CARNIVOROUS PLANTS of the United States and Canada by Donald E. Schnell CARNIVOROUS PLANTS of the United States and Canada Sarracenia flava JOHN F. BLAIR, Publisher Winston-Salem, North Carolina CARNIVOROUS PLANTS of the United States and Canada by Donald E. Schnell Copyright © 1976 by Donald E. Schnell Library of Congress Catalog Card Number: 76-26883 All rights reserved ISBN 0-910244-90-1 Printed in the United States of America by Lebanon Valley Offset Company Incorporated Library of Congress Cataloging in Publication Data Schnell, Donald E 1936- Carnivorous plants of the United States and Canada. Bibliography: p. Includes index. 1. Insectivorous plants—United States. 2. I sectivorous plants—Canada. I. Title. QK917.S36 583'.121'0973 76-26883 ISBN 0-910244-90-1 Sarracenia psittacina For Lee Anne & Kristen Foreword By C. RITCHIE BELL Professor, Department of Botany, University of North Carolina, Chapel Hill and Director, North Carolina Botanical Garden, Chapel Hill Since their discovery by botanists over two hundred of the plant world from rapid extinction through a years ago, the world's carnivorous plants have been the combination of realistic protection from continued col center, from time to time, of much scientific study and lecting, conservation of their specialized habitats, public interest. They have also been the center of a responsible horticultural research, and the general cul number of misconceptions and, in the past few years, tivation of only those plants actually grown from seeds of considerable commercial exploitation. or, where possible, cuttings. Carnivorous plants are both colorful and biologi C. R. B. cally interesting, but they are not house plants. Their July, 1976 specific environmental requirements often involve very high humidity, high light intensities, quite acid soil, pure water, and seasonal temperature changes; such requirements are not met by a pot of generalized dirt, treated tap water, and the usual temperature and hu midity ranges of the average home, office, or apart ment! It is likely that not one "cultivated" carnivorous plant in a thousand lives a full year and probably less than one in ten thousand survives for two years in the hands of most amateur "collectors" or those who, un fortunately, are misled by the commercial advertise ments for these unique plants and attempt to grow them, merely as a novelty, on the kitchen window sill. Given the above background, this book becomes something of a landmark in carnivorous plant publica tion. Not only does it bring together in a very readable fashion the many interesting botanical facts concerning the form, function, and ecology of our carnivorous plants, but the author makes a strong, and very timely, case for the preservation of these unique members vi / Carnivorous Plants of the United States and Canada Preface This book is intended for practical use. It is not con more readily observed plants or those that would best ceived of as a coffee table or bookcase ornament; the illustrate general characteristics, or perhaps grouping entire plan and structure of the work is centered around similar species together so that comparisons could be utility. seen. The photographs have been chosen with a view The carnivorous fungi have been excluded. The book toward sharpness of important features, clarity, color deals only with the green seedplants. Fungi require cul fidelity, and most of all, how well they represent the ture and microscopes for proper observation, and the plant. You will find only a few drawings in this book, species are in a state of taxonomic disarray at present. and these are mainly of a figurative nature. The trouble While fascinating and certainly deserving of further with using only botanical drawings for identification is study, they are not ready for this sort of presentation. that they represent a two-dimensional, colorless av The text is not complete in the classical botanical erage of characteristics in an ideal plant that rarely sense of a monograph, but it was not intended to be. exists. This presents a problem for the beginner in the On the other hand, the occasional or beginning natu field, who has a book of drawings in hand but sees no ralist may find more than he needs and can easily be plants that resemble any of the drawings, or perhaps selective in what he reads. Those interested in further sees too many. study will find references on many levels at the end The photos are intended to show the plants as they of the book. really are, and most of the pictures have been made in • • • the field. If some of the photographs seem pretty or artistic, that is just a bonus. Since no view of a plant I would like to thank John F. Blair, Publisher, and or group of plants can show all the important character his staff for taking on this project in the first place, istics one might wish to see, I have presented multiple and then seeing it through to a quality production. views of them where necessary. Warren P. Stoutamire was kind enough to read por Occasionally, in the field, where the natural back tions of the text and offer many useful suggestions. grounds tended to camouflage, I have had to resort to Letters and discussions with Peter Taylor and Katsu- the use of a neutral gray background, reflectors, or an hiko Kondo were very helpful to me in developing my cillary lighting to make the subjects stand out properly. tentative concepts of the difficult taxonomy of Utricu- Only a few pictures were made under studio condi laria, but I bear full responsibility for the system tions, using live plants from my collection. After hav presented in Chapter 7, including errors. Finally, I ing observed, grown, and worked with carnivorous offer tenderest appreciation to my wife Brenda, who plants for eighteen years, I felt capable of making a has been patient and encouraging during the prep proper selection of cultivated plants for photography. aration of this book and who has accompanied me I have departed from the traditional botanical scheme on many field trips, often serving as my stern early- of presenting species within a genus in alphabetical or warning system for potentially dangerous reptiles, other order, preferring to start with some common, quicksand, and treacherous bogholes. vii / Preface Contents Foreword vi Drosera anglica 61 Drosera intermedia 63 Preface vii Drosera filiformis 64 Drosera capillaris 67 Chapters Drosera brevifolia 69 Difficult Identifications 70 I. Carnivorous Plants—An Introduction 1 Carnivorous or Insectivorous Plants? 4 VI. The Butterworts (Pinguicula) 71 Kinds of Traps 4 Pinguicula vulgaris 74 Carnivorous Plants around the Continent 6 Pinguicula pumila 76 The Future of Carnivorous Plants 9 Pinguicula lutea 77 Flower Structure 10 Pinguicula caerulea 78 How Plants Are Named 13 Pinguicula planifolia 78 Pinguicula primuliflora 79 II. The Venus' Flytrap (Dionaea muscipula) 16 Pinguicula ionantha 80 Pinguicula villosa 81 III. The Eastern North American Difficult Identifications 82 Pitcher Plants (Sarracenia) 22 Sarracenia purpurea 28 VII. The Bladderworts (Utricularia) 83 Sarracenia flava 31 VIII. Growing North American Sarracenia alata 34 Carnivorous Plants 96 Sarracenia oreophila 36 General Principles 96 Sarracenia minor 37 Genus Notes 106 Sarracenia psittacina 40 An Outdoor Home Bog 109 Sarracenia leucophylla 42 Some Commercial Sources of Sarracenia rubra 44 Carnivorous Plants 111 Sarracenia Hybrids 47 Field Collecting 112 Difficult Identifications 51 Additional Reading 114 IV. The California Pitcher Plant (Darlingtonia californica) 52 Glossary 118 V. The Sundews (Drosera) 56 Derivations of Scientific Names 121 Drosera rotundifolia 58 Drosera linearis 59 Index 124 ix / Contents