Catalogue of Palaearctic Coleoptera, Vol. 1: Archostemata - Myxophaga - Adephaga

410 Pages·2003·29.651 MB·English

by  I. Lobl, A. Smetana

Catalogue of Palaeai
Structured knowledge of life, as it evolved and is present on our planet,
reąuires cataloguing. Catalogues summarize the work in systematics, they
Palaearctic Coleoptera
enable assessment of faunal diversity and classification, and provide
information on past taxonomic acts. They are also essential in achieving a
universal language in taxonomy that assures unambiguous transfer of biological
information. Erroneous information due to incorrect use of taxonomic names has
Volume 1
an impact in many fields, including genetics, physiology, ecology, plant
protection, naturę consen/ation, and legislation.
Over the years, the Palaearctic Coleoptera have undoubtedly been studied by morę
students that those of other parts of the world, and morę than Archostemata - Myxophaga - Adephaga
representatives of most other orders of insects. It is therefore remarkable that an
overview of the present knowledge of the about 100,000 species of the Palaearctic
Coleoptera is badly trailing that of, e.g., the Nearctic Coleoptera, or Palaearctic
Diptera.
The aim of Catalogue of Palaearctic Coleoptera is to provide a tool that
addresses the most urgent needs: 1) a list of available names, both valid and
invalid, of taxa occurring in the Palaearctic Region, in their verified original
orthography and with the correct publication dates, 2) a complete list of
verified references to primary descriptions, and 3) informative distributions of the
species and subspecies. Thus, the catalogue is expected to respond to
ąuestions related to biodiversity, and to increase the badly needed
nomenclatorial stability.
Catalogue of Palaearctic Coleoptera is a collecth/e work involving many
authors and will be published in eight volumes. Each volume will group taxa with
respect to the adopted classification, and will be "independent", having its own
Reference section and Index.
Edited by
Apollo Books
DK-5771 Stenstrup I. LÓBL & A. SMETANA
Denmark
Apollo
Books
ISBN 87-88757-73-0 (Vol. 1)
Apollo Books
ISBN 87-88757-71-4 (Vols 1-8) 2003
Catalogue of
Palaearctic Colcoptera
Volume 1
Archostemata - Myxophaga -
Adephaga
Edited by
I. LÓBL & A. SMETANA
Paropisthius indicus chinensis Bousąuet & Smetana, 1996 Apollo Books
Stenstrup, 2003
© Copyright 2003 by Apollo Books, Stenstrup, Denmark. CONTENTS
Introduction 6
AU rights reserved. No part of this book may be reproduced or translated in any form,
by print, photoprint, microfilm, microfiche, disc, CD-Rom or any other means without Taxonomic information 7
written permission from the publisher.
Distributional information 8
Bibliographic information 13
Edited by: I. Lóbl & A. Smetana. Acknowledgements 13
Coordinators 15
Printed by: Yinderup Bogtrykkeri A/S. Authors 15
Mailing and E-mail addresses of the authors 17
Published by: New taxonomic and nomenclatorial acts 18
Apollo Books Catalogue 25
Kirkeby Sand 19
Archostemata 25
DK-5771 Stenstrup
Denmark Crowsoniellidae Iablokoff-Kłmzorian, 1983 25
Telephone + 45 62 26 37 37 Micromalthidae Barber, 1913 25
Telefax + 45 62 26 37 80
Cupedidae Laporte, 1836 25
E-mail: [email protected]
Web site: www.apollobooks.com Sikhotealiniidae Lafer, 1996 25
Myxophaga 25
Chaptersfrom this publication should be cited asfollows: Torrindincolidae Steffan, 1964 25
Nilsson, A. N., 2003: Dytiscidae, pp. 35-78 - In I. Lobl & A. Smetana (editors): Hydroscaphidae LeConte, 1874 25
Catalogue of Palaearctic Coleoptera, Vol. 1. Stenstrup: Apollo Books, 819 pp.
Sphaeriusidae Erichson, 1845 25
Adephaga 26
ISBN 87-88757-73-0 (Vol. 1)
Gyrinidae 26
ISBN 87-88757-71-4 (Vols 1-8
Haliplidae 30
Trachypachidae 33
Datę of publication: 30.05.2003.
Noteridae 33
Amphizoidae 35
Hygrobiidae 35
Dytiscidae 35
Rhysodidae 78
Published with fmancial support from Carabidae 79
References 575
Index to families, subfamilies, tribes, and subtribes 797
to genus-group names 798
AAGE V JENSENS FONDE
INTRODUCTION 1. Old records "Arabia" or "China" may pertain to any states on the Arabian Peninsula and Jordan, or to any part
of the People's Republic of China, respectively.
Structured knowledge reąuires categorisation of observation and phenomena. Stractured knowledge of life, as it 2. Recent field work in the Himalaya and in the mountains of mainland China and Taiwan, resulting in the study
evolved and is present on our planet, reąuires cataloguing. Catalogues summarize the work in systematics, they of new collections from these areas, provides evidence of gradual faunal transitions in Coleoptera, as well as
enable assessment of faunal diversity and classification, and provide information on past taxonomic acts. They are in other insect groups. While subtropical climate with dominant Oriental taxa prevails at Iow elevations,
also essential in achieving a universal language in taxonomy that assures unambiguous transfer of biological significant faunal changes are found already at elevations of 1500 m, and almost "pure Palaearctic" taxa are
information. Erroneous information due to incorrect use of taxonomic names has an impact in many fields, including present at elevations of 2500 m and above. The faunal composition varies along the altitude gradient and,
genetics, physiology, ecology, plant protection, naturę conservation, and legislation. naturally, from one group of insects to the other. Thus, it appears illusory to draw simple biogeographical
The ever increasing number of recognise taxa sińce the tenth edition of Linne's Systema Naturae and the frontiers in any of these parts of the world.
scattered sources of information, were likely the main motivation for compilation of catalogues in the past. Sińce 3. The Catalogue includes morę information this way. This is an obvious advantage in the absence of modern
the middle of the 19* century, numerous catalogues were published, usually covering some parts of the world, that catalogues covering the Afrotropical and Oriental Regions.
were checklists rather than works that we would ąualify today as fuli catalogues. Sińce catalogues are static by their
naturę, while knowledge is dynamie, the need for new, complete catalogues remains. The first, for that time fuli, The Catalogue will be published in a number of volumes. Each volume will be "independent", having its own
catalogue of world Coleoptera was Gemminger & HarokTs Catalogus Coleopterorum issued from 1868 to 1874. It Reference section and Index. The first volume contains the Archostemata, Myxophaga and Adephaga. The following
was followed by Junk & Schenkling' s (editors) Coleopterorum Catalogus, published in separate parts from 1910 to volumes will treat the Polyphaga, grouping the superfamilies with respect to the adopted classification, and the size
1940. At present, even though it is desperately obsolete, the Coleopterorum Catalogus remains the only source of of the groups included.
comprehensive taxonomic information on a world basis for many families of beetles. The only Catalogue covering The authors are responsible for the information provided. A detailed peer review of the parts was not considered
all groups of Coleoptera oceurring in the Palaearctic region is Winkler's 1924 - 1932 Catalogus Coleopterorum to be reasonable, because it would undoubtedly reąuire an amount of work comparable to that of the authors (but see
regionis palaearcticae. The Junk & Schenkling and Winkler catalogues are of different natures, the former one under Acknowledgments).
approaching the definition of a "fuli" catalogue, while the latter is rather a check list. They both share two serious
omissions. The primary sources of taxonomic acts were not consistently checked, and the information on distribution TAXONOMIC INFORMATION
was not detailed enough to satisfy the needs. In addition, many rules subseąuently adopted by the International Code
ofZoologicalNomenclature were obviously not respected. Conseąuently, the language applied in current taxonomic The present Catalogue includes all available names, both valid and invalid, of extant beetle taxa described before
literaturę is inadeąuate in many respects. In particular, unavailable names are regularly used as available and January 1, 2000 and known to oceur in the Palaearctic Region, as it was defined above. The higher classification,
sometimes even regarded as valid, various orthographies are used for one and the same taxon, and references to from suborder down to subfamily, is based on the work of Lawrence & Newton (1995): Families and subfamilies
taxonomic acts are inexact to a high degree for papers published during the 20* century. of Coleoptera (with selectedgenera, notes, references and data onfamily-group names). However, the classification
Undoubtedly, over the years morę students have studied the Palaearctic Coleoptera than those of other parts of proposed in this work is not taken as dogma, and changes are accepted when considered well founded. All taxa below
the World, and morę often than representatives of most other orders of insects. It is therefore remarkable that an subfamily level are arranged alphabetically within the higher taxon and the synonyms follow the respective valid
overview of the present knowledge of the Palaearctic Coleoptera is badly trailing that of, e.g., the Nearctic name alphabetically.
Coleoptera, or Palaearctic Diptera. It may sound like an anachronism, but systematic studies of many groups of Extinct taxa, names rejected by the ICZN, unjustified emendations, misspellings, misidentifications and other
Palaearctic Coleoptera have to start with searches for data scattered throughout the literaturę of the past 250 years. nomina nuda are not included in the Catalogue. Similarly, all infrasubspecific names, such as those established as
A fuli catalogue gives information concerning all published names within a group, their classification, taxonomic "morpha", "natio", or "race", "subvariety" and "aberration", or proposed as yariety and form of a subspecies or
history, means of identification, references to taxonomic acts, type materiał, detailed distribution, habitat, natural another variety, and names published, e.g., as A-us b-us c-us, but specified in the text that they are actually proposed
history, and so on. Ideally, the present Catalogue would include such data. However, considering the number of taxa for a "natio" or "race", etc, are not considered subspecific and are therefore excluded from the Catalogue. Names
and present day competent taxonomists, and the difficulties they have in accessing the sources, a fuli catalogue of proposed as varieties and forms before 1961 are included, except for those established in Letzner's (1847-1852)
all Palaearctic beetles would not be completed within a reasonable period of time. Systematische Beschreibung der Laufkafer Schlesiens and DallaTorre's (1877) Synopsis der Insecten
Having this in mind, the aim of this Catalogue is to provide a tool that addresses the most urgent needs: 1) a list Oberósterreichs. Letzner's work unambiguously reveals that the names were proposed as infrasubspecific entities
of available names, both valid and uwalid, of taxa oceurring in the Palaearctic Region, in their verified original (see ICZN 1999, Art. 45.6.4). The Editors accepted the opinion of a majority of the authors of Carabidae, who
orthography and with the correct publication dates, 2) a complete list of verified references to primary descriptions, consider the new varieties and forms proposed by Dalia Torre (1877) as infrasubspecific. This results in the exclusion
and 3) informative distributions of the species and subspecies. Thus, the Catalogue is expected to respond to of many names that were regularly listed in previous publications. In Adephaga, in particular, it reduced the
ąuestions related to biodiversity, and to inerease the badly needed stability of the nomenclature. enormous list of synonymous names in the taxonomically complex and well-studied genus Carabus.
This Catalogue is a collective work involving many authors. For its realisation, it was necessary to achieve a The International Code ofZoological Nomenclature cannot prevent the appearance of superfluous new names,
reasonable consensus in very diverse fields. The Editors appreciated the understanding of the contributing authors established by descriptions of local populations as subspecies, however the lists of already available names in
of the need for consistency. The most controversial points concern the spelling of the names, and the delimitation synonymy will hopefully help to reduce the popularity of such actions.
of the Palaearctic Region and its main subdivisions. The currently valid names of the family-group taxa include the name of the author and the year of the
The first point is purely formal. The alternatives adopted by the Editors and presented below, are those that found publication. Their synonyms are not listed. Some of the currently used names of the taxa of this group have older
most support during the discussions with colleagues. The Editors are aware of the difficulty in satisfying all opinions, synonyms. One of them is the name Paelobiidae Erichson, 1837 (Pelobiidae sensu auct), based on Paelobius
and appeal to the understanding of the user. Schoenherr, 1808, which has priority over the generally accepted name Hygrobiidae Regimbart, 1878. According
The second point appears morę pertinent. The Palaearctic Region, as it is defined for the purpose of this to the ICZN (Art.40.2) the name Hygrobiidae is to be maintained.
Catalogue, includes some areas that are usually considered to belong to the Afrotropical, Oriental, and Pacific The names of the genus-group taxa are given with the name of the author, and the year and page of publication.
Regions respectively, i.e. the south of the Arabian Peninsula and Socotra, all of Pakistan, the Himalayanpart of India, The page given is the page where the name and the actual description of the taxon is printed. The type species of all
including Arunachal Pradesh, Nepal and Bhutan, the south of China, and the Pacific Islands of Japan. The main genus-group names are given in their original combination. If the type species is currently regarded as a junior
reasons for the inclusion of these areas are as follows: synonym, the valid senior synonym is given in brackets in its original combination.
The names of the species-group taxa are given with the name of the author, and the year and page of publication. specimens in some large museums. The degree of identification reliability and of the records derived from the
The page given is the page where the name and the actual description of the taxon is printed. For species-group taxa identification, is a function of faunal diversity and ąuality of systematic revisions. Thus, the reliability in generał
subseąuently transferred to another genus, the name of the original genus is given in parentheses, following the page increases from south to north and from poorly studied groups to "popular" groups. At present, a number of modern
catalogues or check-lists, covering the beetle faunas of many European and some extra-European countries or
of publication.
archipelagos, are available. The use of data contained in these and other faunistic works is left to the discretion of
Some authors (e.g., V. Apfelbeck, H. John) published the same description twice, or even morę times, in separate
the authors, who may also add unpublished information available to them. The second-level geographic information
papers. Such publication produces, de facto, primary homonyms and objective synonyms. The first publication in
is not necessarily exhaustive, it should rather be considered as a base for future faunal research.
such cases is referred to as indicated above, followed by the mark = , the year and first page of the subseąuent
The third-level geographic information concerns species and subspecies with restricted distribution. Taxa of this
description/s in sąuare brackets. This is particularly important for taxa that are erroneously associated with their
category may be strict endemites, or taxa comparatively widely distributed in one area but restricted in another area.
junior description.
For example, the distributional record of a species widely distributed in North Africa with isolated occurrence on
The following symbols, all given in sąuare brackets following the page of publication, or the original
Pantelleria would appear as follows: E: IT (Pantelleria) N: AG MO TU. The third-level information is facultative.
combination when applicable, are used for taxonomic information: HN for homonyms, RN for replacement names,
It is given in parentheses after the respective second-level symbol. The official language of the respective state is
ND for nomina dubia, NO for nomina oblita, NP for nomina protecta, DA for doubtful assignment, and EA for
used for records in languages using the Latin alphabet, or it is transliterated from the Cyrillic alphabet. Records in
erroneous assignment.
languages using non-Latin or Cyrillic characters (e.g. Chinese or Japanese pictographs) are translated into English,
Nomina dubia and taxa considered incertae sedis are listed separately at the end of the nearest applicable taxon.
and the translated geographical terms are spelled as closely as possible to those used in the Times Atlas®, or in other
Taxonomic and nomenclarorial acts published after January 1,2000 are considered only when they concern taxa
well-known sources. Detailed geographical information may refer to natural geographical features such as islands,
described before that datę.
mountains, lakes, yalleys, caves, or to administrative entities, such as districts.
DISTRIBUTIONAL INFORMATION The extralimital distribution of some Palaearctic species is indicated by three letter symbols in bold,
cosmopolitan species by the symbol COS, all located at the end of the respective geographical information (see Table
The limits of the Palaearctic region, as those of other biogeographical regions, are arbitrarily defined (Map 1). 1). Introductions are indicated by the letter "i" (e.g., Ei: GB). The extralimital regions for the needs of the Catalogue
For practical reasons, the boundaries of the Palaearctic Region, as they were established for the Catalogue (see are defined as follows (see Map 1): Nearctic (NAR): north of Mexico; Neotropical (NTR): south of the United
above), usually follow national boundaries. The region includes Europę, Africa north of the Sahara, and Asia except States; Afrotropical (AFR): south of the North African states included in the Palaearctic Region; Oriental (ORR):
for the part that is arbitrarily defined as belonging to the Oriental Region. areas south of mainland China and Taiwan, areas south of the Himalaya in India, the Philippines, Malaysia and
For each species and subspecies an outline of its present distribution is given. Fossil records are not considered. Indonesia south to the Lydekker linę; Australian (AUS): south of the Lydekker linę, Pacific.
The information is given by means of symbols, presented at three levels.
The first level is the subdivision of the Palaearctic Region into three main parts, Europę (letter E, bold), North
Africa (letter N, bold) and Asia (letter A, bold).
Europę includes the Azores, Iceland and Turkey west of the Bosporus. The eastern boundaries are a matter of
controversy. In the Catalogue, Europę includes Russia west of the main ridge of the Ural mountains, the Permsk
Oblast, Bashkortoskan Republic and Orenburskaya Oblast, and the smali part of Kazakhstan west of the Ural River.
It includes the Caucasian republics of Georgia, Armenia and Azerbaijan. The south-eastern boundaries are the
political boundaries of the Asian part of Turkey, Iran, Kazakhstan, and the Caspian and Black seas.
North Africa includes Morocco (incl. Western Sahara), Algeria, Tunisia, Libya and Egypt west of the Suez Canal,
and the Canary and Madeira islands.
Asia includes Sinai and the Arabian Peninsula (including Socotra), Turkey east of the Bosporus, the Middle East and
Central Asian countries, Russia east of the main ridge of the Ural mountains, Korea, Japan (including Ryukyu Rettó
and the Japanese Pacific Islands), the entire People's Republic of China, Taiwan, Bhutan, Nepal, North India along
the base of the Himalaya (Arunachal Pradesh, Uttar Pradesh, Himachal Pradesh), Jammu & Kashmir and all of
Pakistan. Thus, India is the only state for which the strict political boundaries are not respected. Large parts of Uttar
Pradesh south of Nepal are overpopulated plains. Information on Coleoptera from this North Indian state is based
almost exclusively on its Himalayan districts lying west of Nepal, which are in fact a transitional zone.
The second level of the geographic information is provided by two-letter symbols for countries, major areas of
Russia and North Indian states, and by three-letter symbols for provinces of mainland China and for Taiwan (see
Table 1, Map 3).
The symbols are arranged in alphabetical order within the first-level subdivision. Some, usually older,
distributional records cannot be accommodated within the structure of the Catalogue (e.g., Arabia, Caucasus, North
India, Siberia). Such information is given in ąuotation marks (e.g., "Caucasus") behind the last symbol of the
respective first-level symbol. Russia is subdivided into six major sub-regions (Map 2), each of which has its own
two-letter symbol. These, as well as the symbol RU for Russia, are used only when morę detailed information is not
available. Similarly, the symbol CH for the People's Republic of China, as well as the seven two letter symbols for
China's major regions are used only in the absence of morę detailed geographical information.
In generał, the published distributional information is based on both identified materiał in collections and on
Map. 1: The limits of the Palaearctic Region as defined for the purpose of this Catalogue.
published records, scattered in an enormous number of taxonomic and faunistic papers that are virtually impossible
to review in their entirety. Revisions of collections reveal a high proportion of misidentifications, attaining 40% of
Table 1: GEOGRAPHICAL SYMBOLS
E Europę N North Africa
AB Azerbaijan AG Algeria
AL Albania CI Canary Islands
AN Andorra EG Egypt
AR Armenia LB Libya
AU Austria MO Morocco (incl. Western Sahara)
AZ Azores MR Madeira Archipelago
BE Belgium TU Tunisia
BH Bośnia Herzegovina
BU Bułgaria
BY Byelorussia
CR Croatia A Asia
CT Russia: Central European Territory AE Arab Emirates
CZ Czech Republic AF Afghanistan
DE Denmark AP Arunachal Pradesh
EN Estonia BA Bahrain
FA Faeroe Islands BT Bhutan
FI Finland CE China: Central Territory
FR France (incl. Corsica, Monaco) CH China
GB Great Britain (incl. Channel Islands) CY Cyprus
GE Germany ES Russia: East Siberia
GG Georgia FE Russia: Far East
GR Greece (incl. Crete) HP Himachal Pradesh
Map. 2: Subdivisions of Russia.
HU Hungary IN Iran
IC Iceland IQ Iraq
IR Ireland IS Israel
IT Italy (incl. Sardinia, Sicily, San Marino) JA Japan
KZ Kazakhstan JO Jordan
LA Latvia KA Kashmir (India)
LS Liechtenstein KI Kyrgyzstan
LT Lithuania KU Kuwait
LU Luxembourg KZ Kazakhstan
MA Malta LE Lebanon
MC Macedonia MG Mongolia
MD Moldavia NP Nepal
NL The Netherlands NE China: Northeast Territory
NR Norway NC North Korea
NT Russia: North European Territory NO China: Northern Territory
PL Poland NW China: Northwest Territory
PT Portugal OM Oman
RO Romania PA Pakistan
RU Russia QA Qatar (incl. United Arab Emirates)
SK Slovakia RU Russia
SL Slovenia SA Saudi Arabia
SP Spain (incl. Gibraltar) SC South Korea
SR Svalbard (Spitzbergen) SD Sikkim, Darjeeling (India)
ST Russia: South European Territory) SE China: Southeastern Territory (incl
SV Sweden Macao, Hongkong)
SZ Switzerland SI Egypt: Sinai
TR Turkey SW China: Southwestern Territory
UK Ukrainę SY Syria
YU Yugoslayia (Serbia, Montenegro) TD Tadzhikistan
Map. 3: Provinces of the Peoples' Republic of China.
10 11
WORLD ZOOGEOGRAPHICAL REGIONS: BIBLIOGRAPHIC INFORMATION
A Asia (continued)
References are given to, and only to, primary sources of the genus-group and species-group names included in the
TM Turkmenistan
TR Turkey AFR Afrotropical Region Catalogue. No bibliographical reference is given to family-group names, to type-species, to secondary taxonomie
UP Uttar Pradesh (India) AUR Australian Region acts, or to sources of any other information. Names of all authors, if applicable, are given for each reference. An
UZ Uzbekistan NAR Nearctic Region effort was put forth to unify the spellings of the names. For names that appeared in publications in two or more
WP China: Western Plateau NTR Neotropical Region different orthographies, the one used in the Catalogue is that which was considered to be correct after thorough
WS Russia: west Siberia ORR Oriental Region verification (e.g., the spelling Josef Müller for the author who also published as Giuseppe Müller). In such cases the
alternative spelling follows the original one in brackets. The original name is used for authors that subsequently
YE Yemen (incl. Socotra)
adopted additional names, or honorary epithets (e.g., A. Semenov, and not A. Semenov-Tian-Shanskyi). Initials are
given for all authors, even if they are missing in the original publication. Names published in two or more alternative
spellings, that seem to be equally correct, are spelled following Schmitt, Hiibner & Gaedike (1998): Nomina
Auctorum. AuflösungvonAbkürzungen taxonomischerAutoren-Namen. NovaSupplementaEntomologica 11:3-189.
CHINA: PROVINCES, AUTONOMOUS
An exception is made for F. L. Castelnau de Laporte, who is referred to as Laporte in recent major works. Therefore,
REGIONS OR MUNICIPALITIES, AND
the name Laporte is arbitrarily used in the Catalogue. Names in Cyrillic are transliterated as for any other Cyrillic
TAIWAN
text, unless an alternative spelling was preferred by the author, or the alternative name is generally used in the
literature (e.g., Jakobson, Tschitschérine). It should be noted that lowercase prepositions such as "de", "van" and
AHN Anhui (Anhwei)
"von" are not included with the authors' names in the species catalogue.
BEI Beijing (Peking or Peiping)
Titles of references are given in full, i.e. not abbreviated, with subtitles, as originally published. Text written in
FUJ Fujian (Fukien)
Cyrillic is transliterated. Titles in languages using non-Latin or non-Cyrillic characters are translated into English
GAN Gansu (Kansu)
and are presented in square brackets, followed by the indication of the original language in parentheses.
GUA Guandong (Kwantung)
Titles of periodicals are given in full. Series numbers are given in parentheses, and always precede the boldface
GUI Guizhou (Kweichow)
volume number. The numbers of separate issues are only provided when they are paginated separately, in which case
GUX Guanxi (Kwangsi)
they are in parentheses following the volume number. Unnumbered pages are indicated in square brackets. The titles
HAI Hainan
of some periodicals vary over the years. For these periodicals the titles are given as they appeared for the respective
HEB Hebei (Hopeh)
items referred to (e.g., Coleopterologische Rundschau I Koleopterologische Rundschau). To the contrary, thejournal
HEI Heilongjiang (Heilungkiang)
Entomologische Blatter had several subtitles during its existence, the last of them being Entomologische Blatter für
HEN Henan (Honana)
Biologie und Systematik der Kafer. Since there is no danger of confusion with any other periodical, the journal is
HKG Hongkong
cited simply as Entomologische Blatter. If the year of the volume differs from the actual year of publication, the
HUB Hubei (Hupeh)
former follows the volume number and is given is square brackets. If periodicals have no volume number, the year
HUN Hunan
of the volume is given.
JIA Jiangsu (Kiangsu)
The Bulletin de la Société Entomologique de France is one of the more important periodicals in the field of
JIL Jilin (Kirin)
systematics. The title changed almost continuously for more than 60 years, appearing usually as Bulletin des seances,
JIX Jiangxi (Kiangsi)
Bulletin trimestriel or Bulletin entomologique. In addition, the first numbered volume, 37, was published 100 years
LIA Liaoning
after the creation of the Annales and the Bulletin of the French Entomological Society in 1832. To avoid confusion,
MAC Macao
this periodical is referred to as Bulletin de la Société Entomologique de France, and its volumes from 1832 to 1931
NIN Ningxia (Ningsia)
are identified by the year in which they were published. From the year 1932, the volume number is given. The same
NMO Nei Mongol (Inner Mongolia)
format is used for the Bulletin de la Société Entomologique de Belgique.
QIN Qinghai (Tsinghai)
For ease of access, references of the same year were placed in order based on the first page number of the paper,
SCH Sichuan (Szechwan)
with Roman numerals being placed last. Books were placed first for a given year, unless a page range was cited, in
SHA Shaanxi (Shensi)
which case the first-page rule was followed. Papers of the same year with the same first page numbers were placed
SHG Shanghai
in order based on their last page number. Thus the references are not arranged chronologically within each year.
SHN Shandong (Shantung)
A few items in the "References" are unfortunately given in a variably incomplete form. These represent
SHX Shanxi (Shansi)
references which, even after the combined effort of many workers, could not be completed for various reasons, and
TAI Taiwan (Formosa)
their completion would unduly postpone the publication of the Catalogue.
TIA Tianjin (Tsiensin)
XIN Xinjiang (Sinkiang)
ACKNOWLEDGEMENTS
XIZ Xizang (Tibet)
The generally recognized need for a modern catalogue of the Palaearctic Coleoptera was discussed during a social
YUN Yunnan
meeting of several coleopterists in Prague in October 1996. At this meeting Jan Farkač and David Král came forward
ZHE Zhejiang (Chekiang)
with the idea to establish an editorial committee for such a catalogue in Prague that would organize a collective of
authors for the project. The idea was received quite positively by all the coleopterists present, and shortly afterwards
the project was initiated, with the additional help of Karel Hůrka and Jan Růžička. After many initial difficulties and
personnel changes, the project was eventually brought underway and this volume of the Catalogue is the first result
of the combined efforts of many dedicated coleopterists.
12 13
Compilation of an accurate catalogue requires, in addition to almost endless patience and obsession with detail, COORDINATORS OF THIS VOLUME.
access to an enormous quantity of primary taxonomie literature some of which dates back almost 250 years. The Yves Bousquet, Biodiversity: Systematic Entomology, Agriculture and Agri-Food Canada, Ottawa, Canada
libraries, institutional and private, are therefore the obvious sources for acquiring data for a catalogue. Realizing this, Anthony Davies, Biodiversity: Systematic Entomology, Agriculture and Agri-Food Canada, Ottawa, Canada
the Editors (in most cases on behalf of many contributing authors) acknowledge the invaluable assistance of library Karel Hůrka, Department of Zoology, Charles University, Praha, Czech Republic (assisted by Jan Farkač, Czech
staff and/or colleagues with access to these libraries of all librariers contacted. The list of the libraries is long, but Agricultural University, Praha, Czech Republic)
perhaps a few most important should be mentioned: Agriculture and Agri-Food Canada (Entomology Library), Aleš Smetana, Biodiversity: Systematic Entomology, Agriculture and Agri-Food Canada, Ottawa, Canada
Ottawa, Canada (Mrs. Patricia Madore and Mr. Steve Gamman); American Museum of Natural History, New York,
USA (Lee Herman); Museum d'histoire naturelle, Geneva, Switzerland (Mrs. Anne-Marie Deuss and Mrs. Christelle AUTHORS
Mougin); Naturkunde Museum, Eberswalde, Germany (Reinhard Gaedike). suborder Archostemata Kolbe, 1908
The Editors thank Reinhard Gaedike (Naturkunde Museum, Eberswalde, Germany) who generously made family CROWSONIELLIDAE Iablokoff-Khnzorian, 1983 I. LÖBL
available unpublished bibliographical information. family MICROMALTHIDAE Barber, 1913 I. LÖBL
In addition, the Editors thank many individuals, who in various ways contributed to the completion of this family CUPEDIDAE Laporte, 1836 I. LÖBL
Catalogue: Verena Andres, Berne; M. Barclay, Oxford; Paolo Bonavita, Roma; Daniel Burckhardt, Basel; Holger family SIKHOTEALINIIDAE Lafer, 1996 I. LÖBL
Dathe, Eberswalde; Thierry Deuve, Paris; O. Dumann, Oxford; Claude Jeanne, Langon; Andreas Kaupp, Basel;
Sergej Kazantzev, Moscow; Serge Laplante, Ottawa; Otto Merkl, Budapest; Bernhard Merz, Geneva; Seiji Morita, suborder Myxophaga Crowson, 1955
Tokyo; EugenNitzu, Bucharest; t Ilja Okáli, Bratislava; Kathrin Ruckstuhl, Cambridge; Harald Schillhammer, Wien; family TORRIDINCOLIDAE Steffan, 1964 I. LÖBL
Joachim Schmidt, Rostock; Sharon Shute, London; Alyosha Solodovnikov, Chicago; Luca Toledano, Verona; Hans family HYDROSCAPHIDAE LeConte, 1874 I. LÖBL
Turin, Renkum; Lothar Zerche, Eberswalde. family SPHAERIUSIDAE Erichson, 1845 I. LÖBL
Due to the incompleteness of some of the submitted manuscripts dealing with Carabidae, it was necessary to
establish a "reviewing" group of three taxonomists for this volume, the members of which were Yves Bousquet, suborder Adephaga Clairville, 1806
Anthony Davies and Aleš Smetana, all in Ottawa. After the work of this reviewing group was completed, the family GYRTNIDAE Latreille, 1810 P. MAZZOLDI
amended manuscripts were sent back to the authors for final approval. The part dealing with the tribes Cicindelini family HALIPLIDAE Kirby, 1837 B.J. vanVONDEL
and Carabini posed particular problems. Even after a combined effort of several authors of these sections, some family TRACHYPACHIDAE C.G. Thomson, 1857 K. HŮRKA
problems still remain, particularly in terms of the availability and authorship of some names that are generally family NOTERIDAE C.G. Thomson, I860 A.N. NILSSON
recognized as subspecific at present, but were originally established as infrasubspecific. The names of this category family AMPHIZOIDAE LeConte, 1853 M. JACH
we are aware of are as follows: Cicindela sylvatica rubescens Jeanne, 1967b; Cicindela germanica michaelensis family HYGROBIIDAE Régimbart, 1878 M. JÂCH
VidalyLópez, 1916; CarabusmonticolaroccaeBorn, 1923; CarabusbrandtigansuanusBreuning,1932; Carabus family DYTISCIDAE Leach, 1815 A.N. NILSSON
violaceus HguriensisBremung, 1932; Carabus odoratus kamtschatensisBreuning, 1942; Carabus odoratus melleus family RHYSODIDAE Laporte, 1840 R.T. BELL
Lapouge, 1909; Carabus odoratus septentrionalis Breuning, 1932; Carabus coriaceus joniensis Breuning, 1932; family CARABIDAE Latreille, 1802
Carabus macrogonus akkusensis Breuning, 1954; Carabus varians rugatus Breuning, 1934; Carabus cancellatus subfamily Nebriinae Laporte, 1834
mimus Kolbe, 1913; Carabus yunnanus guerryanus Breuning, 1961. The solution of these problems would require tribe Nebriini Laporte, 1834 J. FARKAČ & M. JANATA
an extensive and time-consuming study of a large number of secondary references, a task that would significantly tribe Notiophilini Motschulsky, 1850 Υ. BOUSQUET & A. BARŠEVSKIS
postpone the publication of the Catalogue. tribe Opisthiini Dupuis, 1912 Y. BOUSQUET
tribe Pelophilini Kavanaugh, 1996 J. FARKAČ
The Editors subfamily Cicindinae Banninger, 1927 P. NAGEL
Ivan Löbl Aleš Smetana subfamily Loricerinae Bonelli, 1810 Y. BOUSQUET
Museum d'histoire naturelle Agriculture and Agri-Food Canada subfamily Cicindelinae Latreille, 1802 A.V. PUCHKOV & A.V. MATALIN
subfamily Carabinae Latreille, 1802
Geneva, Switzerland Ottawa, Canada
tribe Carabini Latreille, 1802 Y. BOUSQUET, B. BŘEZINA, A. DAVIES,
J. FARKAČ & A. SMETANA
tribe Cychrini Laporte, 1834 M. HÂCKEL
subfamily Elaphrinae Erichson, 1837 Η. GOULET
subfamily Omophroninae Bonelli, 1810 Κ. HŮRKA
subfamily Paussinae Latreille, 1807 P. NAGEL
subfamily Brachininae Bonelli, 1810 J. HRDLIČKA
subfamily Siagoninae Bonelli, 1813 M. BAEHR
subfamily Scaritinae Bonelli, 1810 M. BALKENOHL
subfamily Melaeninae Alluaud, 1934 Υ. BOUSQUET
subfamily Broscinae Hope, 1838 Υ. BOUSQUET
subfamily Apotominae LeConte, 1853 D.W. WRASE
subfamily Trechinae Bonelli, 1810
tribe Anillini Jeannel, 1937 J.P. ZABALLOS
tribe Bembidiini Stephens, 1827
subtribe Bembidiina Stephens, 1827
W.A. MARGGI, C. HUBER,
G. MÜLLER-MOTZFELD
& M. HARTMANN
14 15
subtribe Tachyina Motschulsky, 1862 T. KOPECKÝ MAILING AND E-MAIL ADDRESSES OF THE AUTHORS
tribe Patrobini Chaudoir, 1871 A. ZAMOTAILOV Antonio ANDÚJAR, Departamento de Biología Animal, Universidad de Murcia, E-30 071 Murcia, Spain.
tribe Pogonini Laporte, 1834 Y. BOUSQUET E-mail: [email protected]
tribe Trechini Bonelli, 1810 P. MORAVEC, S.-I. UÉNO Martin BAEHR, Zoologische Staatssammlung, Münchenhausenstrasse 21, D-81247 München, Germany.
& LA. BELOUSOV E-mail: [email protected]
tribe Zolini Sharp, 1886 Y. BOUSQUET Michael BALKENOHL, Bertholdstrasse 2, D-79211 Denzlingen, Germany.
subfamily Psydrinae LeConte, 1853 K. HŮRKA E-mail: [email protected]
subfamily Harpalinae Bonelli, 1810 Arvîds BARŠEVSKIS, Baltic Institute of Coleopterology and Department of Biology, University of Daugavpils,
tribe Abacetini Chaudoir, 1872 Υ. BOUSQUET Vienîbas 13- 229, Daugavpils, LV- 5401, Latvia. E-mail: [email protected]
tribe Anthiini Bonelli, 1813 Υ. BOUSQUET Ross T. BELL, Department of Zoology, Marsh Life Science Building, The University of Vermont, Burlington,
tribe Chlaeniini Brullé, 1834 Ε. KIRSCHENHOFER VT 05405, USA. E-mail: [email protected]
tribe Corsyrini Ganglbauer, 1891 Υ. BOUSQUET Igor BELOUSOV, All-Russian Institute of Plant Protection, Podbelskogo 3, St.-Petersburg - Pushkin, 189620
tribe Cyclosomini Laporte, 1834 Υ. BOUSQUET Russia. E-mail: [email protected]
tribe Dryptini Bonelli, 1810 Μ. BAEHR Yves BOUSQUET, Agriculture and Agri-Food Canada, Central Experimental Farm, K.W. Neatby Bldg.,
tribe Galeritini LeConte, 1853 Μ. BAEHR Ottawa, ON Kİ A 0C6, Canada. E-mail: [email protected]
tribe Graphipterini Latreille, 1802 Κ. HŮRKA Boleslav BŘEZINA, U Pentlovky 9/465, CZ- 181 00 Praha 8, Czech Republic. E-mail: [email protected]
tribe Harpalini Bonelli, 1810 Achile CASALE, Istituto di Zoologia dell'Universitâ, Via Muzoni 25,1-07100 Sassari, Italy.
subtribe Amblystomina Fauvel, 1889 Β. JAEGER & D.W. WRASE E-mail: [email protected]
subtribe Anisodactylina Lacordaire, 1854 Ν. İTO Anthony DA VIES, Agriculture and Agri-Food Canada, Central Experimental Farm, K.W. Neatby Bldg., Ottawa,
subtribe Ditomina Bonelli, 1810 D.W. WRASE ON K1A 0C6, Canada. E-mail: [email protected]
subtribe Harpalina Bonelli, 1810 Β.Μ. KATAEV, D.W. WRASE & Ν. ΙΤΟ Jan FARKAČ, Department of Forest Protection, Faculty of Forestry, Czech Agricultural University, CZ-165 21,
(selenophorines) Praha 6- Suchdol, Czech Republic. E-mail: [email protected]
subtribe Stenolophina Kirby, 1837 Β. JAEGER & B.M. KATAEV Henri GOULET, Agriculture and Agri-Food Canada, ECORC, Central Experimental Farm, K.W. Neatby Bldg.,
tribe Helluonini Hope, 1838 Κ. HŮRKA Ottawa, ON K1A 0C6, Canada. E -mail: [email protected]
tribe Hexagoniini G. Horn, 1881 Κ. HŮRKA Martin HÂCKEL, Za zahradami 1431. CZ-253 01 Hostivice, Czech Republic. E-mail: [email protected]
tribe Lachnophorini LeConte, 1853 Υ. BOUSQUET Matthias HARTMANN, Naturkundemuseum Erfurt, Grosse Arche 14, D- 99084 Erfurt, Germany.
tribe Lebiini Bonelli, 1810 Ι. KABAK E-mail: [email protected]
tribe Licinini Bonelli, 1810 M. BAEHR Fritz HIEKE, Zoologisches Museum, Invalidenstrasse 43, Berlin N4, Germany.
tribe Loxandrini Bousquet & Larochelle, 1993 Υ. BOUSQUET E-mail: [email protected]
tribe Melanchitonini Jeannel, 1948 Υ. BOUSQUET Oldřich HOVORKA, Insect Chemical Ecology Unit, Institute of Organic Chemistry and Biochemistry, Academy
tribe Morionini Brullé, 1835 Υ. BOUSQUET of Science of the Czech Republic, Flemingovo náměstí 2, CZ-166 10 Praha 6, Czech Republic.
tribe Odacanthini Laporte, 1834 Υ. BOUSQUET & Ν. İTO Jan HRDLIČKA, Chopikova 22 120 00 Prague 2, Czech Republic.
tribe Oodini LaFerté-Sénectěre, 1851 Υ. BOUSQUET Charles HUBER, Naturhistorişches Museum, Bernastrasse 15, CH- 3005, Bern, Switzerland.
tribe Omphreini Ganglbauer, 1891 Υ. BOUSQUET E-mail: [email protected]
tribe Orthogoniini Schaum, 1857 Υ. BOUSQUET Karel HŮRJKA, Department of Zoology.Charles University, Vim čná 7, CZ 128 44, Praha 2, Czech Republic.
tribe Panagaeini Bonelli, 1810 Μ. BAEHR E-mail: [email protected]
tribe Pentagonicini Bates, 1873 Υ. BOUSQUET Noboru ITO, 1-7-18, Higashiuneno, Kawanishi City, Hyögo Pref. ,666-0117 Japan.
tribe Perigonini G.H. Horn, 1881 Υ. BOUSQUET E-mail: [email protected]
tribe Platynini Bonelli, 1810 Υ. BOUSQUET Manfred JÁCH, Naturhistorişches Museum, Burgring 7, A-1014, Wien, Austria.
tribe Pterostichini Bonelli, 1810 Υ. BOUSQUET E-mail: [email protected]
tribe Sphodrini Laporte, 1834 Bernd JAEGER, Zoologisches Museum, Invalidenstrasse 43, Berlin N4, Germany.
subtribe Atranopsina Baehr, 1982 Ο. HOVORKA & R. SCIAKY E-mail: bernd.j [email protected]
subtribe Calathina Laporte, 1834 O. HOVORKA & R. SCIAKY Miroslav JANATA, V Hůrkách 2141, CZ-158 00 Praha 5, Czech Republic. E-mail: [email protected]
subtribe Dolichina Audouin & Brullé, 1834 O. HOVORKA & R. SCIAKY Ilia KABAK, Zoological Institute, Russian Academy of Sciences, Universitetskaya naberezhnaya 1, 199 034
subtribe Pristosiina Lindroth, 1956 O. HOVORKA & R. SCIAKY St.Petersburg, Russia. E-mail: [email protected] or [email protected]
subtribe Sphodrina Laporte, 1834 A. CASALE Boris M. KATAEV, Zoological Institute, Russian Academy of Sciences, Universitetskaya naberezhnaya 1, 199
subtribe Synuchina Lindroth, 1956 O. HOVORKA & R. SCIAKY 034 St. Petersburg, Russia. E-mail: [email protected]
tribe Zabrini Bonelli, 1810 Erich KIRSCHENHOFER, Otto Eisner Gasse 10-12, A-2380 Perchtoldsdorf, Austria.
subtribe Amarina Zimmermann, 1831 F. HIEKE E-mail: [email protected]
subtribe Zabrina Bonelli, 1810 J. SERRANO & A. ANDÚJAR Tomáš KOPECKÝ, Mánesova 712, CZ- 500 02 Hradec Králové, Czech Republic. E-mail: [email protected]
tribe Zuphiini Bonelli, 1810 M. BAEHR Ivan LÖBL, Museum d'histoire naturelle, Case postale 6434, VH-1211, Geneva, Switzerland.
E- mail: [email protected]
Werner MARGGI, Allmendstrasse 18, CH- 3601 Thun, Switzerland. E-mail: [email protected]
Andrei V. MATALIN, Zoology and Ecology Department, Moscow State Pedagogical University, KibaPchicha
6, Building 5, 129278, Moscow, Russia.
16
Paolo MAZZOLDI, Via Galilei 87, 25128 Brescia, Italy. E-mail: [email protected] Carabidae: Cychrini
Pavel MORAVEC, Tolstého 13, 41201 Litoměřice, Czech Republic. E-mail: moravec@ schkocr.cz M. Hâckel
Gerd MÜLLER-MOTZFELD, Zoologisches Museum und Institut, Ernst-Moritz-Arndt-Universitât, Bachstrasse New synonyms:
11/12, D-17489 Greifswald, Germany. E-mail: [email protected] Cychropsis wittmeriana Deuve, 1983, syn. nov. of C. sikkimensis sikkimensis (Fairmaire, 1901).
Peter NAGEL, Institut NLU - Biogeographie, Universitât Basel, St. Johanns-Vorstadt 10, CH- 4056 Basel, Cychrus inessae O. Berlov, 1997: 32, syn. nov. of C. aeneus aeneus Fischer von Waldheim, 1823).
Switzerland. E-mail: [email protected] Cychrus sharovae O. Berlov, 1997: 32, syn. nov. of C. aeneus aeneus Fischer von Waldheim, 1823.
Anders N. NILSSON, Department of Biology, BIG, Umeâ University, S-90187 Umeâ, Sweden. Cychrus signatus Faldermann, 1835: 13, syn. nov. of C. aeneus aeneus Fischer von Waldheim, 1823.
E-mail: [email protected] Cychrus volkovi O. Berlov, 1997: 28, syn. nov. of C. aeneus aeneus Fischer von Waldheim, 1823.
Alexandr PUCHKOV, Istitute of Zoology KAN, B. Chmelnitzky St., 15252030 Kiev, Ukraine. Cychrus belousovi O. Berlov, 1997: 35, syn. nov. of C. aeneus starcki Reitter, 1888.
E-mail: [email protected] Cychrus makarovi O. Berlov, 1997: 35, syn. nov. of C. aeneus starcki Reitter, 1888.
Riccardo SCIAKY, Via Fiamma 13, 20129 Milano, Italy. E-mail: [email protected] Cychrus milyaevi O. Berlov, 1997: 37, syn. nov. of C. aeneus starcki Reitter, 1888.
José SERRANO, Dept. Zoologia. Facultad Veterinaria, Campus de Espinardo, Aptdo. 4021, 30071 Murcia, Cychrus tillyi O. Berlov, 1997: 35, syn. nov. of C. aeneus starcki Reitter, 1888.
Spain. E-mail: [email protected] Cychrus zamotailovi O. Berlov, 1997: 35, syn. nov. of C. aeneus starcki Reitter, 1888.
Aleš SMETANA, Agriculture and Agri-Food Canada, Central Experimental Farm, K.W. Neatby Bldg., Ottawa, Cychrus costai Emery, 1872: 60, syn. nov. of C. caraboides (Linné, 1758).
ON K1A 0C6, Canada. E-mail: [email protected] Cychrus laticollis Jeanne, 1972: 80, syn. nov. C. caraboides (Linné, 1758).
Shun-Ichi UÉN0, National Science Museum (Natural History), 3-23-1 Hyakunin-cho, Shinjuku-ku, 169-0073 Cychrus zariquieyi Born, 1924: 9, syn. nov. of C. caraboides (Linné, 1758).
Tokyo, Japan.
Bernard J. van VONDEL, Roestuin 78, 3343 CV Herndrik-Ido-Ambacht, The Netherlands. Carabidae: Brachininae
E-mail: [email protected] J. Hrdlička
David W. WRASE, Dunkerstrasse 78, D-10437 Berlin, Germany. E-mail: [email protected] New synonym:
Juan P. ZABALLOS, Departemente de Biologia Animal I, Facultad de CC. Biologicas, Universidad Brachinus matchai Jedlička, 1924, syn. nov. of Brachinus baéticus Rambur, 1837.
Complutense de Madrid, E-28040 Madrid, Spain. E-mail: [email protected]
New replacement names:
Alexandr S. ZAMOTAILOV, Kuban State Agrarian University, Kalinin Str. 13, 350044, Krasnodar, Russia.
Brachinus krynickii, nom. nov. for B. nigricornis Krynicki, 1832 (nee Gebler, 1829).
E-mail: [email protected]
Brachinus motschulskyi, nom. nov. for B. marginipennis Motschulsky, 1864 (nee Laporte, 1835).
NEW NOMENCLATORIAL AND TAXONOMIC ACTS
Carabidae: Siagoninae
As the taxonomie history of many names is complex, these acts are considered new in respect to the latest
M. Baehr
relevant publications, in particular to the Catalogues of Lorenz, 1998 and Kryzhanovskij & al, 1995.
Change in rank:
Siagona oberleitneri Dejean, 1830= A europaea oberleitneri Dejean, 1830.
Carabidae: Nebriini
J. Farkač & J. Janata Carabidae: Paussinae
New replacement name:
P. Nagel
Leistus (Pogonophorus) schilenkovi Farkač, nom. nov. for Leistus (Pogonophorus) richteri Schilenkov, 1999 New synonyms:
[preoccupied by Leistus (Evanoleistus) richteri Farkač, 1995]. Cerapterus singalensis Kolbe, 1926, syn. nov. of C. latipes Svederas, 1788.
New synonyms:
Dimeroderus Westwood, 1847 (type species Melanospilus bensoni Westwood, 1845), syn. nov. of Merismoderus
Leistus {Leistus) kaszabi Horvatovich, 1972, syn. nov. of Leistus piceus piceus Fröhlich, 1799. Westwood, 1845 (type species Melanospilus bensoni Westwood, 1845).
Leistus (Leistus) pseudoalpicola Mařan, 1941, syn. nov. of Leistus piceus piceus Fröhlich, 1799. Paussomorphuspauliani Reichensperger, 1951, syn. nov. of Paussomorphus intuitivus (Kolbe, 1935) [described
Leistus (Leistus) gulickai Schweiger, 1970, syn. nov. of Leistus parvicollis Chaudoir, 1869.
as Paussus intuitivus Kolbe, 1935]. The species, as well as the entire genus, are not Palaearctic. However, the
Leistus (Leistus) rufofemoralis Mařan, 1941, syn. nov. οι Leistus parvicollis Chaudoir, 1869.
synonymy has to be presented to explain the absence of the name Trepopaussus in the list of the synonyms
Leistus (Pogonophorus) spinangulus Reitter, 1913, syn. nov. of i. spinibarbis expansus Putzeys, 1874.
of the genus Paussus.
Leistus (Sardoleistus) romanus K. Daniel, 1903, syn. nov. οι Leistus sardous Baudi di Selve, 1883.
Trepopaussus Kolbe, 1935 (type species Paussus intuitivus Kolbe, 1935), syn. nov. of Paussomorphus Raffray,
Spelaeonebria Peyerimhoff, 1911, syn. nov. oíAlpaeus Bonelli, 1810.
1885 (type species Paussus chevrolatii Westwood, 1852).
Nebria (Nebria) sevanensis Shilenkov, 1983, syn. nov. of Nebria araschinica Reitter, 1892.
Change in rank:
Nebria (Asionebrid) changaica Horvatovitch, 1973, syn. nov. of Nebria nivalis nivalis (Paykull, 1798).
Protopaussus basilewskyi Luna de Carvalho, 1967, is upgraded from subspecific to specific rank.
Changes in rank and new assignments:
Leistus (Pogonophorus) expansus Putzeys, 1874 = Leistus spinibarbis expansus Putzeys, 1874. Carabidae: Bembidiini: Bembidiina
Leistus terskeîensis Belousov & Kabak, 1992 (no subgeneric assignment) = Leistus (Pogonophorus) terskeiensis W. Marggi, C. Huber & G. Müller-Motzfeld
Belousov & Kabak, 1992. (genus Asaphidion: W. Marggi & M. Hartmann)
Nebria (Alpaeus) olympica Mařan, 1938 = Nebria aetolica olympica Mařan, 1938. New replacement names:
Nebria (Alpaeus) asturiensis Bruneau de Miré, 1964 = Nebriapunctatostriata asturiensis Bruneau de Miré, Bembidion (Bembidionetolitzkya) geniculatum claudejeannei Marggi, nom. nov. for Ocydromus (Daniela)
1964. geniculatus nevadensis Jeanne, 1974 [preoccupied by Bembidion nevadense Ulke, 1875].
Nebria (Alpaeus) vseteckai Mařan, 1938 = Nebria taygetana vseteckai Mařan, 1938. Bembidion (Ocyturanes) karokhense Marggi, nom. nov. for Peryphus bifasciatus Schuler, 1959 [preoccupied by
Nebria (Nebria) sitiens Antoine, 1936 = Nebria atlantica sitiens Antoine, 1936. Notaphus bifasciatus Stephens, 1828].
18 19