Checklist of the species of Cyclominae (Coleoptera: Curculionidae) occurring in America south of the United States

14 Pages·2002·0.15 MB·

by  Morrone, Juan José

ISSN 0373-5680 Rev. Soc. Entomol. Argent. 61 (1-2): 11-24, 2002 11
Checklist of the species of Cyclominae
(Coleoptera: Curculionidae)
occurring in America south of the United States
MORRONE, Juan J.
Museo de Zoologia, Departamento de Biologia Evolutiva, Facultad de Ciencias,
Universidad Nacional Autonoma de Mexico (UNAM), Apdo. Postal 70-399, 04510 Mexico D.K, Mexico.
E-mail: [email protected]
• ABSTRACT. The valid names of the genera and species of cyclomines (Co-
leoptera: Curculionidae: Cyclominae) found in America south of the United
States are listed, including their synonyms and distributional summaries.
These taxa belong in the tribes Aterpini (genera Aegorhinus Erichson and
Alastoropolus Kuschel), Gonipterini (genus Gonipterus Schonherr), and
Rhythirrinini (genera Acroriellus Morrone & Ocampo, Acrorius Kirsch,
Acrostomus Kuschel, Adioristidius Morrone, Amathynetoides Morrone, Ande-
sianellus Anderson & Morrone, Antarctobius Fairmaire, Falklandiellus Kuschel,
Falklandiopsis Morrone & Anderson, Falklandius Enderlein, Germainiellus
Morrone, Haversiella Schweiger, Hyperoides Marshall, Lamiarhinus Morrone,
Lanteriella Morrone, Listroderes Schönherr, Listronotus Jekel, Macrostyphlus
Kirsch, Nacodius Morrone, Neopachytychius Hustache, Philippius Germain,
Puranius Germain, Rhigopsidius Heller, Rupanius Morrone, Telurus Kuschel,
and Trachodema Blanchard).
KEYWORDS. Curculionidae. Cyclominae. Aterpini. Gonipterini. Rhythirrinini.
America.
• RESUMEN. Lista de las especies de Cyclominae (Coleoptera: Curculioni-
dae) que se encuentran en América al sur de los Estados Unidos. Se listan
los nombres válidos de los géneros y especies de ciclominos (Coleoptera:
Curculionidae: Cyclominae) distribuidos en America al sur de los Estados
Unidos, incluyendo sus sinonimos y sumarios distribucionales. Estos taxones
pertenecen a las tribus Aterpini (géneros Aegorhinus Erichson y Alastoropolus
Kuschel), Gonipterini (genero Gonipterus Schönherr) y Rhythirrinini (géneros
Acroriellus Morrone & Ocampo, Acrorius Kirsch, Acrostomus Kuschel,
Adioristidius Morrone, Amathynetoides Morrone, Andesianellus Anderson
& Morrone, Antarctobius Fairmaire, Falklandiellus Kuschel, Falklandiopsis
Morrone & Anderson, Falklandius Enderlein, Germainiellus Morrone, Ha-
versiella Schweiger, Hyperoides Marshall, Lamiarhinus Morrone, Lanteriella
Morrone, Listroderes Schönherr, Listronotus Jekel, Macrostyphlus Kirsch,
Nacodius Morrone, Neopachytychius Hustache, Philippius Germain, Pura-
nius Germain, Rhigopsidius Heller, Rupanius Morrone, Telurus Kuschel y
Trachodema Blanchard).
PALABRAS CLAVE. Curculionidae. Cyclominae. Aterpini. Gonipterini.
Rhythirrinini. América.
12 Rev. Soc. Entomol. Argent. 61 (1-2), 2002
INTRODUCTION RESULTS
The subfamily Cyclominae (Coleoptera: Cur- Subfamily CYCLOMINAE Schonherr, 1826
culionidae) comprises several weevil taxa that Tribe ATERPINI Lacordaire, 1863
were assigned formerly by Kuschel (1995) to the Subtribe ATERPINA Lacordaire, 1863
subfamily Brachycerinae (Morrone, 1997a, b,
1998, 2000; Alonso-Zarazaga & Lyal, 1999). A Aegorhinus Erichson, 1834: 261 [type species:
cladistic analysis based on larval features (Mar- A. phaleratus Erichson, 1834, by monotypy]
valdi, 1997) has shown that they are closely re- [= Psuchocephalus Latreille, 1828: 597; Lopho-
lated to the subfamily Entiminae, although their tus Schonherr, 1834: 314 (non Giorna, 1809,
monophyly is doubtful (R. Oberprieler, pers. nec Fischer von Waldheim, 1813); Leucolopus
com.). They are classified into nine tribes: Dejean, 1835: 261; Lophodes Dejean, 1836:
Amycterini, Aterpini, Cyclomini, Diabathrariini, 284; Eublepharus Gay & Solier, 1839: 11; Ce-
Dichotrachelini, Gonipterini, Rhythirrinini, So- ropsis Gay & Solier, 1839: 19; Leucolophus
matodini, and Viticiini (Alonso-Zarazaga & Lyal, Chevrolat, 1846: 322; Orodinus Gistel, 1848:
1999). Three of these tribes (Aterpini, Gonipte- IX; Oegorinus Montrouzier, 1855: 48; Psupho-
rini, and Rhythirrinini) are distributed in the cephalus Imhoff, 1856: 221; Eublephorus Che-
New World. nu, 1860: 225; Lophodus Gemminger & Harold,
O'Brien & Wibmer (1982) and Wibmer & 1871: 2392 (non Symons, 1861, nec Roma-
O'Brien (1986) compiled the most recent chec- nowsky, 1864, nec Newberry & Worthen,
klists of the New World weevils. Over the course 1870); Sublepharus Desbrochers des Loges,
of the last years, however, several new cyclomine 1910: 130; Euplepharus Bruch, 1915:416; Ae-
taxa have been described, new combinations and gorrhinus Bruch, 1915: 416; Micropolus Kus-
synonymies were established subsequent to these chel, 1952a: 243]; Morrone & Roig-Juñent,
checklists, and new data have been added 2000: 421 (revision)].
(O'Brien & Wibmer, 1984; Wibmer & O'Brien, Alonso-Zarazaga & Lyal (1999) recently disco-
1989; Morrone, 1990, 1992a-c, 1993a-d, 1994a- vered that Psuchocephalus is a senior synonym of
d, 1995a-e, 1997b; Morrone & Loiacono, 1992; Aegorhinus. Elgueta & Kuschel (in press), however,
Morrone & Anderson, 1995; Morrone & Ocampo, have recently submitted a proposal to the Interna-
1995; Morrone et al., 1995; Anderson & Morro- tional Commision on Zoological Nomenclature,
ne, 1996; Ocampo & Morrone, 1996; Alonso-Za- proposing that Psuchocephalus has precedence
razaga & Lyal, 1999; Elgueta, 2000; Morrone & over Aegorhinus. While the case is under conside-
Roig-Juñent, 2000). ration by the Commision, I maintain the current
name Aegorhinus.
My objective is to provide a list of the Cyclo- A. albolineatus (Blanchard, 1851: 332). Chile.
minae occurring in America south of the United A. bilineatus Elgueta, 2000: 13. Chile.
States, incorporating changes and additions that A. boviei (Desbrochers des Loges, 1910: 127) [= A.
update O'Brien & Wibmer's (1982) and Wibmer incertus (Germain, 1911: 217)]. Chile.
& O'Brien's (1986) checklists. Basically I follow A. bulbifer (Kuschel, 1952a: 229) [= A. bulbifer
the same format of my previous checklist of the (Germain, 1911: 217)]. Argentina and Chile.
species of Entiminae (Morrone, 1999). Classifi- A. delfini (Germain, 1893: 1031) [= A. boniarde
cation of the taxa into tribes and subtribes fo- Cekalovic, 1970: 56]. Chile.
llows Alonso-Zarazaga & Lyal (1999). The A. fascicularis Kuschel, 1952a: 243. Chile.
arrangement of the genera and species within A. inermis Kuschel, 1952a: 236 [= A. inermis
each tribe and subtribe is alphabetical. Each ge- (Germain, 1911: 217)]. Chile.
neric name is followed by the original citation, A. kuscheli Elgueta, 1974: 133. Chile.
with its type species and synonyms listed chro- A. maestus Kuschel, 1952a: 232. Chile.
nologically between square brackets. Distribu- A. maulinus Elgueta, 2000: 8. Chile.
tional data for each species are summarized by A. nitens Kuschel, 1952a: 229. Chile.
country; an asterisk at the left side of the species A. nodipennis (Hope, 1834:15) [= A. nodinipennis
name denotes that it is present also in America (Solier, 1839: L); A. rouleti (Gay & Solier, 1839:
north of Mexico. 17); A. subrugosus (Philippi, 1859: 665)]. Ar-
MORRONE, J. J. Checklist of the species of Cyclominae 13
gentina and Chile. Goniopterus Gemminger & Harold, 1871:
A. ochreolus Kuschel, 1952a: 223. Chile. 2377(non Agassiz, 1846); Dacnirotatus MarelIi,
A. oculatus Kuschel, 1952a: 235 [= A. philippii 1926:630]
(Germain 1911: 217)]. Argentina and Chile. Although Zimmerman (1994) established the
A. opaculus (Desbrochers des Loges, 1910: 127) synonymy of G. gibberus and G. scutellatus, there
[= A. abbreviates (Germain 1911:217)]. Chile. are two distinct species of Gonipterus in South
A. phaleratus Erichson, 1834: 262 [= A. distin- America (pers. obs.). Until the problem concerning
guendus (Germain, 1854: 331)]. Chile. their valid names is elucidated, I keep Wibmer &
A. schoenherri (Gay & Solier, 1839: 19) [= A. bre- O'Brien's (1986) names.
virostris (Germain, 1911: 217)]. Chile. G. gibberus Boisduval, 1835: 324 [= G. bruchi
A. servillei (Gay & Solier, 1839: 15) [= A. quadri- (Marelli, 1926: 630)]. Argentina, Brazil, and
dentatus (Philippi 1859: 665)]. Chile. Uruguay. Introduced from Australia and Tas-
A. silvicola Kuschel, 1952a: 230 [= A. izquierdoi mania.
(Germain, 1911: 217), A. silvicola cistrans G. scutellatus Gyllenhal, 1833: 458 [= G. platensis
Kuschel, 1952a: 230]. Argentina and Chile. (Marelli, 1926: 640), G. marellii (Uyttenboo-
A. subplanifrons Kuschel, 1952a: 235 [= A. gaart, 1928: 77)]. Argentina, Brazil, and Uru-
schoenherri subplanifrons Kuschel, 1952a: guay. Introduced from Australia and Tasmania.
235]. Chile.
A. superciliosus (Guerin, 1830: 112) [= A. eschs-
choltzi (Boheman, 1834: 316), A germari Tribe RHYTHIRRININI Lacordaire, 1863
(Gay & Solier, 1839: 21), A. fasciatus (Bohe- Subtribe GRONOPINA Bedel, 1884
man, 1842: 131), A. reticulatus (Gyllenhal,
1842: 132), A. perplexus (Desbrochers des Lo- Rhigopsidius Heller, 1906: 7 [type species: R. tu-
ges, 1910: 128), A. vermicuIaris (Desbrochers cumanus Heller, 1906, by original designa-
des Loges, 1910: 129), A. araucanus (Germain, tion]; Morrone & Loiacono, 1992: 130 (revi-
1911: 217), A. ornatus (Germain, 1911: 217), sion).
A. superciliosus ornatus Kuschel, 1952a: 239, R. piercei Heller, 1936: 284 [= R. tucumanus
A. superciliosus germari Kuschel, 1952a: 240, Sasscer & Pierce, 1913: 143 (non Heller,
A. superciliosus philippii Kuschel, 1952a: 241, 1906)]. Argentina, Bolivia, and Peru.
A. superciliosus andinus Kuschel, 1952a: 242, R. tucumanus Heller, 1906: 8. Argentina.
A. superciliosus araucanus Kuschel, 1952a:
242, A. superciliosus laqueatus Kuschel, 1952a: Subtribe LISTRODERINA LeConte, 1876
242, A. superciliosus philippianus Kuschel,
1955: 273]. Argentina and Chile.
Acroriellus Morrone & Ocampo, 1995: 257 [type
A. suturalis (Blanchard, 1851: 331). Chile. species: A. viridisquamosus Morrone & Ocam-
A. transandinus Kuschel, 1952a: 231. Argentina. po, 1995, by original designation].
A. vitulus (Fabricius, 1775: 152) [= A. leprosus A. bobi Morrone & Oeampo, 1995: 260. Ecuador.
(Olivier, 1807: 395)]. Argentina and Chile. A. carinatus Morrone & Ocampo, 1995: 260.
Ecuador.
Alastoropolus Kuschel, 1950: 15 [type species: A. similaris Morrone & Ocampo, 1995: 260.
Curculio strumosus Olivier, 1807, by original Ecuador.
designation]; Morrone & Roig-Juñent, 2000: A. tuberculosus Morrone & Ocampo, 1995: 260.
420 (revision). Ecuador.
A. strumosus (Olivier, 1807: 394) [= A. longipes A. viridisquamosus Morrone & Ocampo, 1995:
(Waterhouse, 1840: 330)]. Argentina and Chile. 261. Peru.
A. vittetae Morrone & Ocampo, 1995: 259. Co-
lombia.
Tribe GONIPTER1N1 Lacordaire, 1863
Acrorius Kirsch, 1889: 25 [type species: A.
Gonipterus Schonherr, 1833: 456 [type species: puncticollis Kirsch, 1889 by monotypy] [=
G. lepidotus Gyllenhal, 1833, by original de- Ocromis Sharp, 1890: 152]; Morrone 1994c:
signation] [= Goniopterus Agassiz, 1846: 165; 102 (revision).
14 Rev. Soc. Entomol. Argent. 61 (1-2), 2002
A. andersoni Morrone, 1994c: 109. Ecuador. A. tuberculatus (Voss, 1954: 258). Bolivia and Peru.
A. bolivianus Ocampo & Morrone, 1996: 25. A. variegatus (Voss, 1954: 244). Bolivia.
Bolivia.
A. cuprinus Morrone, 1994c: 109. Ecuador. Amathynetoides Morrone, 1994b: 28 [type spe-
A. nymphalis Morrone, 1994c: 108. Ecuador. cies: Amathynetes appendiculatus Kuschel,
A. otramas Ocampo & Morrone, 1996: 26. 1949, by original designation] [= Amathynetes
Ecuador. Kuschel, 1949: 43 (non Olliff, 1891)].
A. puncticollis Kirsch, 1889: 25. Colombia. A. appendiculatus (Kuschel, 1949: 47). Bolivia,
A. papallacta Morrone, 1994c: 108. Ecuador. Chile, and Peru.
A. pillahuata Morrone, 1994c: 109. Peru. A. ebeninus (Hustache, 1938b: 287). Peru.
A. plicatifrons Morrone, 1994c: 110. Peru. A. intemperatus Morrone, 1994b: 31. Peru.
A. sisyphus Morrone, 1994c: 105. Colombia. A. longulus (Kuschel, 1949: 49). Chile and Peru.
A. morbeamus Morrone, 1994b: 33. Peru.
Acrostomus Kuschel, 1955: 287 [type species: A. nitidiventris (Hustache, 1938b: 285). Bolivia.
Adioristus bruchi Hustache, 1926, by original A. normae Morrone, 1994b: 34. Bolivia.
designation]; Morrone, 1994d: 404 (revision). A. palustris (Kuschel, 1949: 51). Bolivia, Chile,
A. bruchi (Hustache, 1926: 186). Argentina. and Peru.
A. cruralis Kuschel, 1958: 234. Argentina. A. sparsesetosus (Hustache, 1938b: 283). Bolivia
A. foveicollis Kuschel, 1958: 232. Argentina. and Peru.
A. griseus (Guérin, 1839: 305) [= A. pubescens A. sundrianus Morrone, 1994b: 36. Peru.
(Germain, 1895: 481)]. Argentina and Chile.
A. magellanicus Kuschel, 1958: 233. Chile. Andesianellus Anderson & Morrone, 1996: 260
A. mordor Morrone, 1994d: 409. Argentina. [type species: A. microphthalmicus Anderson
A. vianai Kuschel, 1958: 233. Argentina and Chile. & Morrone, 1996, by original designation]
A. carltoni Anderson & Morrone, 1996: 272.
Adioristidius Morrone, 1994b: 13 [type species: Ecuador.
Adioristus similaris Voss, 1954, by original de- A. cotopaxi Anderson & Morrone, 1996: 276.
signation] [= Adioristidius Voss, 1954: 242; Ecuador.
Anchadioristus Voss 1954: 242; Adioristidus A. fulgidus Anderson & Morrone, 1996: 274.
Edwards & Hopwood, 1966: 5]. Ecuador.
A. anchonoideus (Hustache, 1938a: 181). Bolivia. A. hermani Anderson & Morrone, 1996: 269. Co-
A. carinicollis (Voss, 1954: 255). Peru. lombia.
A. chilensis Morrone, 1994b: 18. Chile. A. masneri Anderson & Morrone, 1996: 268.
A. costulatus (Hustache, 1938a: 182). Bolivia, Ecuador.
A. crassirostris (Hustache, 1938a: 180). Bolivia. A. microphthalmicus Anderson & Morrone, 1996:
A. cuprisquameus {Voss, 1954: 254). Peru. 269. Ecuador.
A. granulatus (Hustache, 1938b: 288). Bolivia. A. minutus Anderson & Morrone, 1996: 264. Peru.
A. hirsutus Morrone, 1994b: 20. Chile. A. planirostris Anderson & Morrone, 1996: 265. Peru.
A. hydanius Morrone, 1994b: 20. Bolivia. A. tricarinatus Anderson & Morrone, 1996: 266.
A. jorgei Morrone, 1994b: 21. Chile. Peru.
A. lidiae Morrone, 1994b: 22. Bolivia.
A. manu Morrone, 1994b: 22. Peru. Antarctobius Fairmaire, 1885: 58 [type species:
A. morio (Voss, 1954: 252). Peru. A. lacunosus Fairmaire, 1885, subsequent de-
A. nivalis (Kuschel, 1949: 45). Chile. signation by Morrone, 1992a]; Morrone,
A. pampaensis (Voss, 1954: 256). Peru. 1992a: 4 (revision).
A. peruvianus (Voss, 1954: 253). Peru. A. abditus (Enderlein, 1907: 57). Argentina (Islas
A. puncticollis (Hustache, 1938b: 286). Bolivia. Malvinas).
A. scrobicollis (Voss, 1954: 256). Peru. A. bidentatus (Champion, 1918: 176). Argentina
A. similaris (Voss, 1954: 250). Peru. (Islas Malvinas).
A. subimpressus (Voss, 1954: 250). Peru. A. falklandicus (Enderlein, 1907: 53) [=A. brac-
A. subtuberculatus (Voss, 1954: 257). Peru. teatus (Enderlein, 1907: 55)]. Argentina (Islas
A. sulcicollis (Hustache, 1938a: 182). Bolivia. Malvinas).
MORRONE, J. J. Checklist of the species of Cyclominae 15
A. germaini (Kolbe, 1907: 105) [= A. griseus (Ger- nis (Germain, 1911: 205); G. fulvitarsis (Husta-
main, 1895: 451) (non Guerin, 1839)]. Argen- che, 1926: 194)]. Argentina and Chile.
tina and Chile. G. laevirostris (Germain, 1895: 583) [= G. quadri-
A. hyadesii Fairmaire, 1885: 58 [= A. laevigatus tuberculatus (Champion, 1918: 51)]. Argentina
(Germain, 1896: 801)]. Argentina and Chile. and Chile.
A. lacunosus Fairmaire, 1885: 59. Argentina and G. lugens (Germain, 1895: 586). Argentina and
Chile. Chile.
A. rugirostris Champion, 1918: 53 [= A. rugicollis G. ovatus (Boheman, 1842: 191) [= G. tristis (Ger-
(Schenkling & Marshall, 1931: 11)]. Chile. main, 1895:481)]. Chile.
A. vulsus (Enderlein, 1907: 50) [= A. gibber (Ender- G. philippii (Germain, 1896: 805). Chile.
lein, 1907: 51); A. griseonotatus (Champion, G. planipennis (Blanchard, 1851: 346). Chile.
1918: 179)]. Argentina (Islas Malvinas). G. punctiventris (Germain, 1895: 596). Chile.
A. yefacel Morrone, 1992a: 10. Chile. G. rugipennis (Blanchard, 1851: 346) [= G. varico-
sus (Blanchard, 1853: 238); G. antarcticus
Falklandiellus Kuschel, 1950: 14 [type species: (Germain, 1895: 581); G. katerensis (Cham-
Falklandius suffodens Enderlein, 1907, by pion, 1918: 52)]. Argentina and Chile.
original designation]; Morrone, 1995b: 106 G. salebrosus (Enderlein, 1907: 44). Argentina (Is-
(revision). las Malvinas).
F. suffodens (Enderlein, 1907: 69) [= F. inaequalis
(Champion, 1918: 184)]. Argentina (including Haversiella Schweiger, 1958: 42 [type species:
Islas Malvinas) and Chile. Haversia albolimbata Champion, 1918, by origi-
nal designation] [= Haversia Champion, 1918:
Falklandiopsis Morrone & Anderson, 1995: 5 185 (non Rower, 1913)]; Morrone, 1994e: 299
[type species: Falklandius magellanicus Mo- (revision).
rrone, 1995 by original designation]. H. albolimbata (Champion, 1918: 186). Argenti-
F. magellanica (Morrone, 1992b: 166). Chile. na (Islas Malvinas) and Chile.
Falklandius Enderlein, 1907: 65 [type species: F. Hyperoides Marshall, 1914: 236 [type species: H.
brachyomma Enderlein, 1907, by original de- fragariae Marshall, 1914, by monotypy]; Morro-
signation] [= Otiorhynchus antarcticus Stierlin, ne, 1993a: 18 (revision).
1903]; Morrone, 1992b: 159 (revision). H. balfourbrownei (Kuschel, 1952b: 134). Ar-
F. antarcticus (Stierlin, 1903: 57) [= F. brachyomma gentina.
(Enderlein, 1907: 66)]. Argentina (including Is- H. fragariae Marshall, 1914: 236 [= H. ellipticus
las Malvinas) and Chile. (Hustache, 1926: 196)]. Argentina and Uruguay.
F. chilensis Morrone & Anderson, 1995: 10. Chile. H. murinus (Germain, 1896: 821). Chile.
F. goliath Morrone, 1992b: 163. Argentina (Islas H. subcinctus (Boheman, 1842: 143) [= H. cine-
Malvinas). rascens (Blanchard, 1851: 348); H. incertus
F. kuscheli Morrone, 1992b: 165. Argentina (Islas (Germain, 1896: 808); H. spoliatus (Germain,
Malvinas). 1896: 817); H. vulgaris (Germain, 1896: 812);
F. peckorum Morrone & Anderson, 1995: 12. H. succinctus (Kuschel, 1950: 14)]. Argentina
Chile. and Chile.
F. turbificatus Enderlein, 1907: 67. Argentina H. victus (Germain, 1896: 815). Chile.
(Islas Malvinas).
Lamiarhinus Morrone, 1992c: 419 [type species:
Germainiellus Morrone, 1993c: 125 [type species: L. aelficus Morrone, 1992c, by original de-
Listroderes dentipennis Germain, 1895, by ori- signation].
ginal designation]. L. aelficus Morrone, 1992c: 421. Chile.
G. angulipennis (Germain, 1895: 593). Chile. L. horridus (Germain, 1896: 741). Chile.
G. attenuatus (Germain, 1895: 598). Chile.
G. dentipennis (Germain, 1895: 589). Argentina Lanteriella Morrone, 1992b: 167 [type species:
and Chile. L. microphtalma Morrone, 1992b, by original
G. fulvicomis (Germain, 1895: 481) [= G. fubricor- designation].
16 Rev. Soc. Entomol. Argent. 61 (1-2), 2002
L. microphtalma Morrone, 1992b: 168. Argentina Kuschel, 1946: 136; L. aequivocus Kuschel,
(Islas Malvinas). 1946: 138; L. costirostris of authors (non
Schonherr, 1826)]. Argentina, Brazil, Chile,
Listroderes Schönherr, 1826: 158 [type species: L. and Uruguay. Introduced into Australia, New
costirostris Schonherr, 1826, by original designa- Zealand, and Norfolk island.
tion] [= Listroderes Schonherr, 1823; Listroderus L. hoffmanni Germain, 1895: 501 [= L gracilicornis
Erichson, 1847: 71]; Morrone, 1993b: 273, Germain, 1895: 504]. Chile.
1993d: 118, 1993e: 16, 1995c: 74 (revisions). L. howdenae Morrone, 1993b: 292. Chile.
L. affinis Hustache, 1926: 197. Argentina and L. leviculus Kuschel, 1952b: 133. Argentina.
L. montanus Germain, 1895: 99. Chile.
Chile.
L. nodifer Boheman, 1842: 194 [= L. chalceatus
L. angusticeps Blanchard, 1851: 341 [= L. proxi-
mus (Germain, 1895: 76)]. Chile. Blanchard, 1851: 342; L. calceatus Lacordaire,
L. annulipes Blanchard, 1851: 340. Chile. 1863: 345]. Chile and Peru.
*L. apicalis Waterhouse, 1841: 123 [= L. argen- L. obliquus Klug, 1829: 6 [= L. obliquus Gyllenhal,
tinensis (Hustache, 1926: 198)]. Argentina, 1834: 277 (non Klug, 1829); L. lugubris Ger-
Brazil, Chile, Paraguay, and Uruguay. Intro- main, 1895: 65; L. obliquatus, Bosq 1935: 331].
duced into USA. Argentina, Brazil, and Uruguay. Introduced into
L. bimaculatus Boheman, 1842: 187 [= L. acutes- South Africa.
quamosus Germain, 1895: 80; L. costulatus L. obrieni Morrone, 1993b: 299. Chile.
Germain, 1895: 86; L. chilensis Germain, L. paranensis Hustache, 1926: 199. Argentina.
1895: 88; L. reticulatus Germain, 1895: 96; L. L. punicola Kuschel, 1949: 42. Bolivia, Chile, and
sticticus Germain, 1895: 93; L. subaeneus Peru.
(Germain, 1895: 97)]. Argentina and Chile. L. pusillus Hustache, 1926: 197. Argentina.
L. brevirostris Germain, 1895: 496. Chile. L. robustior Schenkling & Marshall, 1931: 9 [= L.
L. brevisetis Hustache, 1926: 198. Argentina. robustus Germain, 1895: 64 (non Waterhouse,
L. bruchi Hustache, 1926: 197. Argentina. 1841)]. Chile.
L. charybdis Morrone, 1993b: 281. Argentina. L. robustus Waterhouse, 1841: 122 [= L. arauca-
L. cinerarius Blanchard, 1851: 349 [= L. liliputanus nus (Germain, 1895: 84)]. Chile.
Germain, 1895: 90; L. parvulus (Germain, 1896: L. scylla Morrone, 1993b: 311. Argentina.
823)]. Chile. L. trivialis Germain, 1895: 82. Chile.
L. confusus Hustache, 1926: 196. Argentina and L. tuberculifer Blanchard, 1851: 344 [= L. inae-
Brazil. qualis Blanchard, 1851: 344; L. binodosus
L. costirostris Schonherr, 1826: 158 [= L. costirostris Germain, 1895: 57]. Chile.
Gyllenhal, 1834: 277]. Argentina, Brazil, Para- L. uruguayensis Kuschel, 1952b: 131. Brazil and
guay, and Uruguay. Uruguay.
L. curvipes Germain, 1895: 55. Chile. L. wagneri Hustache, 1926: 195. Argentina and
L. delaiguei Germain, 1895: 63 [= L. magellanicus Uruguay.
Germain, 1895: 62; L. praemorsa (Lea, 1899: L. wittei Hustache, 1926: 195. Argentina.
139); L. delaignei (Germain, 1911: 205)]. Argen-
tina, Chile, and Uruguay. Listronotus Jekel, 1865: 566 [type species: Rhyn-
L. desertorum Germain, 1895: 488 [= L. histrio chaenus caudatus Say, 1824, subsequent desig-
Germain 1895: 491]. Chile. nation by Henderson, 1940] [= Macrops Kirby
*L. difficilis Germain, 1895: 68 [= L. hypocrita 1837: 199 (non Wagler, 1830, nec Burmeister,
Hustache, 1926: 197; L. novica (French, 1908: 1835); Hyperodes Jekel, 1865: 566; Anchode-
754); L. nociva (Lea, 1909: 174); L. vicinus mus LeConte, 1876: 181 (non Motschulsky,
Hustache, 1926: 199]. Argentina, Chile, and 1864); Lixellus LeConte, 1876: 182; Mascarau-
Uruguay. Introduced into USA. xia Desbrochers des Loges, 1898: 52; Relistrodes
L. elegans Hustache, 1926: 194. Argentina and Brèthes, 1910: 209; Aulametopiellus Brèthes,
Uruguay. 1926: 415; Mascaranxia Bosq; 1935: 330;
L. erinaceus Germain, 1895: 499. Chile. Pseudyperodes Hustache, 1939: 49]
L. fallax Germain, 1895: 59. Chile. L. angustatus (Champion, 1902: 122). Belize,
L. foveatus (Lea, 1928: 378) [== L aequicocus Honduras, Mexico, and Nicaragua.
MORRONE, J. J. Checklist of the species of Cyclominae 17
L. apicalis (Hustache, 1926: 209). Argentina and L. laevis (Hustache, 1926: 211). Argentina, Boli-
Chile. via, and Paraguay.
*L. appendiculatus (Boheman, 1842: 192) [= L. flo- L. lineolaticollis (Blanchard, 1851: 372) [= L. affinis
ridensis Blatchley, 1916: 160; L. leucozonatus (Hustache, 1926: 211)]. Argentina, Chile, and
Chittenden, 1926: 341; L. impressus Van Dyke, Uruguay.
1929: 106]. Honduras and Mexico. L. lucens (Hustache, 1926: 208). Argentina and
L. argentinensis (Hustache, 1926: 208). Argentina, Paraguay.
Paraguay, and Uruguay. *L. lutulentus (Boheman, 1843: 165) [= L. angustus
*L. blandus (Henderson, 1940: 282). Honduras. (LeConte, 1876: 181)]. Mexico.
L. bonariensis (Kuschel, 1955: 289) [= L. setosus *L. maculicollis (Kirby, 1837: 200) [= L. mirabilis
(Hustache, 1929: 299), (non LeConte, 1876, (Dietz, 1889: 49); L. myasellus (Dietz, 1889:
nec Hustache, 1926); L. griseus (Marshall, 48)]. Mexico.
1937: 325) (non Hustache, 1926)]. Argentina, *L. manifestus Henderson, 1940: 269. Mexico.
Bolivia, Chile, and Uruguay. Introduced into L. marginalis O'Brien, 1977: 810. Mexico and
Australia and New Zealand. Nicaragua.
*L. borrichiae O'Brien, 1981: 84. Mexico. L. marginicollis (Hustache, 1926: 209). Argentina
L. bosqi (Hustache, 1926: 216). Argentina, Bolivia, and Uruguay.
and Uruguay. L. meridionalis O'Brien, 1977: 812. Costa Rica,
L. breyeri (Brethes, 1910: 210). Argentina and Honduras, and Mexico.
Uruguay. L. minutus (Blanchard, 1851: 373). Argentina, Bo-
L. carinicollis (Hustache, 1926: 214). Argentina livia, and Chile.
and Paraguay. L. nigropunctatus (Suffrian, 1870: 228). Cuba.
L. cinnamoneus (Hustache, 1926: 203). Argentina *L. obtectus (Dietz, 1889: 47). Mexico.
and Uruguay. *L. oregonensis (LeConte, 1857: 55) [= L. impressi-
*L. cryptops (Dietz, 1889: 32). Dominican Re- frons LeConte, 1876: 134; L. tessellatus Casey,
public, Mexico, and Nicaragua. 1895: 828; L rudipennis Blatchley, 1916: 162;
L. cyrticus (Desbrochers des Loges, 1898: 53). L. latiusculus of authors (non Boheman,
Argentina, Bolivia, Chile, and Uruguay. Intro- 1842)]. Mexico.
duced into France. L. ornatipennis (Blanchard, 1851: 371). Chile.
L. dauci (Brèthes, 1926: 416) [= L. ruber (Hustache, L. pampaensis (Voss, 1954: 259). Peru.
1926: 207); L. danei (Bosq, 1935: 328)]. Ar- L. plumosiventris O'Brien, 1977: 813. Mexico.
gentina and Uruguay. L. puncticollis (Hustache, 1926: 213) [= L. puntico-
*L. dietrichi (Stockton, 1963:145). Belize, Costa Ri- llis (Hustache, 1926: 213)]. Argentina, Bolivia,
ca, Honduras, Mexico, Nicaragua, and Panama. and Paraguay.
L. dorytomoides (Hustache, 1926: 212). Argenti- *L. punctiger LeConte, 1876: 135 [= L. gracilis
na and Uruguay. LeConte, 1876: 135; L. bagoiformis Cham-
L. durangoensis O'Brien, 1977: 807 [= L. duran- pion, 1902: 120; L leechi Sleeper, 1955: 156;
goenis O'Brien, 1977: 807]. Mexico. L. teretirostris of authors (non LeConte,
*L. echinodori O'Brien, 1977: 808. Honduras 1857)]. Mexico.
and Mexico. L. pusillus (Hustache, 1926: 215: 215). Argentina
L. elongatus (Hustache, 1939: 49). Argentina, and Paraguay.
Brazil, Paraguay, and Uruguay. *L. rotundicollis LeConte, 1876: 132 [= L. cribri-
L. geminatus (Hustache, 1926: 206). Argentina, collis LeConte, 1876: 134; L. setosus Pierce,
Bolivia, Chile, and Uruguay. 1907: 259 (non LeConte, 1876)]. Costa Rica
L. griseus (Hustache, 1926: 205). Argentina, Chile, and Mexico.
Paraguay, and Uruguay. L. rufomarginatus (Hustache, 1939: 48). Argentina.
*L. grypidioides (Dietz, 1889: 35) [= L. wickhami *L. salicorniae O'Brien, 1981: 115. Mexico.
(Dietz, 1889: 35)]. Mexico. *L. scapularis Casey, 1895: 828. Mexico and Ni-
*L. hirtellus (Dietz, 1889: 46). Guatemala and caragua.
Mexico. L. setosipennis (Hustache, 1926: 205) [= L. setosus
*L. humilis (Gyllenhal, 1834: 284) [= L. subcribra- (Hustache, 1926: 203); L. setossus (Hustache,
tus (Dietz, 1889: 42)]. Mexico. 1926: 203)]. Argentina, Brazil, and Uruguay.
18 Rev. Soc. Entomol. Argent. 61 (1-2), 2002
L. sondondoanus (Voss, 1954: 260). Peru. P. australis Germain, 1896: 747. Argentina and
L. sulcipennis (Boheman, 1834: 346). Brazil. Chile.
L. testaceipes (Champion, 1902: 122). Mexico. P. championi (Kuschel, 1952b: 136) [= P. exsculpti-
L. vulgaris (Hustache, 1926: 206). Argentina, Bo- collis (Champion, 1918: 182) (non Enderlein,
livia, Paraguay, and Uruguay. 1907)]. Argentina (Islas Malvinas).
L. weiseri (Hustache, 1926: 204). Argentina and P. dubius (Germain, 1896: 730). Chile.
Bolivia. P. elguetai Morrone, 1994b: 47. Chile.
P. exsculpticollis (Enderlein, 1907: 60) [= P. insqua-
Macrostyphlus Kirsch, 1889: 25 [type species: M. meus (Enderlein, 1907: 62)]. Argentina (Islas
gualcalae Kirsch, 1889, by monotypy]; Morrone, Malvinas).
1994b: 4 (revision). P. fasciculiger (Blanchard, 1851: 342) [= P. frigidus
M. bilbo Morrone, 1994b: 7. Ecuador. (Germain, 1896: 734); P. sobrinus (Germain,
M. coelorum (Olliff, 1891: 72). Ecuador. 1896: 732)]. Chile.
M. frodo Morrone, 1994b: 9. Colombia. P. hispidus (Germain, 1896: 726). Chile.
M. gandalf Morrone, 1994b: 9. Colombia. P. inaequalis Germain, 1896: 751. Chile.
M. gualcalae Kirsch, 1889: 25. Colombia. P. midas Morrone, 1994b: 52. Chile.
M. howdenorum Morrone, 1994b: 10. Ecuador. P. nigrinus (Fairmaire, 1884: 503) [= P. nigrinus
M. peruvianus Morrone, 1994b: 11. Peru. (Germain, 1896: 801) (non Fairmaire, 1884); P.
M. sturmi Morrone, 1994b: 11. Colombia. obscurus (Germain, 1896: 832); P. obscurusni-
M. transatlanticus (Kirsch, 1889: 24). Ecuador. grinus (Germain, 1911: 205)]. Argentina and
M. venezolanus Morrone, 1994b: 12. Venezuela. Chile.
P. obrienorum Morrone, 1994b: 54. Bolivia and
Nacodius Morrone, 1994a: 3 [type species: Na- Peru.
codius martitae Morrone, 1994a, by original P. pusillus Morrone, 1994b: 54. Chile.
designation]. P. scaber (Enderlein, 1907: 63). Argentina (Islas
N. alectrus Morrone, 1994a: 5. Ecuador. Malvinas).
N. brevirostris (Voss, 1954: 251) [= N. incanus P. sylvanius Morrone, 1994b: 56. Chile.
(Kuschel, 1986: 114)]. Peru. P. torosus Morrone, 1994b: 57. Chile.
N. martitae Morrone, 1994a: 6. Peru. P. tothus Morrone, 1994b: 58. Chile.
N. omissus (Kuschel, 1952b: 129). Ecuador. P. tuberosus Germain, 1896: 749. Chile.
P. verrucosus (Germain, 1896: 728). Chile.
Neopachytychius Hustache, 1939: 55 [type spe- P. vulgaris Morrone, 1994b: 60. Argentina and
cies: N. squamosus Hustache, 1939, by original Chile.
designation] [= Pernotaris Voss, 1943: 232];
Marvaldi, 1994: 58 (revision). Rupanius Morrone, 1995a: 604 [type species: R.
N. squamosus Hustache, 1939: 56 [= N. squami- carinatus Morrone 1995a, by original desig-
ger(Voss, 1943: 233)]. Argentina, Bolivia, Chi- nation].
le, and Uruguay. R. carinatus Morrone, 1995a: 604. Colombia.
Philippius Germain, 1895: 314 [type species: P. Telurus Kuschel, 1955: 288 [type species: Antarc-
superbus (Reed, 1872), subsequent designation tobius laticauda Champion, 1918, by original
by Wibmer & O'Brien, 1986]; Morrone 1990: designation] [= A. dissimilis Fairmaire, 1885];
430 (revision). Morrone & Anderson, 1995: 6 (revision).
P. superbus (Reed, 1872: 354) [= P. insignis Ger- 7. caudiculatus Morrone & Anderson, 1995: 8.
main, 1895: 315]. Argentina and Chile. Argentina and Chile.
T. dissimilis (Fairmaire, 1885: 60) [= T. laticauda
Puranius Germain, 1895: 313 [type species: P. (Champion, 1918: .54)]. Chile.
inaequalis Germain, 1896, subsequent designa-
tion by Morrone, 1994b] [= Puranus Germain, Trachodema Blanchard, 1849: pi. 24 [type species:
1911: 205; Reichertia Enderlein, 1912: 31]; T. tuberculosa Blanchard, 1849, by monotypy];
Morrone, 1994b: 37 (revision). Morrone, 1992c: 417 (revision).
P. argentinensis Morrone, 1994b: 42. Argentina. T. minuta (Hustache, 1930: 269). Chile.
MORRONE, J. J. Checklist of the species of Cyclominae 19
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