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PUBLICATIONS OF THE NATIONAL BOTANICAL INSTITUTE, PRETORIA PUBLIKASIES VAN DIE NASIONALE BOTANIESE INSTITUUT, PRETORIA Obtainable from the National Botanical Institute, Private Bag X101, Verkrygbaar van die Nasionale Botaniese Instituut, Privaatsak X101, Pretoria0001, Republic ofSouth Africa. A catalogue ofall available Pretoria0001,RepubliekvanSuid-Afrika. ’nKatalogusvanallebeskik- publications will be issued on request. barepublikasies kanaangevra word. BOTHALIA Bothalia is named in honour ofGeneral Louis Botha, first Premier Bothalia is vernoem ter ere van Generaal Louis Botha, eerste Eerste and Minister of Agriculture of the Union of South Africa. This MinisterenMinistervanLandbouvandieUnievanSuid-Afrika.Hierdie housejournaloftheNationalBotanicalInstitute,Pretoria,isdevotedto lyfbladvandieNasionaleBotanieseInstituut,Pretoria,isgewyaandie the furtherance of botanical science. The main fields covered are bevorderingvandiewetenskapvanplantkunde.Diehoofgebiedewatgedek taxonomy, ecology, anatomy and cytology. Two parts ofthe journal word,istaksonomie,ekologie,anatomieensitologie.Tweedelevandie and an index to contents, authors and subjects are published tydskrifen ’n indeks vandie inhoud, outeursen onderwerpeverskyn annually. jaarliks. MEMOIRS OF THE BOTANICAL SURVEY OF SOUTH AFRICA MEMOIRS VAN DIE BOTANIESE OPNAME VAN SUID-AFRIKA The memoirs are individual treatises usually ofan ecological nature, ’n Reeks van losstaande omvattende verhandelings oor vernaamlik but sometimes dealing with taxonomy oreconomic botany. ekologiese,maarsomsooktaksonomieseofplantekonomieseonderwerpe. THE FLOWERING PLANTS OF AFRICA DIE BLOMPLANTE VAN AFRIKA / HierdiereeksbiedkleurplatevanAfrikaanseplantemetbygaandeteks. ThisserialpresentscolourplatesofAfricanplantswithaccompanying Die skilderye word meestal deur die kunstenaars van die Nasionale text.TheplatesarepreparedmainlybytheartistsattheNationalBotanical BotanieseInstituutvoorberei. Tallebekendebotaniesekunstenaarshet Institute. Manywellknownbotanicalartistshavecontributedtotheseries, totdiereeksbygedra,soosCythnaLetty(meeras700plate),Kathleen suchasCythnaLetty(over700plates),KathleenLansdell,StellaGower, Lansdell, StellaGower, BettyConnell, PeterBallyenFay Anderson. BettyConnell,PeterBallyandFayAnderson. TheEditorispleasedto Die Redakteur verwelkom lewende plante van algemene belang of receive living plants of general interest or of economic value for ekonomiese waardevirafbeelding. illustration. Vanaf Vol. 50 word een deel, bestaande uit twintig plate, jaarliks FromVol.50,onepartoftwentyplatesispublishedannually.Avolume gepubliseer. ’nVolumebestaanuittweedele.Diepublikasieisbeskikbaar consistsoftwoparts.ThepublicationisavailableinEnglishandAfrikaans. in Afrikaansen Engels. FLORA OF SOUTHERN AFRICA FLORA VAN SUIDELIKE AFRIKA / A taxonomic treatise on the flora ofthe Republic of South Africa, ’nTaksonomieseverhandelingoordiefloravandieRepubliekvanSuid- Ciskei, Transkei, Lesotho, Swaziland, Bophuthatswana, Namibia, Afrika,Ciskei,Transkei,Lesotho,Swaziland,Bophuthatswana,Namibie, BotswanaandVenda.TheFSAcontainsdescriptionsoffamilies,genera, BotswanaenVenda. DieFSAbevatbeskrywingsvanfamilies,genusse, species,infraspecifictaxa,keystogeneraandspecies,synonymy,literature spesies,infraspesifieketaksons,sleutelstotgenusseenspesies,sinonimie, and limited specimen citations, as well as taxonomic and ecological literatuur,verwysingsnaenkeleeksemplare,asookbeknoptetaksonomiese notes. en ekologieseaantekeninge. PALAEOFLORA OF SOUTHERN AFRICA PALAEOFLORA VAN SUIDELIKE AFRIKA / A palaeoflora on a pattern comparable to that of the Flora of ’nPalaeofloramet’nuitlegvergelykbaarmetdievandieFloravansuide- southern Africa. Much of the information is presented in the form likeAfrika.Baievandieinligtingwordaangebiedindievormvantabelle of tables and photographic plates depicting fossil populations. Now enfotografieseplatewaaropfossielepopulasiesafgebeeldword. Reeds available: beskikbaar: MMoolltteennoo FFoorrmmaattiioonn((TTrriiaassssiicc))VVooll.. 12..IGntyrmonduocstpioenr.msDic(reoxiclduiduimn,gbDyi/cdreouirdiJu.mM).,&byH/d.eMu.r JA.nMd.er&sonH..M. Anderson. Prodromus of South African Megafloras. Devonian to Lower Cretaceous, by/deur J.M. & H.M. Anderson. Obtainable from/Beskikbaar van: A.A. Balkema Marketing, Box/Posbus 317, Claremont 7735, RSA. 1 n /9 / 1(0 BOTHALIA ’N TYDSKRIF VIR PLANTKUNDIGE NAVORSING A JOURNAL OF BOTANICAL RESEARCH Volume 21,2 Editor/Redakteur: O.A. Leistner Assisted by B.A. Momberg Editorial Board/Redaksieraad D.F. Cutler Royal Botanic Gardens, Kew, UK B.J. Huntley National Botanical Institute, Cape Town, RSA P.H. Raven Missouri Botanical Garden, St Louis, USA J.R Rourke Compton Herbarium, NBI, Cape Town, RSA M.J. Werger University of Utrecht, Utrecht, Netherlands Editorial Committee Redaksiekomitee O.A. Leistner B.A. Momberg M.C. Rutherford ISSN 0006 8241 Issued by the National Botanical Institute, Private Bag X101, Pretoria 0001, South Africa Uitgegee deur die Nasionale Botaniese Instituut, Privaatsak X101, Pretoria 0001, Suid-Afrika 1991 — CONTENTS INHOUD Volume 21,2 1. Kamiesbergia, a new monotypic genus ofthe Amaryllideae-Strumariinae (Amaryllidaceae) from the north-western Cape. D.A. SNIJMAN 125 2. Studies in the genus Riccia (Marchantiales) from southern Africa. 23. R. bullosa: typification and a full description. S.M. PEROLD 129 3. Studies inthe Ericoideae (Ericaceae). VIII. New species in Erica, sectionPseuderemia from southern Africa. E.G.H. OLIVER and INGE M. OLIVER 137 4. Cyanophyceae associated with mangrove trees at Inhaca Island, Mozambique. S.M.F. SILVA 143 5. Notes on African plants: Asteraceae. New species in the genus Disparago. M. KOEKEMOER 158 Fabaceae. Caesalpiniabracteata, anew species fromthe Onseepkansareaofthenorthern Cape Province. G. GERMISHUIZEN 152 Geraniaceae. Thecorrectauthercitation forPelargonium section Cortusina. L.L. DREYERand J.J.A. VAN DER WALT 157 Lichenes. Two new species in the Agyriaceae (lichenized Ascomycotina, Lecanorales) from southern Africa. F. BRUSSE 154 Liliaceae (Asphodelaceae). The type specimen of Aloe soutpansbergensis. H.F. GLEN and HARDY D.S. 151 Liliaceae. Notes on some Monocotyledoneae described by N.L. Burman, Asparagaceae, Asphodelaceae, Colchicaceae. D.O. WIJNANDS 157 Poaceae. On the origin of Eleusine africana. S.C. HIREMATH and S.S. SALIMATH 161 6. Basic chromosome numbers and polyploid levels in some South African and Australian grasses (Poaceae). J.J. SPIES, E. VAN DER MERWE, H. DU PLESSIS and E.J.L. SAAYMAN .... 163 7. The chromosomes of Chortolirion and Poellnitzia (Asphodelaceae: Alooideae). G.F. SMITH 171 8. Invasive alien woody plants of the northern Cape. L. HENDERSON 177 9. New taxa, new records and name changes for southern African plants. B.C. DE WET, R. ARCHER, L. FISH, G. GERMISHUIZEN, P.P. HERMAN, M. JORDAAN, S.M. PEROLD, C. REID, J. VAN ROOY, W.G. WELMAN and H.F. GLEN 191 10. National Botanical Institute, South Africa: list of staff and publications 215 11. Book reviews 227 Digitized by the Internet Archive 2016 in https://archive.org/details/bothaliavolume2121unse_0 Bothalia 21,2: 125-128 (1991) Kamiesbergia a new monotypic genus ofthe Amaryllideae-Strumariinae , (Amaryllidaceae) from the north-western Cape D.A. SNIJMAN* Keywords: Amaryllidaceae, Amaryllideae, new genus, north-western Cape, South Africa ABSTRACT Kamiesbergia Snijman isanew monotypic genus from raised graniteoutcrops inthe north-westernCape. A member ofthe subtribe Strumariinaeofthe Amaryllideae, itis mostclosely related toHessea Herb, and Namaquanula D. & U. Muller-Doblies. Thedissimilarinnerandouterstamens,theuniquelyclub-shapedinnerfilamentsandthenovel insertion ofthe filamentintheproximalquarteroftheantherconnective are the mainapomorphiesofthegenus. The numberof rare and monotypicgeneraofAmaryllidaceae in this region iscomparable to thatofAndean South America. UITTREKSEL KamiesbergiaSnijmanis ’nnuwemonotipiesegenusvandiehoerliggendegranietriwweindieNoordwesKaap. Ditis 'n lidvandie subtribus Strumariinaevan die Amaryllideaeen isdie naaste verwantaan Hessea Herb, enNamaquanula D.&U.Muller-Doblies.Dieverskilletussendiebinnesteenbuitenstemeeldrade,dieuniekeknuppelvormigebinnestehelmdrade endieongewoneaanhegtingvandiehelmdraadaandieproksimalekwartvandiehelmbindselisdiehoofapomorfevandie genus.DieaantalskaarsenmonotipiesegenussevandieAmaryllidaceaeinhierdiestreekkanvergelykwordmet’nsoortge- lyke verskynsel in die Andes-gebergtes van Suid-Amerika. INTRODUCTION Since this new species lacks all the derived character states of the genera already described within the Stru- The tribe Amaryllideae sensu Traub (1965, 1970) and mariinae, andsinceitpossessesasetofuniquecharacters Dahlgren etal. (1985) is uniquely definedby bisulculate thatjustifies its separation atthe generic level, the taxon pollengrains(Dahlgren&Clifford 1982; Dahlgrenetal. is described here as the new monotypic genus Kamies- 1985; Erdtman 1966; Schulze 1984). Aspresentlycircum- bergia. scribed (Muller-Doblies 1985), thesubtribe Strumariinae oftheAmaryllideaecurrentlyincludesfoursmallandthree MATERIALS AND METHODS monotypicgenera NamaquanulaD. &U. Muller-Doblies, HesseaHerb., Car(polyzaSalisb.,StrumariaJacq.,Bokke- ThisstudywasbasedonherbariummaterialfromBM, veldia D. & U. Muller-Doblies, Gemmaria Salisb. and BOL, K, NBG, PRE, SAM and WIND. Ecological Tedingea D. & U. Muller-Doblies). Character states information was derived from field studies during the common to these genera are the actinomorphic flowers flowering, fruitingandleafingstagesofthespecies. Fresh and the reduced size ofthe plants. Exclusively southern non-acetolyzedpollengrainsforscanningelectronmicro- African,thesubtribecomprisesapproximately35 species, scopy (SEM) were dehydrated in ethanol, critical point which are concentrated in the semi-arid winter rainfall driedandcoatedwithAu/Pd. Photographsweretakenwith regions of the Cape Province and southern Namibia. aCambridge 200 SEM at 10kv. Chromosomedatawere gatheredfromactively growingroottips, pretreatedwith Phylogenetic studies in the Strumariinae (Snijman saturatedalphabromonapthaleneat4°C for24hours,then unpubl.), revealed an undescribed Hessea-like species fixedin 1:3aceticacid/ethanol. Theroottipswerehydro- from the north-western Cape. A character analysis lysedinnormalhydrochloricacidat60°Cforsixminutes, confirmed the presence ofbisulculate pollen grains but stainedwithFeulgenfor30minutes, counterstainedwith indicatedthatthe taxon did not fit into any known genus 2% aceto-orcein and then squashed. Photographs were ofthe Strumariinae. The new species lacks the synapo- taken with a Zeiss Axioskop. morphy ofx = 10 and the adnation ofthe filaments to tithsecsltoyslee,awllhiieschBockhkaervaecltdeirais,esGeCmarmpaorliyzaa,anSdtrTuemdairnigeaaa;nidt nov.,KeaxmiafefsinbietratgeigaesnteernuomsiNpahmoanqSunainjumlaan,D.ge&n.Un.ovM.uletlesrp-. lacks the centrifixed anther insertion which is synapo- Doblies, Hessea Herb., Strumaria Jacq. sensu lato et morphicforHessea; andiswithouttheadaxially-hooked CarpolyzaSalisb. sedcum nullogeneresatiscongruit: ab filaments which characterise Namaquanula. Character CarpolyzaStrumariaqueabsentiaulliuscommisuraeinter statesuniquetothenewtaxonarethedissimilarinnerand stylum et filamenta recedit; ab Namaquanula absentia outer stamens, the club-shaped distal half of the inner hamuliadaxialisfilamentorumdiffert;abHesseaabsentia filaments, and the filament attachment near to the base antherae centrifixae differt. Ab omnibus his generibus ofthe anther connective. filamentis dissimillimis, filamentis interioribus clavifor- mibus supra medium, antheris sub-basifixis differt. — TYPE. Cape, 3018 (Kamiesberg): Kamiesberg, near C*lCaoremmpotnotn7H7e35r.barium, National Botanical Institute, Private Bag X7, Karas on E slopes ofRooiberg, (-AC), 28-4-1988, Snij- MS. received: 1991-03-08. man 1175 (NBG, holo.; K, PRE). 1 126 Bothalia21,2 (1991) i nin 111 1 FIGURE 1.—Kamiesbergiastenosiphon. 1,inflorescence;2,bulbandleaves;2a,'ransversesectionofleaf;3,wholeflower;4,sectionofflower; 5,stigma;6,antherattachmentofinnerstamens,ventralview;6a, lateralview;6b,dorsal view;6c, sectionshowingsub-basifixedinser- tion,connectivehatched; 7,antherattachmentofouterstamens, ventralview;7a, lateralview;7b,dorsal view;7c, sectionshowingsub- basifixed insertion, connective hatched; 8, capsule and seeds (drawn from Snijman 1175). Bothalia 21.2 (1991) 127 mm Bulbous perennial herb, up to 210 tall. Bulb soli- ofKamiesbergia are attached in the proximal quarter of tary,deep-seated, subglobose, 12—30mmacross, withthin the connective (Figure 1.6c & 1.7c). light-brownparchment-likeoutertunics, fleshyandwhitish within, extended into a long slender neck up to 110 mm. The presence of a perigone tube is considered to be Leavesabsentatanthesisor rarely persisting toanthesis, plesiomorphic in the subtribe (Miiller-Doblies 1985). In 2(—3), spreading, narrowly lorate, 60-300 x 1-3 mm, most species ofHessea the perigone tube is reduced and glabrous, withtheadaxial surfaceshallowlycanaliculate, the tubular component is formed by the extension of subtended by a subterranean non-amplexicaul prophyll. the perigone/stamen confluence into a winged tube, a Inflorescenceslightlyspreading, 40—80mmacross; scape featurewhichisconsideredtobederived(Miiller-Doblies stiffly erect, 50—140 mm long, 1—2 mm diam., greyish 1985). Only H. longituba D. & U. Miiller-Doblies has a pinktoleaden-grey,breakingoffatthebaseinfruit; spathe pronounced smooth perigone tube comparable to that of valves linear-lanceolate, 15-30 X 1—2 mm; bracteoles K. stenosiphon which may suggest a close relationship filiform, ap to 5 mm long. Flowers 5—9, ascending, between the tw,o taxa. A critical evaluation of the key hypocrateriform, pale lemon-yellow, usually with a androecial states in Hessea and K. stenosiphon however, greenish to reddish brown tube, flushed dorsally with does not support such a relationship. reddishbrownontheoutertepals, ageing to lightbrown, scentless;pedicelsstraighttoupwardlycurved, 35-60mm The main synapomorphy forHessea is the insertionof long, green. Tepals recurving from a narrow 8—12 mm thefilament intoaconnectivesheath, inwhichthe length long tube, narrowly lanceolate, 6—8 x 2—3 mm, slight- ofthe dorsal wall almost equals that ofthe ventral wall: ly channelled. Stamens in 2 unequal whorls, epi- aconditionknownascentrifixed antherinsertion(Miiller- tepalous, filamentsbasally connateintoagreenishyellow Doblies 1985). This character state is interpreted as the tubeextending to0,5—1,0 mmabove theperigone throat, extremeinamorphologicalseriesrangingfromdorsifixed free above, reduced to a 0,25 mm long filiform free tip to subcentrifixed and centrifixed anther insertion, with intheouterwhorl, prominentand spreading in the inner each state being found in the Strumariinae. In contrast, whorl; the inner filaments free for 3—4 mm, clavate in a transformation series between the centrifixed anther the distal half with a subulate tip; anthers dorsifixed insertion of Hessea and the almost basifixed anther nearthebase, ±2,5 mmlongandmaroonbeforeopening; insertion(withoutaconnectivesheath)ofK. stenosiphon the outer anthers occluding the perigone throat after could notbe established fromcurrent data. Thusa sister dehiscence; pollen cream-coloured. Ovary subglobose, group relationship between the two genera cannot be 2—3 mmacross, with up to4—6ovules per locule. Style inferred. erect, slenderthroughout, upto5—11 mmlong, remaining includedintheperigonetube; stigmashortlytrifid, shortly Thebisulculatepollengrains areglobose, isopolarand penicillateontheinnersurface.Fruitasubglobose, papery, have scattered large spinulae on the surface (Figure 2). loculicidal capsule, 7,5 mm across. Seeds fleshy, ovoid, The karyotype ofK. stenosiphon (Figure 3) comprises a upto2,5 mmacross, reddishbrownwhenripe. Chromo- pair oflarge submetacentric chromosomes; six pairs of some number. 2n = 22. Figure 1. medium-sized metacentric to submetacentric chromo- somes, ofwhichonepairis asatellitechromosome; and Flowering time extends from the end ofApril to May. four pairs of shorter metacentric to submetacentric Inmostbulbsthefoliageleavescommencegrowthshortly chromosomes. The chromosome phenotypes are similar after floweringand subsequently die backat the onsetof tothoseofmostAmaryllideaewithx = 11 (Goldblatt 1972, the summer drought. Occasionally some bulbs which 1976;Jones&Smith 1967). Akaryotypeofx = 11 iscon- occupy wetter, cooler sites have been noted with green sideredbasicinthefamily (Inariyama 1937; Meerow 1984; leaves throughout the summer. Sato 1938). Diagnosticfeatures ofK. stenosiphon are the form of DistributionandhabitatrecordsindicatethatK. stenosi- thestamensandantherinsertion. The innerfilamentsare phonisrare. Knownpopulationsarerestrictedtothenorth- at least twelve times as long as the outer filaments and western Cape and are widely disjunct in the eastern areuniquelyclub-shaped inthedistal halfwithasubulate Kamiesberg and near Pofadder (Figure 4). In the tip. The stamens of the short outer whorl occlude the Kamiesberg, the deep-seated bulbs grow in loamy soils perigone throat after dehiscence. Unlike the medifixed which accumulate in seasonally moist crevices and anthers common to all other Strumariinae, the filaments water-worngulliesonmassiveexposedgranitedomes, at FIGURE2.—Bisulculatepollengrains ofKamiesbergiastenosiphon, Snijman 1175. A, equatorial view,longitudinalposition;B, equatorial view, transverse position.Scalebars =20pm. 128 Bothalia21,2 (1991) ThediscoveryofKamiesbergia, afurthermonotypicgenus intheAmaryllideae, bringsintofocustheparallelswhich exist in this respect between the Amaryllideae of South Africaandthe Andean AmaryllidaceaeofSouth Ameri- ca. Goldblatt’s suggestion (Meerow 1985) that similar evolutionarypatternsexistbetweentheAmaryllidaceaeof these two geographical centres is yet to be examined. CAPE.—2919(Pofadder): Numis [Namies], nearPofadder, (—AC), Lavranos20311(PRE). 3018(Kamiesberg): Kamiesberg,nearKarason EslopesofRooiberg,(—AC),Snijman1175(K,NBG,PRE);nearKaras on E slopes ofRooiberg, (—AC), Snijman 1179(NBG). ACKNOWLEDGEMENTS IwishtothankDrJ.C. Manningforpreparingthepollen micrographs, Mrs J.E. Ward-Hilhorst for the botanical illustrations. DrH.P. Linderforhelpfulcommentsonthe FIGURE3.—MitoticmetaphaseinKamiesbergiastenosiphon,Snijman manuscript and Dr J.C. Paterson-Jones for assistance in 1175, 2n = 22, x 665; the arrow indicates the satellitechro- the field. mosome. REFERENCES DAHLGREN, R.M. &CLIFFORD,H.T. 1982. Themonocotyledons', 12 14 16 18v-2L0 L 2.2 24 6 28 ^30Tij32i 3r4 r DAHLGaREcoNm,paRr.aNt.i,veCLstIuFdyF.OARcDa,deHm.Ti.c&PreYsEsO,,LoP.nFd.o1n9.85. Thefamilies 18-Hix a Trji-p18 ofthemonocotyledons. Springer-Verlag, Berlin. t ERDTMANN,G. 1966.Pollenmorphologyandplanttaxonomy.Hafner, 20tftii I —11 PIin (Ao 20 GOLDBNLeAwTTY,orPk..1972.Chromosomecytologyinrelationtoclassifica- 22 i22 tioninNerineandBrunsvigia(Amaryllidaceae).JournalofSouth 24 U —f7 r^tTTtTfe1r~\/ni24 GOLDBLACfaArrTipcToal,nyzPB.aot1(a9A7nm6ya.rC3y8hllr:io2dm6a1oc-es2ao7em)5e..cAyntnoallosgyoofftHheesMsiesas,oSutrriuBmoatrainaiacnadl 26 \l—r 26 Garden 63: 314-320. INARIYAMA,S. 1937.KaryotypestudiesinAmaryllidaceae1.Science 28 • 28 JONES,ReKp.or&tSTMokIyToHB,unJr.Bi.ka19D6a7.igCahkruomseocstioomneBev3o,lu5t2i:on95i—n1t1h3e.genus 32—i—t—\ i p -1VV- V J 27 TTilflf3302 MMEEEERROOHCWWer,,ribneuAArm..t.WWi..aCa411r9098:y85o4.l1.o3Ag9Ki—ana1re5y2w40o.:tsyppe1ec6i3e—es1v7oo9lfu.tEiuocnroisniathaendAmaarnyelwlindaacmeeaei.n —— Stenomesson (Amaryllidaceae). Brittonia 37: 305-309. 34\10 \ 12 14 1|6 1 18 20 22 4 26 26 _ 30 312 1 3J4 (Ti3U6wT MEEROMWa,thiAe.uWa. K1l9o87t.zsTchheaniddenPtliatgieisolairnidonsyBsatkemearti(cAmraerlyaltlioindsahciepase)o.f FIGURE 4.—The known geographical distribution ofKamiesbergia Taxon 36: 566-572. stenosiphon. MiiLLER-DOBLIES, D. & U. 1985. De Liliifloris notulae 2. De taxonomiasubtribusStrumariinae(Amaryllidaceae).Botanische JahrbiicherfurSystematik 107: 17—47. elevationsofapproximately610m. Dwarfsucculentshrubs SATO, D. 1938. KaryotypeevolutionandphylogenyIV. Karyotypesin andthefernspeciesCheilanthesmultifida(Swartz)Swartz AmaryllidaceaewithspecialreferencetotheSAT-chromosome. constitutetheassociatedplantcommunity. Kamiesbergia Cytologia9: 203—242. SCHULZE,W. 1984.BeitragezurTaxonomiederLiliiflorenXTV.Der stenosiphon occupies similar habitats at Namies near Umfang der Amaryllidaceae. Wissenschaftliche Zeitschrift, Pofadder, onlargegraniteoutcropsamidthesandyplains Friedrich-Schiller Universitat, Jena, Math.-Naturwiss. 32: ofBushmanland (J.J. Lavranos pers. comm.). 985-1003. TRAUB, H.P. 1965. Addendato Traub’s ‘Thegeneraofthe Amaryl- Meerow (1987) has drawn attention to the number of TRAUB,lidHa.cPe.ae1’97(01.96A3n).inPtlraondtucLtiifeon2t1:oH8e8r,be8r9t.’s‘Amaryllidaceae,etc.’ small or rare monotypic genera amongst the pancratioid 1837andrelatedworks.InW.Herbert,Amaryllidaceae.Reprint AmaryllidaceaeinthenorthernAndeanofSouthAmerica. J. Cramer, Lehre.

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