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Case 2851. Nacaduba Moore,[1881] (Insecta, Lepidoptera): proposed precedence over Pepliphorus Hübner,[1819] PDF

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Preview Case 2851. Nacaduba Moore,[1881] (Insecta, Lepidoptera): proposed precedence over Pepliphorus Hübner,[1819]

. BullelinofZoologicalNomenclature50(11March1993 35 Case2851 NacadubaMoore,|1881|(Insecta,Lepidoptera):proposedprecedence overPepliphorusHiibner,|1819| ToshiyaHirowatari EntomologicalLaboratory,CollegeofAgriculture. UniversityofOsaka Prefecture,Sakai.Osaka,593Japan Abstract.ThepurposeofthisappHcationistoconservethenameoftheOrientaland AustraHanlycaenidbutterflygenusNacadubaMoore.[1881],Itisproposedthatitbe given precedence over the senior subjective synonym Pepliphorus Hiibner. [1819], rarelyusedinthissense. 1 PepliphorusHiibner.[1819](p.71)wasestablishedfortwotaxonomicspecies,the first being '698. Pepliphorus Euchylas. HylasCram." and the second '699. P[apilio] cyamisEabr.[errorforFabricius].CyaneaCram.';abbreviatedreferencesweregiven leadingtoPapiliohylasCramer,[1781](p. 142,pi. 363)andPapiliocyaneaCramer, [1775] (p. 120, pi. 76), ofwhich P. cyanusFabricius, 1781 (p. 116)isan unjustified emendation.AsnotedbyFruhstorfer{1916a,p.33)Hiibner'sspecificnameeuchylas was a replacement name forhylas Cramer: there are at least three senior primary homonymsofthelatter, theoldestbeingPapiliohylasLinnaeus, 1758(p.486).The synonymyofeuchylasandhylasCramerwasmentionedbyRothschild(1915,p. 138), Toxopeus(1927,p.267)andD'Abrera(1990,p.356). 2. Scudder(1875,p.245)listedPapiliocyaneaCramer,[1775]asthetypespeciesof Pepliphorus:thisisavaliddesignation,butsubsequentlyWaterhouse&Lyell(1914, p. 101)designatedPapiliohylasCramer,[1781].AsnotedbelowPepliphorushasbeen usedonlyinthelatter{euchylas)sense.ItisScudder'slargelyignoreddesignationwhich hascausedthepresentdifficulty. 3. Moore([1881],p.88)proposedthegenusNacadubawithtypespeciesbyoriginal designation LampidesprominensMoore. 1877(p. 341); thisiscurrentlytreatedasa subspeciesofLycaenakuravaMoore,[1858](p.22). 4. ThenameNacadubahasbeenverywidelyusedinbothprimaryresearchpubli- cationsandrevisionalworks(e.g.Fruhstorfer,1916b;Corbet,1938;Eliot.1955,1973; Tite,1963;1 referencespublishedsince1981andadditionaltothosementionedinthis applicationareheldbytheCommissionSecretariat).Nacadubapresently(Hirowatari, 1990)contains 38 species from the Oriental and Australian Regions, including the South Pacific Islands. Recently two species of Nacaduba. N. berenice (Herrich- Schaefier, 1869)andN.normaniEliot, 1969,havebeenrecordedaspestsofcocoain Sabah,EastMalaysia,byFujii(1987)andPan&Morishita(1990)respectively. 5. Although Scudder'sdesignation (seepara. 2)ofcyaneaas the type species of Pepliphorus was noted by Toxopeus (1927, p. 267) and Hemming (1967, p. 351), Pepliphorushasscarcelybeenrecordedinthissense;perhapsBridges's1988Catalogue entryistheonlyinstance. Butler(1900,p.442)referredto'...Pepliphorus(P.hylas. ) 36 BulletinofZoologicalNomenclature50(I)March1993 Cramer,andallies).,.",andasalreadymentioned(para.2)Walerhouse&Lyell(1914) invalidly designated hyla.s (=euchylas) as the type species. Rothschild (1915) and Friihstorfer(1916a)usedPepliphorusforeiichylas,asdidJoicey&Talbot(1917,under thename P.hylas).Toxopeus(1929,p. 230)attemptedtodealwiththesituationby erecting the new genus Pvploclyia with Papilio cyanea Cramer as the type, despite Scudder'sidenticaldesignationforPepliphorus.InrecentyearsPepliphorushashardly beenusedatall,andeuehylasandalliedspecieshavebeenincludedinJumiclesHiibner, [1819)(p.71)(see,forexample,D"Abrera, I990,p.355). 6. BecauseofitsexternalappearancePapiliocyaneaCramerhascustomarilybeen placedinDanisFabricius, 1807(ortheobjectivelysynonymous ThysonoiisHiibner, [1819],p.20)(e.g.Common&Waterhouse, 1981,p.576)orinPsychonoiisToKopeus, 1930(p.129)(e.g.D"Abrera,1990,p.326);thelatterassignmentignorestheseniorityof Pepliphorus. I (Hirowatari, 1987. p. 35) pointed out theclose relationship between cyanea and species ofNacacluha, but temporarily retained Pepliphorus for cyanea alone,followingScudder'stypespeciesdesignation.Currentstudies(Hirowatari,1992) haveshownthatthereisnomorphologicaldifTerencebetweencyaneaandNacacluha species,sothatPepliphorusandNacaduhaaresubjectivesynonyms.SinceNacaduhais awidelyusednameandPepliphorusisunusedforthespeciescurrentlyattributedto Nacaduha.IproposethatthenameNacaduhabegivenprecedenceoverPepliphorus wheneverthetwoareconsideredtobesynonyms. 7, The International Commission on Zoological Nomenclature is accordingly asked: (1 touseitsplenarypowerstogiveprecedencetothenameNacaduhaMoore,[1881] overthenamePepliphorusHiibner,[1819],wheneverthetwoareconsideredto besynonyms; (2) toplaceontheOfficialListofGenericNamesinZoologythefollowingnames: (a)Nacaduha Moore, [1881] (gender: feminine), type species by original designationLampidesprominensMoore, 1877,withtheendorsementthatit istobegivenprecedenceoverPepliphorusHiibner,[1819]wheneverthetwo namesareconsideredtobesynonyms; (b)PepliphorusHiibner,[1819](gender:masculine),typespeciesbydesignation byScudder(1875)PapiliocyaneaCramer,[1775],withtheendorsementthat itisnottobegivenpriorityoverNacaduhaMoore,[1881]wheneverthetwo namesareconsideredtobesynonyms; (3) toplaceontheOfficialListofSpecificNamesinZoologythefollowingnames: (a)prominensMoore, 1877,aspublishedinthebinomen Lampidesprominens (specificnameofthetypespeciesofNcuaduhaMoore,[1881]); (b)cyaneaCramer,[1775],aspublishedinthebinomenPapiliocyanea(specific nameofthetypespeciesofPepliphorusHiibner,[1819]); (4) toplaceontheOfficialIndexofRejectedandInvalidGenericNamesinZoology thenamePeplodyiaToxopeus,1929(ajuniorobjectivesynonymofPepliphorus Hiibner,[1819]). References Bridges,C.A.1988.CatalogueofLycuenidae&Riodinidac(Lcpidopicra:Rhopaloccraj.816pp. Author,Urbana. BulletinofZoologicalNomenclature50(1)March1993 37 Butler,A.G.1900M.AlistofthespeciesofCyaniris,awell-knowngroupofthefamilyLycaenidae. Annalsand agazineofNaturalHistory,(7)5:441^51. Common,I.F.B.&Waterhouse,D.F. 1981.ButterfliesofAustralia,(revisededition).682pp. Angus&Robertson.Sydney. Corbet, A.S. 1938. A revision ofthe Malayan species ofthe Nacaduha group ofgenera (Lepidoplera,Lycaenidae).TransactionsoftheRoyalEntomologicalSocietyofLondon,87: 125-146. Cramer,P.[1775],DeUillandscheKapellenvoorkommendeindedrieWaereld-deelenAsia.Africa enAmerica,vol.I.155pp.,96pis.Baalde.Amsterdam&Wild.Utrecht. Crameenr,AmP.er[i1c7a8,1]v.o£l).e4U.i2t5l2a,nd2s9chppe.K,a1pe1l1lpeins.vBoaoarlkdeo,mmAemnsdteeirnddaemdr&ieWWialedr,eUltdr-edcehetl.enAsia.Africa D'Abrera, B.L. 1990. Butterflies of the Australian Region, Ed. 3. 416 pp. Lansdowne, Melbourne. Eliot,J.N.1955.NotesontheNacaduhahermtis(C.Felder)complex(Lepidoptera:Lycaenidae). ProceedingsoftheRoyalEntomologicalSocietyofLondon,(B)24:153-158. Eliot,J.N.1973.ThehigherclassificationoftheLycaenidae(Lepidoptera):atentativearrange- ment.BulletinoftheBritishMuseum(NaturalHistory),Entomology,28:371-505. Fabricius,J.C.1781.SpeciesInseclorum,vol.2.517pp.Bohn.HamburgietKilonii. Fruhstorfcr,H.1916a.RevisionderGattungLum/7/rfejaufGrundanatomischerUntersuchungen. ArchivfurNaturgeschichte,81A(6):1^6. Fruhstorfer, H. 1916b. Revision der Galtung Nacaduha aufGrund der Morphologic der Generationsorgane.ZoologischeMededeelingen,2:103-140. Fujii,H.1987.BehaviourofbutterfliesinSabah.Pp.\5-^in:Studiesonthebehaviourandthelife- cyclestrategiesofarborealinsectsinthehumidtropics.1985.supplement.KyotoUniversity, Kyoto. Hemming,F.1967.Thegenericnamesofthebutterfliesandtheirtype-species.BulletinoftheBritish Museum(NaturalHistory),Entomology,Supplement9:1-509. Hirowatari, T. 1987. Systematic position of Pepliphorus cvanea (Cramer) (Lepidoptera. Lycaenidae).TyotoGa.38(1):31-37. Hirowatari,T.1990.OverviewonthegenusNacaduhaMoore(Lycaenidae).Yadoriga,142:2-11. [InJapanese.] Hirowatari,T. 1992. Agenericclassificationofthe tribe Polyommatini oftheOrientaland Australianregions.(Lepidoptera,Lycaenidae,Polyommatinae).BulletinoftheUniversity ofOsakaPrefecture.(B)44supplement:1-102. Hiibner,J.[1819].VerzeichnisshekannlerSchmettlinge,part5.Pp.17-176.Augsburg. Joicey,J.J.&Talbot,G.1917.NewLepidopterafromWaigeu.DutchNewGuinea,andBiak. AnnalsandMagazineofNaturalHistory,(8)20:216-229. MLiononraee,usF,.C[.1815785)8..ISn:ysHtoersmfaieNladt,urTa.e.&EMdo.o1r0e,,voFl..A1.8c2a4taplpo.guSealovifi,thHeollempiiaden.pterousinsectsofthe museumoftheHon.East-IndiaCompany,vol.1.277pp..18pis.Allen.London. Moore,F.1877.DescriptionsofCeylonLepidoptera.AnnalsandMagazineofNaturalHistory. (4)20:339-348. Moore,F. [1881]. TheLepidopteraofCeylon, vol. 1, part 3. Pp. 81-136, pis. 37-54. Reeve, London. Pan,T.-S.&Morishita,K. 1990.NotesonNacaduhanormaniEliotinSabah,NorthBorneo (Lepidoptera.Lycaenidae).TyotoGa.41(3):149-154. Rothschild,W.1915.OnLepidopterafromtheislandsofCeram(Seran),Buru,Bali,andMisol. NovitatesZoologicae.22:105-144. Scudder,S.H.1875.Historicalsketchofthegenericnamesproposedforbutterflies:acontribution tosystematicnomenclature.ProceedingsoftheAmerican.AcademyofArtsandSciences,10: 91-293. Tite, G.E. 1963. A synonymic list ofthe genus Nacaduha and allied genera (Lepidoptera: Lycaenidae).BulletinoftheBritishMuseum(NaturalHistory).Entomology,13:67-116. Toxopeus,L.J. 1927. EineRevisionderjavanischen.zuLycaenopsisFelderundverwandten Genera gehorigen Arten. Lycaenidae Australasiae II. Tijdschrift voorEnlomologie. 70: 232-302. 38 BulletinofZoologicalNomenclature50(I)March1993 Toxopeus,L.J.1929.DeRiodinidaeenLycaenidaevanheteilandJava.(LycaenidaeAustralasiae VI),TijdschriftvoorEiUomolugie,11:215-244. Toxopeus.L.J. 1930.DesoonalsfunclievanplaalsenlijdgetoelstaandeLycaenidaevanhel Aiistralaziatischgebied.198pp..4pis.Paris.Amsterdam. Waterhouse,G.A.&Lyell,G. 1914. ThebutterfliesofAustralia.239pp.Angus&Robertson, Sydney.

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