Butterflies of the central Lao P.D.R. in the low temperature and dry season (1997-2001)

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esttw T}'ansI.epidS.oc.Japan 56 (1)3:6-50,January2005

Butterfiie sof the central Lao RD.R. in the low temperature and dry season
(1997-2001)
Masayuki NiMuRAi )and Hiroyuki WAKAHARA2)
]' Department of Developmental Biology, Shimane Vniversit Iyz,umo, 693-850 1Japan
2}P.O. Box 6217,Vientianet,heLau P.DR

Abstract IIhe distributie nof butterfli eisn the central Lao P.D,R. was k]yestigat iend the low tem-
perature and dry season, We recerded in tota l221 species, corresponding to approximately 25% of
876 species recorded in `ttln itlustrate dChecklist of the Buttetfiie osfLaos P. D. R," (Osad aet al,,
1999).
Key words Butterfiy fauna, the central Lao P.D.R., low temperature and dry season.
Intreductio"
There are many reports about the butterfl ioefs the Lao RD.R. (Motono & Negishi, 1989;
Masui & Uehara,1994,1995,1996,1997;Uehara& Ybshida, 1995;Sashida& Miyamoto,

1996,1998;Koiwaya & Osada,1998).Based thesefindingsO,sada (1999)
on et al. pro-
duced a summary report of the butterfi ioefs the Lao RD.R, Recently ,Masui & Uehara
(1999 2,000) and Sashida (2000 r)eported several newly recorded species in the Lao P.D.R.
In addition, detail sof morphelogical changes, larva lfbod plant and behavior from egg to
adult were examined in several species of butterfl iiens the Lao RD,R. (Koiway eat al.,
1996, 1997, 1998, 1999a,1999b, 2002).

Jn the presen tstudy, the distributi oofn adult butterfl iiens the central Lae RD.R. was inyes-
tigated in four separate periods at the end of December (1997 ,1998 and 1999) and
November (2001 )[,Ib obtain a reproducible result on the distributio nof butterfi iiens the
short term, 4 locati-o onns the Mekong River plain and 3 non-riverine location wsere select-
ed fOr sampling of as many species of butterfl iaess possibl e,It had been expected that only
a few species would have been collected during this cold season. However, we collected
Nymphalidae fycaenidae.
relatively many species, especially of and
In this report, we established a lis tof identifi ebdutterfii feosr the central Lao RD.R. in the
cool and dry season from 1997 to 2001,
Investigation (Fig,1)
areas
We established the following investigatio nareas, (1 )Mekong River plains (1-4 ;altitude:
180-200 m) and (2 )inland areas (1-3 ;altitude: 250-700 m).
(1)-1T.habok(Tb)

This area is locate dapproxirnately 1 20 km east along the Mekong River from Vientian eand
mainly consists of mixed fbrest sof evergreen and deciduou strees. Altitude: approximately
200 m.
(1)-2N.arndhua (Nd)

This area is locate dapproximately 150 km southeast along the Mekong River from Thabok.
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Butterfli oefs the central Lao P.D.R. 37
Lk
R13
Tb Nd Nh Vi etf ailn
R8
VienSiane
Nk

Tk
X

Rl2Tk
Mekong/
River

[Irkaiiazzd
Fig .1. Investigat iaorneas. Tb: Thabok, Nd: Namdhua, Th: Thabeng, Tk: Thakhek, Nh: Nhahin,
Lk:Laksao,Nlc:Nakai.

This fores tmainly consists of evergreen trees. Altitud ea:pproximately 200 m.
(1)-3T.habeng (Th)

This area is located approximately 40 krn southeast along the Mekong River from
Namdhua, This fores tmainly consists of evergreen trees. This area is also ca]Ied
"Vienthone". Altitude a:pproximately 180 m.
(1)-4T,hakhek (Tk)

Thakhek is a relatively big city of the Lao P.D.R. and approximately 90 km seutheast along
the Mekong River from Thabeng. The fbrest sjn and around Thakhek mainly consist of
evergreen trees on limestone-zone. Altitude a:pproximately 200 m.
(2)-1N.hahin (Nh)

This area is a limeston ezone locate dapproximately 40 km east aleng Route 8 from
Namdhua. Altitude: approximately 250 m to 700 m.
(2)-2L.aksao(Lk)

This is a platea uarea locate dapproximately 50 km east along Route 8 from Nhahin. We
exarnined the periphery of this tQwn and the area around the power station close to the Lao-
Vietnamboundary,Altitude: 600
approximately m.
(2)-3N.akai (Nk)

This is also a platea uarea located approximately 50 krn northeast along Route 12 from
Thakhek. Altitude: approximately 600 m.
Foridentification Pinratana(198'119,83,1985,1988),Motono &
of most samples we used
Negishi(1989)P,inratana& Eliot(1992,1996),Smith (1994) Osada (1999).
and et aL
However, for some diMcult sarnples, we requested identificat ifornom the followin gre-
searchers.
Pieridae:Dr Haruki Sashida

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38 Masayuki NiMuRA arid Hiroyuki WAKAHARA
SatyridaeM:r [[latsuMoiyamoto

fycaenid Mare [:latsu oMiyamoto and Mr Satoshi Koiwaya
Investigation
period
In central Laos, hardly any rain fall sin the cool and dry season from the middle of
November to March. TThe seasonal temperature is the lowest at any point in the year, We
investigate dthe distributi oofn butterfii iens the 4 period sas fo11ows.
1)25-30December 1997. Thabok,Namdhua, Thakhek,Nhahin

2)26-31December 1998. Thabok,Namdhua, Thakhek,Nhahin,Laksao

3)25 December 1999-3January2000,Namdhua, Thabeng,Thakhek, Nhahin,Laksao

4)25-30November 2001.Thabok,Namdhua, Thabeng, Thakhek, Nakai

The lis tof collection
We categorized the samples from the Mekong River plain sas `low land' (Thabok,
Namdhua, Thabeng and Thaldiek )and the non-riverine areas as Cinland' (Nhahin L,aksao
and Nakai) .The data from each year were added together .We excluded specimens without
identification.
complete
In total 221 species were recorded in thi sinvestigati oOnf, 221 species, 199 species were
collected in lowland and 79 in inland areas. Fifty-seven species were recorded in both
areas, 142 in lowland only (show nas * in list )an,d 22 in inlan donly (shown as # in list )re-
The details follows,
speetively. are as
Papilionidae 12 species, Pierida e17 species, Danaidae 14 species, Nymphalidae 61 specjes,
Satyrida e11 species, Amathusiidae 7 species, Rjodinida e7 species, Lycaenidae 67 species,
and Hesperiidae 25 species.
The 221 species in the presen tcollection correspond to approximately 25% of the 876
species recorded in `tAn itlustrate dChecktist of the Butteijlie sofLao sP. D. R." (Osada et
1999).
al.,
Papilionidae
Byasalaos(Riley& Godfrey,1921)'

[lowland]Tk:1g,28. 1997,
xii.
Lesaria doubledayi(Wallac1e8,65)'
coon
[lo wland] Nd: 1X, 30. xi, 2001. Th: 18, 1. i .2000,
Ptichliqpt aaristolochiae goniopertis (Rothsch i1l9d08,)
[lo wland] Nd: 18,30. xii.1997. Tk: 1X,29. xii.1997; 18, 30, xii,1999,
[inlanNdh]: 18,27. 1997.
xii.
lkpitiodemole"sdemoleusLinnaeus,1758#

[inianNdh:] 18,27, 1997.Nk: 18,28, 2001.
xii, xi.
lhpiliodemotiondemotionCramer,1776"

[lo wland] Tb: 18, 26. xii. 1997. Nd: 18, 30. xii. 1997;2 S, 1. i. 20eO.
Ilapiliohelen"s helenus Linnaeus, 1758

[lo wland] Tb: 1e, 25. xi. 2001.
[inlan Ndk]: 2g, 28. xi. 2001.
Papilio polytes romulus Cramer, 1775*
[IQ wland] Nd: 1 \, 30. xi. 2001.
Ptipitio Linnaeus,1758*
memnon agenor
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Butterfii oefs the eentral Lao P.D.R. 39
[lo wland ]Nd: 28,29-30. xi. 2001. Tk: 18,31, xii. 1999.
Lci,uproptera curius curius (Fabric i1u78s7,)
[lowland]Nd: 1g, 30. 2001.
xi.
[inlanNdh:]18 27. 1997.Lk:18,28, 1999.
xii. xii.
Lampmptera meges virescens (Butl e1r87,0)
Ilow land ]Tb:38, 26. xii. 1997, Nd: 18,3. i .2000. Th: 18, 26, xi. 2001, Tk:18, 26.
xii. 1999.
[inlan dNh]: 28, 28. xii. 1998.
Graphium sarlpedon luctati u(sFruhstQr 1f9e0r7),"
[lo wland] Tb: 28, 26. xii. 1997.
Gvaphium agamemnon agamemnon (Linnaeu s17,58)*
[lo wland] Nd: 1r1g, 30. xii. 1997; 19, 26, xiL 1998; 19, 3. i .2000; 1g, 26. xi. 2001.
Pieridae
Deliaspasithoethyra Fruhstoder, 1905"

flo w!a"d] Nd: 18 1g, 26. xi. 2001.
Deliash)tparetiendica(Wallac1e86,7)"

[lo wLand ]Tb: 18, 26. xii. 1997, Tk: 1\,3e. xii. 1997.
Leptosia (Fabrici1u7s93,)"
nina nina
[lowland]Nd: 1g, 26. 1998;18, 2.i.2000;18, 26. 2001. Tk: 18, 28.
xii. xi. xii,
1997.

7kelboti anaganum pamsi (Vita ldie sSaivaza ,1921) #
[inlanLdk]:1g,28. 1999,
xii.
Ceporzz (Lucas1,852)
nadina nadina
[lowland]Tb:18,26. 1997.Nd: 1S1\, 26. 2001.
xii. xi.
[inla nLdk:] 1 8, 28, xii. 1999.
Ampiaslyncidu (Boisduval,1836)
eleonora
[lowland]Nd: 28, 27-30. 1997;281?,3. i,2000;28, 26. 2001. Tk:14, 30.
xiL xi.
1997;2?,27, 2001.
xii, xi.
[inla nNdh]: 18, 27. xii. 1997. Nk: 28, 28. xi. 2001.
Ampias albina dartid a(C .& R. Felder ,186S)*
[lo wland] Nd: 1\, 30. xii. 1997; 18, 2. i .2000 .Tk: 18, 28. xii. 1997; 1S, 27. xii.
1998;19,27. 2001.
xi.
Al?piainsdrzzthrvnion Fruhstorfer,191O"

[lo wland ]Tb: 18, 26. xii. 1997. Nd: 1g, 27. xii. 1997.
Ixias pyrene vema H. Druce, 1 874 '1997;
[lo wland] Nd: 4 819,27-30, xii. 28, 26-27. xii. 1998; 28, 26-29. xi. 2001.
Th: 2c7i 2,6. xi. 2001, Tk: 29, 28. xii. 1997; 19, 27. xii. 1998; 4c7i ,26. xii. 1999; 3,3 2"
?,27, 2001.
xi.
[inla nNdh:] 381e,27. xii. 1997;1g, 30. xii. 1999. Nk: 28, 28. xi. 2001,
Hebomoia glaucippe glaucippe (Linnae u1s75,8)"
[Io wland] Nd: 28, 30. xii. 1997; 18, 2. i ,2000; 28, 26. xi. 2001. Th: 18, 25. xii.
1999. Tk: 1 cl4,31. xii. 1999.
Prionerisphiionomeheijler(iC.& R. Felder,l865)"

[lo wland] Tb: 2g, 26. xii. 1997. Nd: 28, 30. xii. 1997; 1\, 3, i .2000; 1g, 26. xL
200L

I]kereronia (Lesson18,37)*
anais anais
[lo wland] Nd: 1t, 30. xii. 1997; 28, 2. i .2000; 28, 29-30. xi. 2001. Tk: 18, 31. xji.
1999.

IlareroniaparavatarBingham, 1907

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[low land] Tk: 1 8, 28. xii. 1998.
[inlan dNh]; 18, 27, xii .1997.
Catopsilia (Fabrici1u7s87,)
scylla comelia
[lowland]Tk:1\, 27. 2001.
xi.
[inlanNdk]: 19,28. 2001.
xi.
Catopsilia (Fabrici1u7s75,)
pomona pomona
[lo wland ]Nd: 1\ (f p.omona), 26. xii. 1998; 1･9 ･(f .catilla (Cramer l,777)"), 3. i. 2eOO.
Tk: 1 8 (f h.ilari a(Ste l1l7,81)) ,26. xii. 1999.
[inla nNdh:] 1 8 (f h,ilari a(Stol L1781)) 29 (f p.omona (Fabrici u1s77,5)) ,28. xii. 1998.
Lk: 28 (f .hilaria )2,8-29. xii. 1999.
E"rema hecabe hecabe (Linnaeus17,58)*

[io wland ]Tb: 2l, 25, xi. 200L Tk: 1r19,27-28. xii, 1997;18, 27. xi. 2001,
Gandaca harinab"rmana Moore, 19e6*

[lo wland] Tb: 1 8, 26. xii' .1997.
Danaidae
Danaus genutiagen"tia(Cramer17,79)

[lo wland] Tb: l ,7', 25, xii. 199g. Th: 2 ,gn, 25-27. xii, 1999.
[inlan dLk]: 1 g, 29. xii. 1999,
Danaus hegesippus(Cramer1,777)*
melanippus
[lowland]Nd: 18,26, 2001,
xi.
nrumala septentrionis sqptentrionis (Butl e1r8,74)
[]owland]Tk:184,27. 1998.
xi.i,
[inlandNh]: 18,30. 1999.
xii.
nrumala limniacelimniace(Cramer[,1775]#)

[inlan dNh]: 1 8, 30. xii. 1999.
ldeopsis (Fruhstor1f9e04r),"
vulgaris macrina
[lowiand]Tb: 18, 24. 1999.Nd: 28, 26. 2001.Tk: 1S,31. 1999;13, 27.
xii. xi. xii,
2001,
xi.
ldeopsis simitis persimilis (Moore ,1879)"
[low land] Tk: 18, 27. xi. 2001.
Ptzranti caaglea melanoides Moore, 1883
[low land] Nd: 1r. 27. xii. 1997. Tk: 18, 28. xii. 1997; 1r, 31. xii. 1999.
[inlan dNhl: 1g, 27, xii. 1999. Nk:18, 28. xi. 2001.
Ptirantic amelaneus (Cramer 1,775) #
[inlan dLk]: 28, 28, xii. 1999.
Euptoea mulciber mulciber (Crame r1,777)
[low land] Tb: 1g, 26. xii. 1997; 133 9, 24, xii. 1999, Nd: 29, 26. xii, 1998. Tk: 1g,
28. xii, 1997,
[inlan dNh]: 18l9,27. xii. 1997. Nk: 1e, 28. xi, 2001.
Eaptoea But]er,1866
eamaralzeman camaralzeman
[low iand] Tb: 1 8i ,26. xii. 1997. Th: 1 6f i2,5. xii. 1999.
[inlan dNhl: 1 ?, 27. xii. 1997.
Euploea dottbteddyidoubtedoyi C. & R, Felder,1865"

[lowland]Nd: 19,2.i.2000.

Euploea tulliolusdehaaniLucas,1853*

[low iand] Tb: lr, 24, xii. 1999. Nd: 18, 2. L 2000,
Euploea radainanthus radamanthus (Fabric i1u7s93,)
[lo wland] Tb:18, 25. xii. 1998;18, 25. xi. 2001, Nd: 2824,26-29. xi, 2001. Th: 18
15i,1.i.2000.

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[inla nNdh:] 18, 27. xii. 1997; 588?, 30. xii. 1999. Lk: 18, 30. xii. 1998. Nk: 1X,
28.xi.2001.

Eaptoea eunice leucogoni s(Butle r18,79)
[lo wland ]Nd: l\.26. xi. 2001. Tk: 18, 26. xii. 1999.
[inla nNdh]: 1 8, 30, xii. 1999.
Nymphalidae
Ariadne (Fruhstor1f9e12r),
merione pharis
[lo wland] Th: 1 g, 26. xi. 2001,
L'inl aLnk:d ]1 ,g, 30. xii. 1998.
Phalanta alcippe alcippoides (Moore ,1 900)"
[lo wland] Th: 1 8i ,27. xii. 1999, Tk: 18", 26. xii. 1999; 1 8` ,27. xi. 2001.
Cupha lotis(Sulze1r77,6)"
erymanthis
[lo wland] Nd: 3 8, 26-29. xi. 2001.
Cirrochro atyche mithila Moore, 1872
flo wland] Nd: 1 8, 3. L 2000; 1 8, 30. xi. 2001.
[inla nNdh]: 19,27, xii, 1997; 1g, 27. xii. 1999.
1,lndul aerota erota (Fabric i1u79s3,)'
[lowland]Nd: 18,26. 1998.Th:1?,27. 1999.
xii. xii.
7brinos clarissafalcata Fruhstorfe r1,898*
[low land] Nd: 3g,3. i .2000;38', 26-29. xi. 2001.
71irin otserpander intermedia Godfrey ,1916#
[inlan dNh,:1 1 8, 30. xii. 1999.
Cethosia bibli stisamena Frehstorfer ,l 912
[lo wland] Nd: 28, 27. xii. 1997;2g, 25. xii. 1999; 28, 3, i .2000,
[inlan Ndh]: 14,30. xii. 1999. Nk: 1g1?,28, xi, 2001.
Cethosia cyane euanthes Fruhstorfe r1,912
[lo wland] Nd: 1 9, 30. xii. 1997.
Linlan Ldk]: 2{la 2,4-28. xii. 1999. Nk: 1 8, 28. xL 2001.
.htnoniaiphitiaphita(Cramer1,779)*

[low land] lk: 1 4, 27. xii. 1998,
Jdenonia (Linnaeu1s75,8)
atmana almana
[low land] Nd: 18, 27, xii. 1997; 18, 26. xi. 2001. Th: 18, 25. xii. i998; 18, 24. xii.
1999.Tk:18,27.xL 2001.

[inla nNdh]:18, 28. xii. 1998. Lk: 18, 28, xii. 1999. Nk:18, 28. xi. 2001.
Junonia atlites atlites (Linnae u1s76,3)*
[lo wland] Nd: 1,7 i2,8. xii. 1997;1,?i, 26. xii, 1998; 1clA ,26. xi. 2001. Th: 18i ,25. xii.
1998.Tk:18,29, 1997.
xii,
.htnonialemoniaslemonias(Linnaeus1,758)*

[low land] Tk: 1 8, 31. xii. 1999; 1?,28. xii. 1997.
hiertahierta(Fabriciu1s79,8)
.htnonia
[lo wland ]Tb: 14,25. xii.1998. Tk: 2,7i 2,9. xii. 1997.
[inla nNdh:] 1 8, 28. xii. 1998.
Kliniska (Linnaeu1s76,3) #
canace canace
[inla nLdk]: 1 8, 31. xii. 1998; 8, 29. xii. 1999.
Symhrenthliitaaealilaea(Hewitson1,864)

[low land] Tk: 18, 26, xii. 1999. Nd: l8, 3, i. 2000.
[inlan dLk]: 1 X, 28. xii. 1999.
Cynest ithsemiTe siamensis Fruhstorfer ,1898
[lowlandJNd: 18",26. 1998.Tk:18i,28. 1997.
xii, xii.
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[inl Nh: 18,27. 1997.
andJ xii.
Ciyrest icoscles coctes (Fabric i1u78s7,)"
[]Q wland] Tk: 1 6n ,26, xii, 1999.
Ciyrest iths{yodamas thyodamas Doyere, 1 840 #
[inlan dNh]: 1 g, 27. xiL 1997.
Chersonesia risa transiens (Manin 1,903)
[lowland]Th:18,26. 1997.
xii.
[inlanLdkl: 2r, 28, 1999.
xii,
Chersonesia internzedia Moore,1899*
rahrioides
[low land] Th: 1 ,3a, 26. xii. 1997. Tk: 1 cln, 27. xi, 2001.
Rhinopaipapolynice hirmanaFruhster[fer1,897*

[lo wland] Nd: 1?, 29. xi. 2001.
Iilypolimnabsolinaholina(Linnaeu1s75,8)"

[lowland]Th:19,25. 1998.
xii.
Doleschalli haisaltid econtinentalis Fruhstorfer ,1 899*
[lo wland] Nd: 19,2, i ,2000;29,29. xi. 2001.
Neptis Moore, 1872*
clinia susr"ta
[low land ]Nd: 2g, 26. xi. 2001,
AJepti ssappho astola Moore, l872
Llow land] Tb: 18L ,25. xii. 1998. Nd: 1cl a2,6. xii. 1998; 1c7 1\,26. xi. 2001.
[inlan dLkl: 1 8' ,30, xii, 1998.
Neptis, Moore, 1872"
nata adipata
[lewland]Nd: 18,29. 2001.
xi.
NeptisharitaharitaMoore,1875'

[lowland]Tb: 18,24. 1999.Nd: 19,29. 2001.
xii, xi.
NeptisIrylaksamarerpaMoore, 1874#

[inlanNdk:]1\,28. 2001.
xi,
IVlept miisah nolana H. Druce, 1874#
[inlan Ndkl: 14, 28. xi. 2001.
Lasippa tiga (Moore ,1858), ssp."
[lo wland] Nd: 1 8, 26. xi. 2001.
Ptintoporiahontoniahortlonia(Stol1l7,90)

[low land] Nd: 18, 26. xii. 1997; 1e, 30. xi. 2001. Th: 28, 25-27, xii, 1999.
[inlan dNk]: 1 g, 28. xi. 2001.
AthymaPravara indosinica(Fruhstorf1e9r06,)"

[low land] Nd: 18, 3, i ,2000; 18, 26, xi, 2001,
Athyma (Linnaeus1,758)
perius perius
[low land] Nd: 28, 30. xii. 1997,
[inlan dNh]: 1 g, 28. xii. 1998.
Athyma kanwa phorkys (Fruhstor 1f9e12r),*
[lowland]Tk: 1 27. 2001.
ci", xi.
Athyma tarymna (Fruhstor1f9e06r),"
siamensis
riow land ]Nd: 1 8, 26. xii. 1998.
Athyma selenophora gatijinsc i(aTalbo t19,36)
[low landJ Nd: 1 8, 2. i. 2000; 18" ,29. xi. 2001.
Linlan dN]k: 1\,28, xi. 2001.
Moduza pivcrisprocris(Cramer1,777)*

rlo wland] Nd; 19,3. i .2000;1\, 29. xi. 2001.
Lebadea martha martha (Fabrici u1s78,7)*
[low land] Nd: 18, 26. xii. 1998; 18, 25. xii, 1999; 1r, 26, xi. 2001. Th:18,1. i.
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2000, Tk: 1\,27. xii. 1998; 1 E7 1L 4, 26. xii, 1999; 1 gL ,27. xi. 2001.
Ptirtheno ssylvia gambrisius (Fabric i1u78s7,)
[low land] Tb: 1 8, 25, xii, 1998. Th: 1 l, 25. xii. 1.999.
Linlan dLkl: 1 8, 28. xii. 1999.
7innaeciajuliijuli(iLesson18,37)

[lo wland] Nd: 18 14, 27. xii, 1997; 18, 26. xii. 1998; 2S14, 2, i. 2000; 7g, 26-30,
xi. 2001. Th: 14,25. xii. 1999; 1c7'1\, 1. i .2000. Tk: 18, 28. xiL 1997; 16n ,26. xii.
I999; 2 4i,27 x.i. 2001.
[inlan Ndh]: 1r, 27.xii. 1997. Nk: 1\, 28. xi. 2001.
lhnaeciajah,nu (Moore,1858)'

[lo wland] Nd: 1?, 26. xii. 1998; 1\,2, i. 2000. Th: 2g, 25-27. xii. I999. Tk: 1?, 29.
xiL 19971 19, 31. xii. 1999.
Cynit itqapide acegnata Moore, 1897"
[lo wland] Nd: 18, 27. xii. 1997; 2r, 26. xii. 1998; 28, 2-3, i ,2000; l9, 29. xi. 2001.
Clynitia (Fmhstode1r9,13)"
cocytus ambrysus
[lo wland] Nd: 281 ?, 26. xii, 1998; 28, 2-3. i .2000; 1$, 26. xi. 2001. Th: 18, 1. i.
2000. Tk: 2 cli, 28-30, xii .1997; 1 r, 27. xii. 1998; t8, 31. xii. 1999,
Euthatia kesava(Moore,18S9)*
monina
[lo wland] Nd: 28, 30. xii. 1997; 18, 26. xii. 199g; 2829, 26-29. xi. 2001. Th: 181
g,25- 27, xii, 1999;1 \,26. xi, 2001. Tk: 19,31. xii. 1999.
Euthalia cijata Distant & Pryer ,1887, ssp,"
[low land] Tk: 1 oi, 27. xi. 2001.
Euthalia eriphylae tionel Firuhstorfer ,1905*
[low land] Nd: 1 8, 26. xii. 1998.
Euthaliateuta ira(Moore1,897)"

[lo wland] Tk: 1 I? - 29. xii, 1997.
,
Euthalia recta monilis (Moore 1,897)*
[lowland]Tk:28, 29. 1997.
xii,
Euthaliadunya dunya(Doubleday1,848)"

[lo wland] Nd: 1 g, 30. xii. 1997.
Euthali aanosia anosia (Moore ,1858) #
[inlan Ndk]: 1 8, 28, xi. 2001.
Lexias dirteabontouxi (VitaldiesSalvaza,1924)'

[lo wland] Nd: 1 8, 26, xii. 1998; 3g29, 29-30, xi. 2001. Tk: 19, 27, xi. 2001.
Lexiaspantalisjudeitina (Fruhstor1f91e3r),

[lo wland] Nd: 5Sl,30. xii. 1997; 18, 29, xi. 2001. Th: 18, 1. i. 2000, Tk: 1829,29.
xii. 1997; 1 8" ,31. xii. 1999.
[inlan Ldk]: 1\, 31. xii. 1998.
Lexias bonealisHanafusa, 199e*
albqpunctata
[Iow landl Nd: 28, 26-29, xi, 2001.
Chararcesbernant"s hierex(C.& R. Felder,1867)"

[low land] Tb: 18, 26, xii. 1997. Nd: 1g, 3, i. 2000.
Rohana tonkiniana sia}nensis (Fruhstor f1e9r13,)*
[low land] Tk: 1 8, 29. xii. 1997.
Eutaceura osteria (Westwood ,1850), ssp.*
[low land] Nd: 1g, 25. xii. 1999; 19,26. xi. 2001.
Euripus nyctetius e"pioeoides C. & R. Felder, 1 867
[Io wland ]Th: 1 8, 27. xii. 1999. Nd: 28, 3. i. 2000.
[inlan dNk]: 1 9, 28. xi. 2001.
Fblyura (C.& R. Felder,1867)"
aija ai:ia
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44 Masayuki NiMuRA and Hiroyuki WAKA}iARA
Llo wlandl Nd: 1 (l", 2. i. 2000; 2c;' ,26-29. xi. 2eOl. Tk:1 8X,31 .xii. 1999,
Pblyutu (Drury1,773)
athamas athamas
Llowland]･Nd:1cln3,0, 1997;1 3,i.2000;4,?i2,6-30. 2001. Tk:18n,28.
xiL c?, xi, xii.
1997.

LinlanNdh]: 1g,30, 1999.
xii,
Pbyura schreiber tisamenus (Fruhstor f1e9r1,4)*
[low land] Nd: 1 8, 29, xi. 2001.
Satyridae
Melanitistedotedu(Linnaeu1s7,58)

[lo wland] Nd: 1g1?,30. xii. 1997;1g'19,2-3.L 2000;18, 26. xi. 200]. Tk: 18,
27. 1998;48',26-31, 1999.
xii. xii.
[inlan dNh]: 1,ft 2,7. xii, 1999. Lk:1 8,31. xii. 1998; 18, 28. xii. 1999.
Ml71anitisphedima ganmpatiFruhstorfer,1908

[low land] Nd: 18, 30. xi. 2001, Tk: 18, 27, xi. 2001.
[inla nLdk:] 1g, 29. xii. 1999.
Etymnias h}tpermnestra Fruhstorfer,1902*
violetta
[low land ]Nd: 1l, 2, i. 2000;1r19-,26-29. xi, 2001. Tk:18, 28, xii. 1997.
Lethe mekara crtinana Fruhstorfe r1,91 1 *
[lowland]Th:18, 1.i.2eOO,

Erites deNic6ville1,893"
medura rotundata
[lo wland] Tb: 1 8a ,24, xii, 1999.
Otsoiriaena (Fabrici1u77s5,)"
med"s medus
tlow Iand] Tb: 18, 26. xiL 1997. Nd: 48, 26, xii. 1998;18, 29. xi, 2001.
Mycalesis mineus s"bj7isciata (Moore 1,882)"
L]ow land] Nd: 1 cln, 26. xii, 1998,
Mycagesis intennedia (Moore1,892)*

[lowlandlNd: 18,26. 2001,Tk: 18,29. 1997.
xi. xii.
Mycatesis visala visala Moore, 1858
[low land] Nd: 1 S, 27. xii. 1997; 2r, 25-26. xii. 1998.
[inlan dLk]: 18, 30. xii, 1998.
1())thimbaald"sbaldus(Fabriciu1s7,75)

[low land] Nd: 5r, 26-29, xi. 2001 ,
[inla nLdk:] 3 r, 28. xii. 1998.
Ragadia Riley & Godfrey, 1921"
critias
Llow land ]Nd: 2[?, 26. xiL 1998; 1cl" 2,5. xii. 1999; 18 l, 26-30. xi, 2001. Th: 2c]n 2,6,
2001.
xi.
Amathusiidae
Fbu"ni seume"s incerta (Stauding e1r8,87)*
[lo wland] Tk: 1X, 29. xii. 1997;58,31. xiL 1999. Th: 1g, 1, i .2000. Nd: l8, 26, xi.
2001.

fuunis canens arcesitas Stichel 1,933*
[lo wland] Nd: 281\, 27-30. xii. 1997; lr, 26, xii. 1998; 1g, 25. xii. 1999; 2S, 2. i.
2ooo; 4 g, 26-3o. xi. 2ool,
lhaumantis diores sptendens TYtler ,1939*
[low Iandl Nd: 39, 29-30, xi. 2001,
Amathuxidia amythaen amythaon (Doubleda y1 ,847)*
[lowland]Th: 19,27. 1999.
xii.
711tauri alatlr ysiiamensis Rothschild ,1916"
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TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan
Butterfii oefs the central Lao P.D.R. 45
[]owland]Tk: 18,27. 2001.
xi.
Discophora soncftzica zat Westwood, l851*1
[lo wland] Th:18, 1. i .2000. Tk: 4, 27. xi.
DiscophoratimoTu timorei Westwood
,1850*2001.
[Iow land] Th: 18, 1. i .2000,
Riodinidae
Zemerosfiegyas.flegy(aCsramer1,780)

[lo wland] Nd: 2 8, 30. xii. 1997;1 8, 26, xii, 1998; 1r, 3. i .2000.
[inlan Ndhl: 1 r, 27, xii. 1999.
Abisatu echerius (Stol l17,90) ,ssp.
Ilo wland] Tb: 19, 24, xii. 1999. Nd: 1\,3. i ,2000; 18, 29. xi. 2001. Th: 18, 26, xii.
1998. Tk: 14,27. xi. 2001,
[inlan Ldk]: 28, 30-31. xii. 1998.
Abisara bijitsci aMtoaore, 1877, ssp."
[lo wland] Tk: 1 clfi, 27. xi, 2001.
Abisara neophron chetina (Fruhstor 1f9e0r4), #
[inla nLdk]: 1 8, 29. xii. 1999.
Laxita th"isto ephorus (Fruhstor 1f9e0r4),"
[low landl Nd: 1 824, 2030. xi, 2001. Tk: 1?, 27. xi. 2001.
Tarcilahaq"inusberthaeFruhstorfbr,1904*

[lo wiand] Nd: 1r, 25. xii. 1999; 18 l?, 26-29. xi. 2001. Tk: 1 8, 27. xii.1998.
Taxita dora Fruhstorfe r1,904, ssp. #
[inlanLdk]:l4,30. 1998,Nk:19,28. 2001.
xii, xi.
Lycaenidae
POritiahevvitsonhiewitsoniMoore, 1866

[lowland]Nd: 19,29. 2001.Tk:18,29. 1997.
xi. xii,
rinlanLdk]: 1\,31. 1998.
xii.
Simiskina binghami(Fruhstor1f9e1r2),
phalia
[lowland]Nd: 1Si,26. 2001.Th:1EP,26. 2001,Tk:1･?,27. 1998.
xi. xi, xiL
[inlanNdk:]2?,28. 2001.
xi.
Miletus (Godfre1y9,13)*
ancon siamensis
[lowland]Th:1r, 1,i.2000.

Aglotinus (Moore1,884)"
suhstrigosus suhstrigosus
[lowland]Th:18, 25. 1999.Nd:38,3. i.2000.
xii.
Allotinus Ri]ey& Godfrey,]921'
unicotor rekkia
[lowland]Tb:1r,26. 1997.Tk:l8l?, 28-29. 1997.
xii. xii.
Altotin"s drumita aphtonius Fruhstorfer ,1914#
[inlanNdk]:29, 28. 2001.
xi.
Castatius rosimon rosimon (Fahrici u17s7,5)
[lo wland] Tb: 1r, 25. xii. 1998; 18, 24. xii. 1999. Nd: 18, 30. xii, 1997; 38, 26-30.
xi. 2001. Tk: 1 8,31. xii. 1999; 2 ,T, 27. xi. 2001.
[inla nNdh]: l8, 28, xii. 1998. Nk: 18, 28. xi. 2oo1.
Cageta (deNiceville1,897)*
roxus roxana
1lowland]Nd: 2r, 3,i.2000;18,26. 2001.
xi,
Ctileta (Fruhstor1f9e1r8),
elna noliteia
[lowland]Tb:1\,25.xii.1998;18,25. 2001.Nd: 18,3.i.2eOO.
xi.
[inlan dNh]: 8,30. xii. 1999.
Zzeeria karsandra (Moore,1865) #

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