubei.
Quedius(Microsaurus)chromesSMETANA
QuediuschromesSMETANA,1996a le.
[Shaanxi]:DabaShan,mtn rangeNpass22kmNWZhenPin9,
New records.
32°01'N10g°21'E,2850m,14.VII 2001,A.Smetana[CIO3],2(5,1e,intheSMETANA
co11ectjon; Qjnljng shan,mtn rangeWpassonroadXi'an-Shagoujie,45kmSSW
xj'an,33°52'N108°46'E,2675m,A.Smetana[CII9],1(3,2?,M.SChiilke[C01-20],
2 3 intheScHULKEandSMETANACollections.
, The, specjmenscodedCIO3weretaken,togetherwithspecmi ensofQhuenn,ina
remnantofanoriginalAblesforestbysiftingvariousdebris,rottingWoodandfallen
mouldybarkaroundbasesofdead,standingorfallenAblestrees;thoseCodedCII9
and cOl_20 were sjfted from floor litter andvegetationunderAbles, Betula and
Rhododendrontrees
“ed加s(MierosaMrus)adJacensCAMERON
QuediusadJacensCAMERON,1926,368.
chjna: [Shaanxi]:Qinlingshan,12kmSWXunyan9ba,1900-
New ,ecol・ds
intheNaturhistoriSCheSMuseum,Wien,
2250m, 14_186 2000, C Holzschuh, 1
Austrja; Qjnlingshan, 6km E Xunyangb, a, 1000-1300m, 235・一13・6・2000, C・
.
jn theNaturhistorisches Museum, Wien,Aust「Iaand inthe
Ho1zschuh, 4 6
, ,
l40 AlesSMETANA
SMETANAcOllection,Ottawa,Canada.
Comments. OnlyonerecordofthisspeciesfromShaanxiwaspreviouslyknown
(SMETANA,1999,214).
Quedius(Microsaurus)holzschuhiSMETANA
QuediusholzschuhiSMETANA,1999,220.
Net,、ノ record. [Shaanxi]: Qinlingshan, 6km E Xunyangba, 1000-1300m,
23.5.-13.6.2000,C.Holzschuh,1 intheNaturhistorischesMuseum,Wien,Austria.
Comments. Thisspecimenismarkedlysmaller thanthetwomalespecimensof
theoriginalseries(7.4mm),butitagreesinallcharacters,includingtheshapeofthe
aedoeagusandofthestemite9ofthemalegenitalsegment(figs.12-16inSMETANA,
1999a,221)withtheholotype.Thisisthefirst recordofthisspeciesfromShaanxi.It
waspreviouslyknownonlyfromcruelShan,Sichuan(SMETANA,2001b,199).
Quedius(Microsaurus)schuelkeiSMETANA
QuediusschuelkeiSMETANA,1997b,455.
Nevi, records. [Shaanxi]: Qinlingshan, 6km E Xunyangba, 1000-1300m,
23.5.-13.6.2000,C.Holzschuh,6 8 intheNaturhistorischesMuseum,Wien,
, ,
AustriaandintheSMETANAcollection,Ottawa,Canada.
Comments. Thespeciesisatpresentknownonly fromQinlingshan,butsinceit
Seemstooccurmainlyat lowermountainelevations, itmaybemorewidelydistrib_
uted.
Quedius(Microsaurus)germattorumSMETANA
QuediuSgermanorumSMETANA,1997b,457.
No、,l' record. [Shaanxi]:QinlingShan,passonroadZhouzhi_Foping,105km
SWXi'an,Nslope,33°44'N107°59'E,1990m,4.VII2001,A.Smetana[cg3],1e;
M・SChiilke[COI-01],1e,19,intheSMETANAandScHULKEcollections.
TheSpecmi ensweretakenbysiftingvariousdebrisanddeadwoodalongasmall
creekinamixeddeciduousforest.
Quedius(Microsaurus)przewalskiiREITTER
QuediusprzewalskiiREITTER,1887,211.
New 「ecord. [Sichuan]:northernSichuan,Zoige,13.VI.1996,Benes&stepaj
Ie9.,1 intheSMETANAcOllection.
,
Comment ThisisthefirstrecordofthisspeciesfromSichuan.Itwaspreviously
:
knownf「omQmghai,TibetandYunnan(SMETANA,2001b,206).
ContributionstotheQuediinaofChina,22 141
Quedius(Microsaurus)duhSMETANA
QuediusduhSMETANA,2001,186.
New records. [Shaanxi]: Qinling Shan, mt range W pass on road Xi'an-
Shagoujie,45kmSSWXi'an,33°52'N108°46'E,2600m,25.VII2001,A.Smetana
[CII8],1e,59;samebut2675m,26.VII 2001,A.Smetana[CII9], 1e,5e;same
but2675m,25.VII 2001,M.Schiilke[C01-20],2e,2e;samebut25.-26.VII2001,
pitfalltraps,M.Schiilke[C01-20B],39,3e;samebut2675m,25.VII2001,Wrase
[20],1e;allspecimensintheSCHULKEandSMETANAcollections.
Comments. Theabovespecimenscome fromthesameareaasthetwospeci-
mensoftheoriginalseriesofthisspecies.Thehabitatisanareaofsubalpinemeadows
withmoreor lesslargepatchesofAbles,Betula,RhododendronandsomeLarixtrees
withshrubbyundergrowth.Quediusduhisfairly frequentthere,mostspecimenswere
takenbys量l ingvarious forest floor debris, rottingpiecesofwoodandbark around
basesoftrees,andbysiftingvariousforestfloorvegetation.Itoccurstogetherwith
.
chromesinthehabitatundertheRhododendrontreesonthenorthslope.
Quedius(Microsaurus)zheduoSMETANA
QuediuszheduoSMETANA,1999,235.
New record. [Sichuan]:DaxueShan,TshetoLaPass,30°05'N101°48'E,4300-
4350m(alpinemeadows),25.VII i999,D.W.Wrase,1 in theSCHuLKEcollection.
,
Comment. Judgingfromthecoordinates,thiscollectingsiteissituatedcloseto
thetypelocality(seeSMETANA,1999,236)andthename“TshetoLa”mustbeanOthe「
name forthesamepass.
Quedius(Microsaurus)euanderSMETANA
9uediuseuande,SMETANA,1997a,63.
New record. [Sichuan]: LudingCo.,ErlangShanPass,road318,cca3000m,
8kmSELuding,190kmSWChengdu,21.-29.VI.1999,D.W.Wrase,13,,2?,inthe
SCHULKEcOllection.
Thisadditionalrecord fromErlangShan(seeSMETANA,2001a, l84)
comment.
confjrmsthatQeuanderisfairlycommoninthismountainran9e.
Quedius(Microsaurus)gueySMETANA
(Fig.l)
QuediusgueySMETANA,2001a,188.
Newrecords. [shaanxi]:QinlingShan,Autoroute93kmSZhouzhi,108kmSW
xj'an,107°56'E33°45'N,1650m,mountainforest,1.-2.IX.1995,leg.A.PiitZ,1j in
thepoTzcollection;QinlingShan,passonroadZhouzhi-Fopin9,105kmSWX1'an,
142 AlesSMETANA
N-slope, 107°59'E 33°44'N, 1990m, 2.VII 2001, A. Smetana [C89], 4e, in the
SMETANAcollection;samebut4.VII2001and[C93],2!,intheSMETANAcollection.
Comments. ThespecimencollectedbyPuTzwasactuallytakenat thetypelo-
cality(seeSMETANA,2001a,190);theremainingspecimens[C89,C93]comefromthe
sameareaandthesame forest,thecollectingsiteliesjusthigherup,abovethepass.
Theywerecollectedinamixeddeciduousforestbysl価ngvariousforest floordebris
alongasmallcreek.
Basedontheshapeofthetergite10ofthefemalegenitalsegment,aswellason
thefact thatbothcollectingsites [C89andC93] lieclosetothetypelocalityofthe
species,thereislittledoubtthatthespecimensbelongtoQguey.
Sincethetergite10ofthefemaleallotypeofthisspecieswasbadlydamagedwith
onlythecharacteristicapicalportionpresent, theentiretergite10is加ustratedhere
(Fig. 1).
Quedius(Microsaurus)raa,tspnov.
(Figs2-8)
Description. Piceouswithpiceous-blackhead,apicalmarginofelytranarrowly
paler, apical margins ofabdominal tergites andapex of abdomen paler, abdomen
slightly iridescent;maxillaryandlabialpalpi testaceobrunneoustobrunneous, legs
brunneouswithpaler tarsi,medial facesofmiddleandhind femoradarkened.Head
relativelynarrow,ofroundedquadrangularshape,aboutaslongaswide(butappear_
ing,duetoitsconfigurationbehindeyes,slightlylongerthanwide),lateralmarginsbe_
hindeyesevenly,graduallynarrowedtowardneck,headthereforeentirelylackjngeven
tracesofposteriorangles;eyesmoderatelylargeandconvex,temperaaboutaslongas
eyes Seen from above; noadditionalsetiferouspunctures between anterjor frontal
Punctures;Posteriorfrontalpuncturesituatedawayfromposteromedialmarginofeye,
butmarkedlycloser toit thantoposteriormarginofhead,oneadditjona1setjferous
Puncturepresentatposteriormarginofeyeandnumerousadditionalpuncturespresent
ante「ladandposteriadofposteriorfrontalpuncturebetweentwousualsetiferouspunc_
tu「eSatPosteriormarginofhead;temporalpuncturesituatedaway fromposteriormar_
9inofeye,markedlyclosertoposteriormarginofheadthantoposteriormargjnofeye;
tempo「awithnumerous finepunctures;surfaceofheadwith very fine, densemi_
C「oSCulPtu「eoftransversewaves.Antennamoderatelylong,slightlywidenedtoward
apex,Se9ment3somewhat longer thansegment2(ratio125),segments4_6longer
thanwide,graduallybecomingshorter,segments7-10aboutaslongaswjde,lastseg_
Fi9S・ 1-9・ - 1.Quediusguey:tergite10offemalegenitalsegment. - Fjgs.2_8 Quedjuslaan:2,
apicalPO「tiOnofmalesternite8;3,tergite10ofmalegenitalsegment;4;sternjte9ofmalegenjla1
Se9ment;5,aedOeagus,ventralview;6,apicalportionofmedianlobe,paramereremoved;7,apjca1
PO「tOinofundersideofparamere;8,tergite10offemalegenitalsegment. _
g.Quedlustzwu:api_
calportionofmalesternite8.
ContributionstotheQuediinaofChina,22 143
l44 AlesSMETANA
montaboutaslongastwoprecedingsegmentscombined.Pronotumaboutaslongas
wide,widestataboutposteriorthir markedlynarrowedanteria withlateralmargins
continuouslyarcuatewithbroadlyroundedbase, transverselyconvex,lateralportions
notexplanate;dorsalrowseachwiththreepunctures;sublatera1rowseachwiththree
or four punctures,posteriorpuncturesituatedbehindlevelof largelateral puncture;
oneadditionalpuncturebetweendorsalandsublatera1rows;surfaceofpronotumwith
microsculpturesimilar tothatonhea butmarkedly fineranddenser.Scutellumim-
punctate,withveryfineanddensemicrosculptureoftransversestriae.Elytrarelatively
long,atbasenarrowerthanpronotumatwidestpoint,slightlywidenedposteria atsu-
tureaboutaslongas,atsidessomewhatlongerthanpronotumatmidline(ratio1.15);
punctationmoderately fine, dense, transverseinterspacesbetweenpuncturesmostly
aboutaswideasdiametersofpunctures;pubescencebrownish;surfacebetweenpunc-
tures withoutmicrosculpture.Wings fully developed.Abdomenwith tergite7(fifth
visible)withfinewhitishapicalseamofpalisadefringe;punctationandbrownishpu-
bescenceofabdominaltergitesfineandmoderatelydense,almostevenlycoveringeach
tergite,ingeneralbecomingindistinctlysparsertowardapexofabdomen;surfacebe-
tweenpunctureswithexceedinglydenseandfinemicrosculptureoftransversestriae.
Male. First foursegmentsof front tarsusmarkedlydilate sub-bi1obe each
denselycoveredwithmodifiedpalesetaeventrally;segment2aboutaswideasapexof
tibia;segment4narrowerthanprecedingsegments.Sternite8withfivelongsetaeon
eachside,withmoderatelywideanddeep,arcuatemedioapica1emargination,small
triangular areabeforeemargination attenedandsmooth (Fig 2). Genital segment
withtergite10triangular,evenlynarrowedtowardnarrowlyarcuateapex,withnumer-
ous longsetaeatandnearapical margin, andwithseveralshortersetaein front of
them(Fig3);sternite9withrathershortbasalportion,apicalportionwidenedbasally
andthennarrowedintosubtruncate,mediallyemarginateapex,withtwoindistinctly
differentiatedsubapicalsetaeoneachsidebeforeemargination(Fig 4). Aedoeagus
(Figs 5-7)narrowandelongate;median lobesubpara11e1-side anteriorly narrowed
intonarrowlyarcuateapex,withmarkedlydeve1ope longmedialcarinaonfaceadja_
centtoparamere.Paramereelongate,narrow,withnarrowlyarcuateapexdistinctlynot
reachin9apexofmedianlobe; fourfinesetaeatapex,twoslightlyfinersetaeateach
lateral marginbelow apex; undersideofparamerewith threesensory pegsetae far
belowapex.Internalsacwithoutlargersclerotizedstructures.
Female. Firstfoursegmentsoffronttarsussimilartothoseofmale,butlessdi_
lated;segment2slightlynarrowerthanapexoftibia(ratio0.87).Genitalsegmentwjth
tergite10narrowlytriangular,withslightlydifferentiated,subacuteapicalportjon,wjth
SeVe「allen9setaeatandnearapicalmargin,andwithseveralmarkedlyshortersetaein
frontofthem(Fig8).
Length7.8-8.00mm.
Typematerial. Holotype(male)andallotype(female):China:“China:Shaanxj,
QinLingShan110°06'E,34°25'NHuaShanMt.,S.-top,1950-2000mForest,sifted
19.08.1995,1e9.Piitz”.InthePUTzcollection,Eisenhiittenstadt.Holotypetemporarily
ContributionstotheQucdiinaof(、hina,22 l45
intheSMETANAcOllection,Ottawa.
Paratype:China: [Shaanxi]:samedataasholotype,but leg. M. Schiilke, l !, in
theScHOLKEcollection,Berlin.
Geograp/1!caf fsf1'10zlffo'7. zled1zfs1,aanisatpresentknownonly fromthetype
localityinQinlingShaninsouthernShaanxi.
Bionomics. Thespecimensweres量l edinapresumablymixeddeciduousforest,
butnodetailsareknown.
ecogn加onan comments. lfedlzls1-aan is another member of the ,mi ｽuensl
Itdiffersfromailothermembers,inadditiontothemalesexualcharacters(par-
group.
ticularlytheverydistinctive,longmediancarinaofthemedianlobeonthe faceadja-
cent totheparamere),by thedistinctiveshapeoftheheadcombinedwith itschaeto-
taxyandthepresenceofnumerousadditionalpunctures(seeabove),andbythepres-
enceoffivelongsetaeoneachsideofthemalesternite8.Theshapeoftheheadof
.
1・aanresemblesthatoftheNearcticspeciesQ.pete'g1'inus(GRAvENHoRsT,1806).
Thefemaleparatypeoftheoriginalseriesismarkedlyteneral.
Etymology. ThespecificepithetistheChineseword“raan”(meaning“gradu-
ally”).Itreferstotheshapeofthehea thatnarrowsgraduallytowardtheneckbehind
theeyes.
Mefl加s(M'crosaM'MS)tZ'- Sp nov.
(Figs 9-15)
Desc1-1pt1o11. Inallcharacterssimilar toQep、'tusSMETANA,1995anddifferent
mainlybybothmaleandfemalesexualcharacters.0naveragesmallerandlessrobust、
antennaeandlegsslenderer,elytrashorter,atsutureslightlyshorter (ratio0.87),at
sjdesaboutaslongaspronotumatmidline(correspondingratios forQepytus:1.07
and1.15).
First foursegmentsof fronttarsusmarkedlydilated,sub-bilobed,each
Male.
withmodifiedpalesetaeventrally;segment2aboutaswideasapexoftibia,se9ment4
narrowerthanprecedingsegments.Sternite8withthreelongsetaeoneachside;With
moderatelywideandshallow,arcuatemedioapica1emargination,smalltrian9ula「a「ea
beforeemarginationflattenedandsmooth(Fig 9).Genitalsegmentwithbothto「9ite
10andsternite9similartothoseofQep、・ttls(Figs.10,11).AedoeaguS(Fi9S.12-14)
narrowandelongate,similar tothatofQepytus;medianlobenarrower,pa「tiCula「ly
anteriorly,paramerenarrower,subparalle1-side notquitecoveringmedianlobe,With
vaguelyemarginateapex,apexappreciablynotreachingapexofmedianlobe;With
fourfjnesetaeatapex,medialpairlongerthanlateralsetae,andwithtwominuteSetae
ateachlateralmargjnclosetoapex; undersideofparamerewiththreeSense「yPe9
setaenearapicalmargin.InternalsacwithoutlargersclerotizedSt「uCtu「oS.
Fjrstfoursegmentsoffronttarsussimilartothoseofmale,butlessdi-
Female
segment2slightlynarrowerthanapexoftibia(ratio0.90).GenitalSe9mentWith
late
tergjte10sjmjlartothatofQep)・ttls,but ingeneralshorter,andwithditto「entiated
146 AlesSMETANA
apicalportionmarkedlyshorter(Fig.15).
Length6.7-6.9mm.
Typematerial. Holotype(male) andallotype(female):China: “CHINA: S-
Shaanxi(DabaShan)mountainrangeNpass22kmNWZhenping,32°01'N109°21'E
2850m, l4.VII 2001,legM.Schiilke[COI-12].IntheScHuLKEcollection,Berlin.
Paratype: [Shaanxi]:samedataasholotype,1e. IntheSMETANAcollection,Ot-
tawa.
Geographicaldistribution. Quediustzwuisatpresentknownonlyfromthetype
localityinDabaShaninsouthernShaanxi.
Bzonomlcs. Thespecimenswere collected in anoriginal Ables forest on the
northernslopenear thetop of themountain; they bear the followinghabitat data:
“Ables,bushes,deadwood(sifted)”.
ecogmtzonandcomments. uedzus fzwuisanothermember of themti eKszs
group(seeSMETANA,2001b,201).TheaedoeagusofQ tzwuismostsimilar tothose
ofQ epytusandQanton1.ItdiffersfromthatofQ anton1bytheparamereoftheae-
doeagusreachingmarkedlycloser totheapexofmedianlobeandbythelowernumber
ofsensorypegsetaeontheparamere(figs.12,14,andfigs.10,13inSMETANA,1995,
235).TheaedoeagusofQepytusdiffersbythewiderparamere,whichisslightly,ar-
cuatelydilatedanteriorlyandcoverstheentireanteriorportionofthemedianlobe,ex-
cept for theverytip,andbythelargernumber ofsensorypegsetaeontheparamere
(figs.17,19inSMETANA,1995,235,237).
Thereisanextrapunctureintheleftdorsal rowonthepronotuminthemale
paratype.
Etymology. ThespecificepithetistheChineseword“tzwu”,whichinoneofits
meaningsmeans“relativesinagroup”.Itreferstotheapparentcloserelationshipof
.
tzwutotheabovetwomembersofthemukuenslsgroup.
Quedius(Microsaurus)huennspnov.
(Figs.16-21)
Description. Inallexternalcharacterssimilar toQ. liauSMETANA,1999a,but
different as follows: somewhat larger andmore robust, coloration darker: piceous
black,antennapiceouswithfirstthreesegmentspaler,legspalebrunneouswithinner
facesofmiddleandhindtibiaedarkened;palercolorofpronota1marginsmuchless
appa「entandmostlyappreciableonlyatbasal margin;headwider andlessnarrowed
Fi9S.10-22. - 10-15.Quediustzwu:10,tergitel0ofmalegenitalsegment;11,sternite9ofmalegen_
italSo9ment.12,aedoeagus,ventralveiw;13,apicalportionofmedianlobe,paramereremoved; l4,
apical Portion ofundersideofparamere; 15, tergite 10 of female genital segment. _ 16_21
Quediushuenrl:16,apicalportionofmalestemite8;17,tergite10ofmalegenitalsegment;18,ster_
nife9ofmalegenitalsegment;l9,aedoeagus,ventralview;20,apicalportionofundersjdeofpara_
me「e;21, to「9ite10of femalegenitalsegment. - 22.Quedius,,eng:tergite10offemalegenjtal
segment.