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yX 25^^3u.x_ ( Bull. nut. Hist. Mas. Loiul. (Bot.)32(1): 13-59 Issued27June2002 Catalogue of the lichens of Thailand PATRICIA A. WOLSELEY DepartmentofBotany, TheNaturalHistoryMuseum, CromwellRoad, LondonSW75BD BEGONAAGUIRRE-HUDSON 58 WillowhayneDrive, Walton-on-Thames, SurreyKT122NR PATRICKM. McCarthy AustralianBiologicalResourcesStudy, GPOBox787, Canberra, A.C.T. 2601,Australia Synopsis. AcatalogueofthelichensofThailandisprovidedbasedonpublishedreports.Itincludespublicationdetailsof554 speciesandinfra-specifictaxatogetherwithinformationondistributioninThailand,occurrenceinothersouthandsouth-east Asiancountries,literaturereferencesandthelocationsofnamedspecimens. INTRODUCTION BKF: Forest Herbarium, Royal Forest Department, Bangkok, Thailand. BM: Herbarium, Botany Department, The Natural History Before the 1990s, knowledge ofthe diversity and distribution of Museum, London, UK. the lichen flora ofThailand was based almost entirely on collec- C: Herbarium,BotanicalMuseum,UniversityofCopenhagen, tions made by a handful offoreign visitors in the first halfofthe Denmark. twentieth century (Wolseley & Aguirre-Hudson, 1991). During a CANB: Australian National Herbarium, CSIRO, Canberra, research project on lichens as environmental indicators based at Australia. the Natural History Museum, London, acomprehensive surveyof CMU: Herbarium,BiologyDepartment,ChiangMaiUniversity, literature concerning lichens in south-east Asia was made and a Thailand. d(aAtgauibrarsee-Hcuodnssotrnuc&tedW,olswehliecyh, f1o9r94m)s.tIhtewabassinsotofpotshsiisblcehedcukrliinsgt CDOULKOE:: HHeerbbaarriiuumm,,BCootlaonryadoDeUpnairvtemresintty,,CDoulkoeradUnoi,veUrSsAit.y, Dur- thistimetotracealltypesdescribedintheliterature,ortoconfirm ham, NorthCarolina, USA. the accuracy ofmany earlier identifications. Since then there has E: Herbarium, Royal BotanicGarden,Edinburgh,UK. been a rapid and productive expansion oflichenological research FH: Farlow Refence Library and Herbarium ofCryptogamic by Thai practitioners, especially at the Lichen Research Unit, Botany, Harvard University, Cambridge, Massachusetts, Ramkhamhaeng University, Bangkok, further expanding the list USA. and including many newly described species (Boonpragob et al., GB: Herbarium,BotanicalMuseum, Gothenburg, Sweden. c1h9a9n8t;arPaoo&prCaonpgpinets,al2.,0021)9.99T;heVopnrgessheentwacrhaetckleitstali,s c1o9n9c9e;ivHeodma-s H: Herbarium, Botanical Museum, University of Helsinki, Finland. a working document for all researchers in south-east Asia to KF: Herbarium, ForestResearchInstitute, Seoul, Korea. provide both the literature sources and herbaria in which speci- KYO: Herbarium,BotanyDepartment,KyotoUniversity,Japan. mens from Thailand may be seen. L: Rijksherbarium, Leiden,Netherlands. LG: Herbarium,BotanyDepartment,LiegeUniversity,Belgium. OSA: Herbarium, OsakaMuseumofNatural History,Japan. SCOPEANDARRANGEMENT PSU: Herbarium, Department of Biology, Prince of Songkla University, Haad Yai,Thailand. Thiscataloguedocumentsall554currentlyacceptedtaxathathave RAMK: Herbarium, Biology Department, Ramkhamhaeng Uni- beenreportedinthe literature fromThailand, aswell as synonyms versity, Bangkok,Thailand. previously applied to Thai specimens. Genera and species are TNS: Herbarium,DepartmentofBotany,NationalScienceMu- arranged in alphabetical order, and generic and family concepts seum,Tokyo,Japan. followHawksworthetal.(1995).Wherenewcombinationshavenot TUR-V: Vainio herbarium. Biology Department, University of beenmade,taxahavebeenplacedinthecurrentgenuswithinsquare Turku, Finland. brackets []. Accepted names are shown in bold and synonyms in UPS: Botanical Museum,UppsalaUniversity, Sweden. italics. Distribution within Thailand is given where possible by US: United States National Herbarium, Botany Department, provinceincludingnewprovinces andtheirformernames (Fig. 1). SmithsonianInstitution, WashingtonDC, USA. Literature pertaining to distribution in Thailand is cited and refer- Distributionoftaxaininothercountriesinadjacentareasofsouth ences provided. Location ofspecimens in herbariais indicatedby and south-east Asia is indicated underthe subheading 'Asia'. We abbreviationsfollowingIndexHerbariorum(Holmgrenetal., 1990) haveavoidedmakingtaxonomicjudgementsonspecimenswehave as shownbelow: notexamined.Consequently,anumberofhighlydubiousidentifica- AAU: HerbariumJutlandicum,BotanicalInstitute,Universityof tionsremain,asdospecieswhichwillalmostcertainlybetransferred Aarhus,Denmark. toothergenera. ®TheNaturalHistoryMuseum.2002 14 P.A.WOLSELEY,B.AGUIRRE-HUDSONANDP.M.MCCARTHY KeytoprovincesofThailand Northernregion CATALOGUEOFTHELICHENSOFTHAILAND 15 Specimens. CANB. Arthonia obscuratula Zahlbr., Cat. Lick. Univ. 2: 71 Asia. India, Papua New Guinea, Philippines, Sri Lanka. (1922). Arthoniastenographellasubsp. obscurata Vain. Anthracothecium Hampe ex A. Massal.; Pyrenulaceae Thailand. Trat (Koh Chang I.). Vainio (1909: 140, asArthonia stenographella subsp. obscurata). Anthracothecium macrosporum Miill. Arg., Linnaea 63: Specimen. TUR-V (holotype). 44(1888). — Asia. Anthracotheciumpyrenuloidesauct. non (Mont.) Miill. Arg. Arthoniaochrodes Nyl., in Willey, Syn. Gen. Arthonia: 10 Thailand. Chumphon (Koh Tao I.). Paulson (1930: 99, as (1890). Anthracotheciumpyrenuloides). Specimens. BM. Thailand. Trat(KohChang I.). Vainio(1909: 142). Asia. India, Indonesia. Specimen. TUR-V. — Asia. Anthracotheciumparameroides (Vain.) Zahlbr., Cat. Lick. t/mv. 1:466(1922). Arthoniaperpallens Nyl.,Ann. Sci. Nat., Bat., sen 4, 20: Bottariaparameroides Vain. 231 (1863). THAILAND. Trat (Koh Chang I.). Vainio (1907: 179; 1909: 143, Thailand. Trat(KohChang L). Vainio(1909: 141). bothasBottariaparameroides). Specimens. TUR-V. Specimen. TUR-V (holotype). — Asia. Asia. Indonesia. Arthonia rhizophorae Vain., Bat. Tidsskr. 29: 141 (1909). Anthracotheciumprasinum (Eschw.) R.C. Harris, in Egan, Thailand. Trat (KohChang I.). Vainio(1909: 141). Bryologist90: 163(1987). Specimen. TUR-V (holotype). Thailand. Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani — (Huay Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre- Asia. Hudson (1997a: 333), Boonpragobetal. (1998: 216). Arthonia varia Nyl. van stenographella (Nyl.) Vain.,Ann. Specimens. BM, RAMK. Acad. Sci. Fenn., sen A, 15(6): 393 (1921). Asia. PapuaNewGuinea. ArthoniastenographellaNyl. Anthracothecium roseum (Vain.) Zahlbr., Cat. Lick. Univ. Thailand. Trat (Koh Chang I.). Vainio (1909: 140, as A. 1: 468 (1922). stenographella). Bottaria roseaVain. Specimen. TUR-V (holotype). Thailand. Trat (Koh Chang I.). Vainio (1907: 179; 1909: 143, Asia. Philippines, Sri Lanka. bothasBottaria rosea). Arthopyrenia A. Massal.; Arthopyreniaceae Specimen. TUR-V (holotype). — Asia. Arthopyreniasiamensis Vain., Suom. Elain-ja Kasvit. Seuran Van. Julk. 1(3): 55 (1921). Arthonia Ach.; Arthoniaceae Thailand. ChiangMai (DoiSuthepN.P). Vainio(1921: 55). Arthonia complanata Fee, Essai Crypt. Ecorc. Offic.: 54 Specimen. TUR-V (holotype). (1824). — Asia. Thailand. Trat (KohChangI.). Vainio (1909: 139). Arthothelium A. Massal.; Arthoniaceae Specimen. TUR-V. Asia. China, Indonesia,PapuaNewGuinea, Philippines. Arthothelium monosporum (Vain.) Zahlbn, Cat. Lick. Univ. 2: 129(1922). Arthonia aff. mira R. Sant., Symb. Bat. Upsal. 12(1): 77 Arthoniamonospora Vain. (1952). Thailand. Nakhon Ratchasima(KhaoYaiN.P.). Boonpragobet Thailand. Chiang Mai (Doi Suthep N.P). Vainio (1921: 54, as Arthoniamonospora). al. (1998:216). Specimens. RAMK. Specimen. TUR-V (holotype). — — Asia. Asia. P.A.WOLSELEY,B.AGUIRRE-HUDSONANDP.M.MCCARTHY Arthothelium ruanum (A. Massal.) Korb., Parerg. Lich.: Specimens. CANB. 263(1861). Asia. India, Myanmar. Arthonia ruana A. Massal. Brigantiaea Trevis.; Brigantiaeaceae Thailand. Trat (KohChang I.). Vainio (1909: 139, asArthonia ruana). Brigantiaea leucoxantha (Spreng.) R. Sant & Hafellner, in Specimen. TUR-V. Hafellner& Bellemere, Nova Hedwigia 35: 246 (1981). — Asia. Thailand. Chiang Mai (Doi Suthep N.P.), Nakhon Ratchasima (KhaoYai N.P), Uthai Thani (Huay Kha Khaeng Wildlife Sanctu- Arthothelium ruanum var.phaeosporum (Vain.) Zahlbr., ary). Wolseley &Aguirre-Hudson(1997a: 333), Boonpragobetal. Cat. Lich. Univ. 2: 132(1922). (1998: 216). Arthonia ruana var.phaeospora Vain. Specimens. BM, RAMK. Thailand. Trat (Koh Chang I.). Vainio (1909: 139, asArthonia Asia. India,Malaysia,PapuaNewGuinea,Philippines,SriLanka, ruana var.phaeospora). Taiwan. Specimen. TUR-V (holotype). — Brigantiaea tricolor(Mont.) Trevis., Spighe e Paglie: 9 Asia. (1853). Thailand. Chiang Mai (Doi Suthep N.P, Chiang Dao N.P), Arthotheliumspectabile Flotow ex A. Massal., Ric. Auton. Lampang(DoiKhunTanN.P).Wolseley&Aguirre-Hudson(1997a: Lich. Cro5r.: 54 (1852). 333). Thailand. Chumphon (KohTaoI.). Paulson (1930: 99). Specimens. BM, CMU. Specimen. BM. Asia. India,Indonesia,Malaysia,PapuaNewGuinea,SriLanka, Asia. China, India, Malaysia, Sri Lanka. Taiwan. Bacidia De Not.; Bacidiaceae Buellia De Not.; Physciaceae Bacidiaaffinis Vain.,Acta Soc. Fauna Fl. Fenn. 53(1): Buellia blastenioides Vain., Hedwigia 46: 171 (1907). & 146 154(1922). Thailand. Trat(KohChangI.). Vainio(1909: 114). Bacidiaatrosanguinea Th. Fr. Specimen. TUR-V (holotype). Thailand. Chumphon(KohTaoI.).Paulson(1930:99,asBacidia Asia. — atrosanguinea). Specimen. BM (notlocated). Buelliapleiophoroides var. meiospermoides Vain., Sot. — Tidsskr. 29: 113(1909). Asia. Thailand. Chiang Mai (Doi Suthep N.P), Trat (Koh Chang I.), Bacidiamanilensis var. siamensis Vain., Suoml. Elain-ja UthaiThani(HuayKhaKhaengN.P).Vainio(1909: 113),Wolseley Kasvit. Seuran Van. Julk. 1(3): 46 (1921). &Aguirre-Hudson (1997a: 339). Thailand. Chiang Mai (Doi SuthepN.P). Vainio (1921: 46). Specimens. BM,TUR-V (holotype). — Specimen. TUR-V (holotype). Asia. — Asia. Buelliastramineoatra Vain., Hedwigia 46: 172 (1907). Bacidia umbrina var. turgida Th. Fr., Lichenogr. Scand. 1: Thailand. Trat (KohChangI.). Vainio (1909: 115). 365 (1874). Specimens. TUR-V (syntypes). Thailand. Chumphon (KohTaoI.). Paulson (1930: 99). — Asia. BM Specimen. (not located). — Buelliasubdives Vain., Hedwigia 46:172 (1907). Asia. Thailand. Trat(KohChangI.).Vainio(1907: 172; 1909: 114). BatheliumAch.; Trypetheliaceae Specimens. TUR-V (syntypes). — Bathelium madreporiforme (Eschw.) Trevis., Spighe e Asia. Paglie: 20 i\S53). Bulbothrix Hale; Parmeliaceae Laureramadreporiformis (Eschw.) Riddle. Thailand. NakhonRatchasima(KhaoYai N.P.). Boonpragobet Bulbothrixgoebelii (Zenker) Hale, Smithsonian Contr. Bat. al. (1998: 217, asLaureramadreporiformis). 32: 14(1976). CATALOGUEOFTHELICHENSOFTHAILAND 17 Parmeliaaddenda Vain. Bulbothrixqueenslandica (Elix & G.N. Stevens) Elix, Mycotaxon 47: 126 (1993). Thailand. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P., QueenSirikitBotanical Garden),Trat(KohChang I.), UthaiThani Thailand. Chiang Mai (Queen Sirikit Botanical Garden), Loei (Huay Kha Khaeng WildHfe Sanctuary). Vainio (1909: 107, as (NahaewN.P.),Phitsanulok(PhuHinRongklaN.P.).Pooprangetal. Parmelia addenda). Hale (1976a: 14), Mongkonsuk et al. (1996: (1999: 119). 16),Wolseley & Aguirre-Hudson (1997a: 333). Specimens. RAMK. Specimens. C(lectotype),TUR-V(isolectotype),BKF,BM,CMU, — Asia. US,Queen SirikitBotanical Garden Herbarium. Asia. India, Indonesia, Malaysia, Papua New Guinea, Philip- Bulbothrixsensibilis (J. Steiner&Zahlbr.) Hale, pines,Taiwan. Phytologia 2S: 4Sl (1974). Thailand. Loei (NahaewN.R). Pooprangetal. ( 1999: 119). Bulbothrixhypochraea (Vain.) Hale, Phytologia 28: 480 (1974). Specimens. RAMK. THAILAND. Chiang Mai (Queen Sirikit Botanical Garden). Asia. India, Sri Lanka. Pooprangetal. (1999: 118). Bulbothrixsetschawensis (Zahlbr.) Hale,Phytologia 28: Specimens. RAMK. 481 (1974). — Asia. Thailand. Chiang Mai (Doi Inthanon N.P.), Lampang, Uthai Thani(HuayKhaKhaengWildlifeSanctuary).Wolseley&Aguirre- Bulbothrixisidiza (Nyl.) Hale, Phytologia 28: 480 (1974). Hudson (1997a: 333). Thailand. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P., Specimens. BKF, BM,CMU. ChiangDaoN.P.,Maetang),Lampang(Doi PaeLuang),Loei (Ban Asia. China, India, Nepal. Sui, Phu Kradung), Phitsanulok (Phu Hin Rongkla, N.P.), Uthai Thani(HuayKhaKhaengWildlifeSanctuary).Wolseley&Aguirre- ByssolomaTrevis.; Pilocarpaceae Hudson (1995: 318; 1997a: 333; 1997b: 355-356), Moon et al. (2000a: 99). Byssolomachlorinum (Vain.) Zahlbr., Cat. Lich. Univ. 8: Specimens. BKF, BM, CMU. 233 (1932). Asia. China, India, Indonesia, Malaysia, Papua New Guinea, Thailand. NakhonRatchasima(KhaoYai N.P.). Boonpragobet Philippines, Sri Lanka,Taiwan. al. (1998: 216). Specimens. RAMK. Bulbothrixlaevigatula (Nyl.) Hale, Phytologia 28: 480 (1974). Asia. India,Indonesia,PapuaNew Guinea. ParmeliahookeriTaylor. Byssoloma leucoblepharum (Nyl.) Vain., DanskSot. Ark. Thailand. Chiang Mai (Doi Suthep). Vainio (1921: 37, as 4: 23 (1926). Parmeliahookeri). Thailand. Nakhon Ratchasima (Khao Yai N.P.). Wolseley & Specimen. TUR-V. Aguirre-Hudson (1997a: 333), Boonpragobetal. (1998: 216). — Specimens. BM, RAMK. Asia. Asia. India, Indonesia, Malaysia, Papua New Guinea, Philip- Bulbothrixmeizospora (Nyl.) Hale, Phytologia 28: 480 pines, Singapore, Vietnam. (1974). Byssoloma tricholomum (Mont.) R. Sant., Symb. Bot. Thailand. Chiang Mai (Queen Sirikit Botanical Garden), Loei Upsal. 12(1): 480 (1952). (PhuRhoaN.P.),NakhonNayok(KhaoYaiN.R).Boonpragobetal. (1998: 216), Pooprangetal. (1999: 118). Thailand. Nakhon Ratchasima(KhaoYai N.P.). Boonpragobet al (1998: 216). Specimens. BM, RAMK. Specimens. RAMK. Asia. India, Indonesia, Nepal, SriLanka. Asia. China, Philippines, Singapore. Bulbothrixpigmentacea (Hale) Hale, Phytologia 28: 480 Calenia Mull. Arg. emend. R. Sant.; Gomphillaceae (1974). Thailand. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.), Caleniagraphidea Vain.,Ann. Acad. Sci. Fenn., sen A, 15: Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley & 163(1921). Aguirre-Hudson (1995: 318; 1997a: 333; 1997b: 355-56). Thailand. NakhonRatchasima(KhaoYai N.P.). Boonpragobet Specimens. BKF, BM, CMU. al. (1998: 216). Asia. Malaysia, Philippines. Specimens. RAMK. 18 P.A.WOLSELEY,B.AGUIRRE-HUDSONANDP.M.MCCARTHY Asia. Indonesia, Malaysia, PapuaNewGuinea, Philippines. Rimeliellasubsumpta (Nyl.) Kurok. Calenia aff.phyllogena (Miill. Arg.) R. Sant., Symb. Bot. Thailand. Chiang Mai (Maetang). Kurokawa (1991: 9, as Upsal. 12(1): 349 (1952). Rimeliellasubsumpta); Moonetal. (2000a: 99). Specimen. TNS. Thailand. Nakhon Ratchasima (KhaoYai N.P.). Boonpragobet al. (1998:216). Asia. India, PapuaNewGuinea. Specimens. RAMK. Canomaculina subtinctoria (Zahlbr.) Elix, Mycotaxon 65: Asia. India, PapuaNew Guinea, Philippines, Sri Lanka. 477 (1997). Thailand. Chiang Mai (Queen Sirikit Botanical Garden), Loei Calopadia Ve^zda; Ectolechiaceae (Nahaew N.P.). Mongkonsuk et al. (1996: 17), Pooprang et al. (1999: 119). Calopadiafusca (Miill. Arg.) VeSzda, Folia Geobot. Phytotax.ll: 2\5{\9m). Specimens. RAMK. Thailand. Nakhon Ratchasima (Khao Yai N.P.). Boonpragobet Asia. China, India, Indonesia, Nepal. al. (1998:216). Canoparmelia Elix & Hale; Parmeliaceae Specimens. CANB. Asia. Indonesia, PapuaNew Guinea, Philippines, Sri Lanka. JCoahnnosptaornm&eliVearaddosnp,eMrysaco(tVaaixno.n)2E7l:ix2&78H(a1l9e8,6)i.n Elix, Caloplaca Th. Fr.; Teloschistaceae Parmeliaadspersa Vain. Caloplacaaequata (Hue) Zahlbr., Cat. Lich. Univ. 7: 211 Thailand. Trat (Lem Ngob). Vainio (1907: 168, as Parmelia adspersa). (1931). Thailand. Chumphon (KohTaoI.). Paulson (1930: 99). Specimens. C (isolectotype),TUR-V (lectotypeandsyntype). Specimen. BM (notlocated). Asia. PapuaNew Guinea, Philippines. — Asia. Canoparmelia concrescens (Vain.) Elix & Hale, Mycotaxon 21: 21^ (19^6). Caloplacaflavorubescens (Huds.) Laundon, Lichenologist 8: 147(1976). Thailand. Phitsanulok (Phu Hin Rongkla, N.P.). Moon et al. (2000a: 100). Caloplacaaurantiaca (Lightf.)Th. Fr. Specimens. TNS. Thailand. Chiang Mai (Doi Inthanon N.P.), Chumphon (Koh — Asia. Tao I.). Paulson (1930: 99, as Caloplacaaurantiaca). & Specimens. BM. Canoparmelia ecaperata (Miill. Arg.) Elix Hale, in Elix, Johnston & Verdon, Mycotaxon 27: 278 (1986). Asia. Sri Lanka,Taiwan. Parmeliamalaccensisvan laeteflavensVain. Caloplaca holocarpa (Hoffm.) A.E. Wade, Lichenologist Thailand. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P., 3: 11 (1965). Chiang Dao N.P., Maetang), Loei (Phu Kradung N.P.), Nakhon Caloplacapyracea (Ach.)Th. Fr. Ratchasima(KhaoYaiN.P.),Phitsanulok(PhuHinRongkla,N.P.), Thailand. Chumphon (Koh Tao I.). Paulson (1930: 99, as U38t,haaisTPhaarnmiel(iHaumaaylaKchcaenKshiasevnagrlWaieltdelfilfaeveSnasn)c,tWuoalrys)e.leVyai&niAogu(i1r92r1e:- Caloplacapyracea). Hudson(1995: 318; 1997a: 333; 1997b:355-56),Boonpragobetal. Specimen. BM (notlocated). (1998: 216), Moonetal. (2000a: 100). Asia. China, India. Specimens. BM, CMU, RAMK, TNS, TUR-V (lectotype and syntype ofP. malaccensisvarlaeteflavens), US. [Caloplaca] Placodium testaceorufum Vain., Hedwigia 46: 170(1907). Asia. India, Nepal. Thailand. Trat (Lem Ngob). Vainio (1909: 111, as Placodium Canoparmelia owariensis (Asahina) Elix, Mycotaxon47: testaceorufum). 127 (1993). Specimen. TUR-V (holotype). Pseudoparmeliaowariensis(Asahina) Hale. — Asia. Thailand. ChiangMai(DoiSaget),Loei(BanSui,PhuKradung), Phitsanulok (Phu Hin Rongkla, N.R). Hale (1976b: 39, as Canomaculina Elix & Hale; Parmeliaceae Pseudoparmeliaowariensis).Moonetal. (2000a: 100). Specimens. TNS. Canomaculinasubsumpta (Nyl.) Elix, Mycotaxon 65: 477 (1997). Asia. China(Hong Kong). CATALOGUEOFTHELICHENSOFTHAILAND 19 Canoparmeliapustulescens (Kurok.) Elix, Mycotaxon 47: Asia. China, Indonesia, Nepal, Philippines,Taiwan. 127(1993). Thailand. Chiang Mai (Doi Saget), Loei (Phu Kradung). Moon Cetrelia W.L. Culb.& C.F. Culb.; Parmeliaceae etal. (2000a: 101). Cetrelia braunsiana (Mull. Arg.) W.L. Culb. & C.F. Culb., Specimens. TNS. Contr. U.S. Natl. Herb. 34: 493 (1968). Asia. India. Cetrariacollataf. isidiata Asahina. Canoparmeliasalacinifera (Hale) Elix & Hale, in Elix, Thailand. Chiang Mai (Doi Inthanon N.P). Sato (1962: 3, as Johnston & Verdon, Mycotaxon 27: 279 (1986). Cetraria collata f. isidiata), Wolseley & Aguirre-Hudson (1997a: 333, 337). Pseudoparmeliasalacinifera(Hale) Hale. Specimens. BM,TNS. Thailand. Chiang Mai (Doi Saget, Doi SuthepN.P.), Loei (Phu Kradung N.P.). Hale (1976a: 46, as Pseudoparmelia salacinifera), Asia. China, India, Malaysia, Papua New Guinea, Philippines, Wolseley&Aguirre-Hudson(1995:318),Moonetal. (2000a: 101). Taiwan. Specime—ns. BKF, BM, CMU, TNS, US. Cetreliajaponica (Zahlbr.) W.L. Culb. & C.F. Culb., Contr. Asia. U.S. Natl. Herb. 34: 511 (1968). JCoahnnosptaornm&eliVeartdeoxna,nMay(cToutcakx.)onEl2i7x:&27H9al(e1,98i6n).Elix, TYahiaiNl.aPn).d.WolCsheilaenyg&MaAigu(iDroireI-nHtuhdansoonnN(.1P9)9,7aN:ak3h33o)n,NBaoyoonkpr(aKghoabo etal. (1998:216). Thailand. Chiang Mai (ChiangDaoN.P,Queen SirikitBotani- Specimens. BM, RAMK. cal Garden), Lampang (Doi Khun Tan N.P), Loei (Phu Kradung N.P). Hale(1976a: 53), Moonetal. (2000a: 101). Asia. Indonesia, Malaysia, Taiwan. Specimens. BKF, BM, CMU, TNS. Cetrelia nuda (Hue) W.L. Culb. & C.F Culb., Contr. U.S. Asia. India, Malaysia, Sri Lanka. Natl. Herb. 34: 513 {196S). Thailand. ChiangMai(DoiInthanonN.P).Wolseley&Aguirre- CatillariaA. Massal.; Catillariaceae Hudson (1997a: 337, 339). Catillariatestaceolivens (Vain.) Zahlbr., Cat. Lich. Univ. 4: Specimen. BM. 79 (1927). Asia. China, Taiwan. Lecidea testaceolivensVain. Cetreliaolivetorum (Nyl.) W.L. Culb. & C.F. Culb, Contr. Thailand. Trat (Koh Chang I.). Vainio (1907: 173; 1909: 117, U.S. Natl. Herb. 34: 515 (1968). bothasLecidea testaceolivens). Thailand. ChiangMai(DoiInthanonN.P).Wolseley&Aguirre- Specimen. TUR-V (holotype). — Hudson (1997a: 337, 339). Asia. Specimen. BM. Catillaria unicolor (Vain.) Zahlbr., Cat. Lich. Univ. 4: 83 Asia. China, India, PapuaNew Guinea,Taiwan. (1927). Lecidea unicolorVain. Chaenotheca (Th. Fr.) Th. Fr.; Coniocybaceae Thailand. Trat (Koh Chang I.). Vainio (1907: 173; 1909: 118, Chaenothecatrichialis (Ach.) Th. Fr., NovaActaRegiae bothasLecidea unicolor). Soc. Sci. Upsal, sen 3, 3: 251 (1860). Specimen. TUR-V (holotype). Thailand. NakhonRatchasima(Khao YaiN.P.). Boonpragobet — Asia. al. (1998:216). Specimen. RAMK. Cetrariopsis Kurok.; Parmeliaceae Asia. India. Cetrariopsis wallichiana (Taylor) Kurok., Mem. Nat. Sci. Mus., Tokyo U: 140(1980). Chiodecton Ach.; Roccellaceae Cetrariawallichiana (Taylor) Miill. Arg. Chiodecton congestulum Nyl., Bull. Soc. Linn. Thailand. Chiang Mai (Doi Suthep). Vainio (1921: 36, as Ce- Normandie, ser. 2, 2: 106 (1868). traria wallichiana), Kurokawa (1980: 140), Wolseley & Thailand. Trat(KohChangI.).Thor(1990:41). Aguirre-Hudson (1995: 318; 1997a: 333). Specimens. C, TUR-V. Specimens. BM, PSU, TNS. Asia. India, Indonesia,Malaysia,PapuaNewGuinea,Taiwan. 20 P.A.WOLSELEY,B.AGUIRRE-HUDSONANDP.M.MCCARTHY Chiodecton subsphaerale Nyl., Bull. Soc. Linn. Cladoniadidyma var. vulcanica (Zoll. & Moritzi) Vain., Normandie, sen 2, 2: 106 (1868). Acta Soc. Fauna Fl. Fenn. 4: 145 (1887). Thailand. Trat(KohChangI.). Vainio(1909: 138). Cladonia vulcanicaZoll. Specimens. TUR-V. Thailand. ChiangMai (DoiInthanon N.P.), Loei (Phu Kradung N.P.). desAbbayes(1956: 259,as Cladonia vulcanica),Yoshimura Asia. Malaysia. (1978: 38, as Cladonia vulcanica). Specimens. TNS. Chroodiscus Miill. Arg.; Thelotremataceae Asia. India,Indonesia,PapuaNewGuinea,Philippines,SriLanka, Chroodiscus mirificus (Kremp.) R. Sant., Symb. Bot. Taiwan. Upsal. 12(1): 311 (1952). Thailand. Nakhon Ratchasima(KhaoYai N.P.). Boonpragobet Cladoniafurcata (Huds.) Schrad., Spicil. Fl. Germ.: 107 (1794). al. (1998: 216). Specimens. RAMK. Thailand. NakhonRatchasima(KhaoYai N.P.). Boonpragobet al. (1998:216). Asia. India, Indonesia, Malaysia, Papua New Guinea, Philip- Specimens. RAMK. pines, SriLanka, Vietnam. Asia. China, India, Indonesia,Taiwan. Chrysothrix Mont.; Chrysothricaceae Cladoniagymnopoda Vain.,Acta Soc. Fauna Fl. Fenn. 10: Chrysothrixcandelaris (L.) Laundon, Lichenologist 13: 172(1894). 110(1981) Thailand. Loei (Phu Miang). Yoshimura(1978: 38). Leprariaxanthina Vain. Specimens. AAU, C. TTrhaatil(aKnodh.ChCahniganIg.),MUatiha(iDoTihaSnuith(eHpuaN.yP.,KhDaoiKhIantehnagnoWnilNd.lPi.f)e, Asia. India, Indonesia, Malaysia, Papua New Guinea, Philip- Sanctuary). Vainio (1909: 151, asLeprariaxanthina), Wolseley & pines, Taiwan. Aguirre-Hudson (1997b: 355). & Cladonia modesta Ahti Krog,Ann. Bot. Fenn. 24: 88 Specimens. BM,TUR-V. (1987). Asia. China(Hong Kong), India, Philippines, Sri Lanka. Thailand. Nakhon Ratchasima(KhaoYai N.P.). Boonpragobet al. (1998:216). Cladia Nyl; Cladoniaceae Specimens. RAMK. — Cladiaaggregata (Sw.) Nyl., Compt. Rend. Hebd. Seances Asia. Acad Sci. S3: 8(1876). Cladonia ramulosa (With.) Laundon, Lichenologist6: 225 Thailand. Loei (Phu Miang, Phu Kradung N.P.). Yoshimura (1984). (1978:37). Specimens. AAU, C. Cladoniapityrea(Florke)Fr. Thailand. Chiang Mai (Doi Suthep N.P.), Khu Kawng, Loei Asia. China (Hong Kong), India, Indonesia, Malaysia, Nepal, PapuaNew Guinea, Philippines,Taiwan. (Phu Luang N.P.). Vainio (1921: 46, as C. pityrea), Yoshimura (1978:38, as C.p/ryr^'a). Cladonia P. Browne; Cladoniaceae Specimens. C. Asia. China(incl.HongKong),India,Indonesia,Malaysia,Papua Cladonia calyciformis Nuno, J. Jap. Bot. 47: 161 (1972). NewGuinea, Philippines, Sri Lanka. Thailand. Loei ( Phu KradungN.P.). Yoshimura(1978: 37). Cladonia rangiferina (L.) F.H. Wigg., Primit. Fl. Holsat. Specime—ns. C,TNS (holotype). 90: 29 (1780). Asia. Thailand. Loei (Phu Kradung N.P). Yoshimura (1978: 37), Kurokawa& Kashiwadani (1988: Lich. Ran Crit. Exs. No. 60). Cladonia ciliata var. tenuis (Florke) Ahti,Ann. Bot. Fenn. 15: 8 (1978). Specimens. C, TNS. Cladoniatenuis (Florke)Harm. Asia. China, India,Taiwan. Thailand. PhuMieng.Yoshimura(1978:37,asCladoniatenuis). Cladonia rappii A. Evans, Trans. ConnecticutAcad. Arts Specimens. AAU, C. 38:297(1952). Asia. China, India. CladoniacalycanthaDeliseex Nyl. CATALOGUEOFTHELICHENSOFTHAILAND 21 Thailand. Chiang Mai (Doi Suthep N.P.), Loei (Phu Kradung Suthep N.R, Doi Inthanon N.P.), Khiri Khan, Uthai Thani (Huay N.P.). Yoshimura(1978: 37, as C. calycantha). KhaKhaeng Wildlife Sanctuary). Mongkonsuketal. (1996: 16). Specimens. AAU, BM, C. Specimens. BM,QueenSirikitBotanical Garden Herbarium. Asia. China, India, Indonesia, Malaysia, Papua New Guinea, Asia. Malaysia, PapuaNewGuinea, Philippines. Philippines,Taiwan. Coccocarpia erythroxyli (Spreng.) Swinscow & Krog, Cladoniasiamea des Abbayes, KewBull. 2: 262 (1956). Norweg. J. Bat. 23: 256 (1976). Thailand. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.), CoccocarpiamolybdaeaPers. Loei (Phu KradungN.P.). Yoshimura(1978: 37), Huovinen&Ahti (1986: 182),Stenroos(1986: 164),Kurokawa&Kashiwadani(1988: Thailand. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.), Lich. Ran Crit. Exs. Nos. 61, 62). Nakbon Ratchasima (Khao Yai N.R), Trat (Koh Chang I.), Uthai Thani (Huay Kha KhaengWildlife Sanctuary). Paulson (1930: 99, Specimens. BM (holotype), C, H (isotype), RAMK,TNS. as C. molybdaea), Arvidsson (1982: 62), Boonpragobetal. (1998: Asia. Indonesia, PapuaNewGuinea. 216). Specimens. BM,GB, L, RAMK,TNS. Cladoniasiamea f. evoluta des Abbayes, KewBull. 2: 262 (1956). Asia. China(incl.HongKong),India,Malaysia,Myanmar,Papua NewGuinea, Philippines, Singapore, SriLanka,Taiwan. Thailand. Loei (PhuKradungN.P.). desAbbayes (1956: 262). Specimens. BKF, BM (holotype). Coccocarpiaglaucina Kremp.,Nuovo Giorn. Bat. Ital. 7: — 14(1875). Asia. Thailand. ChiangMai(DoiSuthepN.R),NakhonNayok(Khao Cladoniasiamea f.pulvinata des Abbayes, KewBull. 2: Yai N.R). Boonpragobetal (1998: 216). 263 (1956). Specimens. BM, RAMK. Thailand. Loei (Phu KradungN.P). desAbbayes (1956: 263). Asia. Malaysia, PapuaNewGuinea, Philippines. Specimens. BKF, BM (holotype). — & Coccocarpiapalmicola (Spreng.) Arv. D.J. Galloway, Asia. Bat. Not. 132:242(1979). Cladoniasquamosa (Scop.) Hoffm., Deutschl. Fl. 2: 125 Thailand. Chiang Mai (Queen Sirikit Botanical Garden, Doi (1796). SuthepN.R, Doi InthanonN.R,ChiangDaoN.R), Nakhon Nayok (KhaoYaiN.R),Tak(TonKrabakyaiN.R),UthaiThani(HuayKha Thailand. NakhonRatchasima(KhaoYai N.P.). Boonpragobet Khaeng Wildlife Sanctuary). Arvidsson (1982: 76), Wolseley & al. (1998: 216). Aguirre-Hudson (1997a: 333; 1997b: 355), Mongkonsuk et al Specimens. RAMK. (1996), Boonpragobetal. (1998: 216). Asia. China, India, Indonesia, Papua New Guinea, Philippines, Specimens. BM, L, RAMK, TNS, Queen Sirikit Botanical Gar- Taiwan. den Herbarium. Cladoniasubpityrea Sandst., in Zahlbruckner,Ann. Asia. China(incl.HongKong),India,Malaysia,Myanmar,Papua Naturhist. Mus. Wien 42: 61 (1928). NewGuinea, Philippines, Singapore, SriLanka,Taiwan. Thailand. Loei (Dan Sai). Yoshimura(1978: 38). Coccocarpiapellita (Ach.) Miill. Arg. emend. R. Sant., Specimens. C. Symb. Bat. Upsal. 12(1): 420 (1952). Asia. PapuaNewGuinea, Philippines, Taiwan. Thailand. Nakhon Nayok (Khao Yai N.R), Uthai Thani (Huay KhaKhaengWildlife Sanctuary). Boonpragobetal (1998: 216). Cladonia transindica Ahti,Ann. Bat. Sac. Zool-Bot. Fenn. Specimens. BM, RAMK. Tflnamo' 32(1): 72 (1961). Asia. China (Hong Kong), India, Indonesia, Malaysia, Papua Thailand, localitynotknown, desAbbayes (1964), Ahti (1984: New Guinea, Philippines, SriLanka. 44), Stenroos(1988:212). Specimen. ? CoUemaF.H. Wigg.; Collemataceae Asia. China, Indonesia, Malaysia, Vietnam. Collemacoccophyllum Nyl., Syn. Meth. Lich. 1: 112 CoccocarpiaPers.; Coccocarpiaceae (1858). Thailand. ChiangMai (Doi Suthep). Degelius(1974: 69). Coccocarpiadissecta Swinscow & Krog,Norweg. J. Bat. 23:254(1976). Specimen. TNS. Thailand. Chiang Mai (Queen Sirikit Botanical Garden, Doi Asia. India. 22 P.A.wolseley,B.AGUIRRE-HUDSONANDP.M.MCCARTHY Collema coilocarpum (Miill. Arg.) Zahlbr., Cat. Lich. Dichosporidium Pat.; Chiodectonaceae (//j/v. 3:34(1924). Dichosporidium boschianum (Mont.) Thor, Opera Bot. Thailand. ChiangMai(DoiInthanonN.P.),NakhonRatchasima 103:64(1990). (Khao Yai N.P.), Uthai Thani (Huay Kiia Khaeng Wildlife Sanctu- ary). Wolseley &Aguirre-Hudson (1997a: 333), Boonpragobetal. Thailand. NakhonRatchasima(KhaoYaiN.P). Boonpragobet (1998:217). al. (1998: 217). Specimens. BM, RAMK. Specimen. RAMK. Asia. India, Papua New Guinea, Philippines, Sri Lanka. Asia. China (Hong Kong), India, Malaysia, Philippines, Singa- pore, Sri Lanka. ConotremaTuck.; Stictidaceae Dictyonema C. Agardh ex Kunth; Meruliaceae Conotrema lumbricoides Sipman, in Aptrootet al, Biblioth. Lichenol. 64: 52 (1997). Dictyenema sericeum (Sw.) Berk., J. Bot. (London) 2: 639 (1843). Thailand. Nakhon Sri Thammarat (Khao Luang N.P.). Vongshewaratetal. (1999: 227) Dictyonemasericeumf. thelephora (Spreng.) Parm. Specimens. RAMK. Thailand. Trat (Koh Chang I.). Vainio (1909: 151), Parmasto (1978:110,111). Asia. PapuaNew Guinea. Specimen. TUR-V. & Cresponea Egea Torrente; Roccellaceae Asia. China (incl. Hong Kong), Indonesia, Papua New Guinea, Philippines, Taiwan. Cresponea chloroconia (Tuck.) Egea & Torrente, Mycotaxon^: 3X0 {\992>). Dirinaria (Tuck.) Clem.; Physciaceae Lecanactisdeminuens(Nyl.) Vain. Dirinariaaegialita (Ach.) B. Moore, Bryologist71: 248 Thailand. Trat(KohChangI.).Vainio(1909: \2\,as,Lecanactis (1968). deminuens). Thailand. Chiang Mai Doi Inthanon N.P, Chiang Dao N.P), ( Specimen. TUR-V. Lampang (Doi Khun Tan N.P, Doi Pae Luang N.P). Wolseley & — Aguirre-Hudson (1997a: 333). Asia. Specimens. BM, CMU. Crocynia A. Massal.; Lecideaceae Asia. China (Hong Kong), India, Indonesia, Malaysia, Philip- pines, Sri Lanka, Taiwan, Vietnam. Crocyniapyxinoides Nyl., Sert. Lich. Trop. Labuan Singapore: 31 {\^9\). Dirinariaapplanata (Fee) D.D. Awasthi, in Awasthi & Thailand. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.), Agarwal, J. IndianBot. Soc. 49: 135 (1970). Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani (Huay Kha Thailand. KhiriKhan,UthaiThani(HuayKhaKhaengWildlife Khaeng Wildlife Sanctuary). Wolseley & Aguirre-Hudson (1995: Sanctuary). Wolseley & Aguirre-Hudson(1997a: 333). 320; 1997a: 333, 339; 1997b: 356), Boonpragobetal. (1998: 217). Specimens. BM. Specimens. BM, RAMK. Asia. China(incl.HongKong),India,Indonesia,Malaysia,Papua Asia. Malaysia, Singapore. New Guinea, Philippines, Sri Lanka,Taiwan, Vietnam. Cryptothecia Stirt.; Arthoniaceae Dirinaria confusa D.D. Awasthi, Biblioth. Lichenol. 2: 56 (1975). Cryptothecia Candida (Kremp.) R. Sant., Symb. Bot. Upsal. 12(1): 65 (1952). Thailand. Fang (northernThailand). Yoshimura(1978: 35). Thailand. NakhonRatchasima(KhaoYai N.P.). Boonpragobet Specimen. C. al. (1998:217). Asia. China, PapuaNew Guinea,Taiwan. Specimen. RAMK. Dirinaria consimilis (Stirt.) D.D. Awasthi, in Awasthi & Asia. India, Indonesia, Malaysia, Sri Lanka. Agarwal, J. Indian Bot. Soc. 49: 135 (1970). & Thailand. Chiang Mai (Doi Suthep N.P, Doi Inthanon N.P, Cryptothecia obtecta Patw. Sethy, Biovigyanam 13(2): 46(1987). Chiang Dao N.P), Sara Buri, Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Awasthi (1975: 93), Wolseley & Aguirre- Thailand. NorthernThailand. Wolseley (1991: 240). Hudson (1995: 321; 1997a: 333; 1997b: 355). Specimen. BM. Specimens. BM, CMU. Asia. India. Asia. India, Indonesia, Malaysia, Myanmar, Vietnam.

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