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Birds of Fazao-Malfakassa National Park, including the first record of White-browed Forest Flycatcher Fraseria cinerascens PDF

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Birds of Fazao-Malfakassa National Park, including the first record for Togo of White-browed Forest Flycatcher Fraseria cinerascens PaulM. Radleya andGenevieve Campbell Les oiseauxdu ParcNational de Fazao-Malfakassa, incluantlapremiere mention pourle Togo du Gobemouche a sourcils blancs Fraseria cinerascens. Le Parc National de Fazao-Malfakassa (08°40’N 00°43’E) comporte unevegetation variee, allant de laforet dense alasavane. Bien que le pare soit une Zone d’Importance pour la Conservation des Oiseaux, peu de donnees sont disponibles sur les especes d’oiseaux presentes. En six semaines de travail sur le terrain, du 27 octobre au 13 decembre 2003, nous y avons recense 199 especes d’oiseaux, dont une, le GobemoucheasourcilsblancsFraseriacinerascens n’avaitpasencoreetesignaleeauTogo. Letotal , des especes d’oiseaux recensees jusqu’a ce jour dans le pare de Fazao-Malfakassa est de 242 especes. Summary. Fazao-MalfakassaNational Park (08°40’N 00°43’E) has avariedvegetation, from for- esttosavanna. Despitethattheparkis an Important BirdArea, fewdataonits birds areavailable. During six weeks offield work, between 27 October and 13 December 2003, we recorded 199 birdspecies, amongstthemone,White-browedForestFlycatcherFraseriacinerascens, thathadnot been recorded previously in Togo. Our work brings the total number ofspecies known from Fazao-Malfakassa National Park to 242. Covering 192,000 ha, between 300 and 800 m the local human populations, these buffer zones in elevation, Fazao-Malfakassa National Park are considered within park boundaries and juris- (08°40’N 00°43’E) is the largest ofTogo’s three diction, and species observed in these areas were national parks. It is located in the transition zone included in our list. between forest (Upper Guinea Forest block) and ThedryseasoninTogo isNovember-February savanna, and comprises the most extensive area of and the rainy season May-September, peaking in relatively undisturbed vegetation in the country August (Cheke & Walsh 1996). The typical (Cheke 2001). north-south pattern of rainfall in West Africa is With relatively low rainfall (compared to disrupted by the effects ofthe Togo and Atakora neighbouring Ghana), resulting in primarilyxero- Mountains, resulting in higher average rainfall in phytic vegetation, Togo almost completely lacks central Togo compared to the southern, coastal tropical forest, and savanna (primarily grassland, zone (Cheke & Walsh 1996). For example, mean wooded grassland, and woodland) extends almost annualrainfallforSokode (08°59’N), c.20kmeast to the coast (Jenik 1984, Cheke & Walsh 1996, of park, is 1,441 mm, whereas the mean for mm & Cheke 2001). Habitats within Fazao-Malfakassa coastal Lome (06°08’N) is 924 (Cheke National Park are varied, but consist primarily of Walsh 1996). Guinea savanna and woodland, semi-evergreen Although designated an Important Bird Area galleryforestand forestpatches, andgrass-covered (Cheke 2001), few bird surveys have been under- hilltops (Figs. 1-3; Cheke 2001). The park’s ter- taken inFazao-MalfakassaNationalPark, themost & rain is rugged, including the Monts de Malfakassa recent in March-April 1984 (Cheke Walsh inthe north, the centralMonts du Fazao, andpre- 1996). Herewe present the results ofan avifaunal cipitous rock and cliff faces associated with the survey conducted at the onset ofthe dry season, Falaise de Boulowou at the western edge (Cheke between 27 October and 13 December 2005. We 2001). Buffer zones at the park borders comprise also listadditionalbirdspecies previouslyrecorded moderately to heavily cultivated areas ofgrassland byotherobservers butnotbyus, to produce apre- andwoodland, andwetlands and rivers fringed by liminary bird list ofthe park. dense reedbeds (Fig. 4). Createdas aconcession to BirdsofFazao-MalfakassaNationalPark, Togo:Radley&Campbell BullABCVol15No2(2008)-203 Methods not figure on the report’s list (Dowsett-Lemaire & We endeavoured to cover as much of the park’s Dowsett 2007). As Latham’s Forest Francolin was areaas possible, usingfourvillages atits borders as claimed from Fazao by an observer of unknown starting points for our survey: Fazao (08°4l’N experience andis notyetdocumentedfrom neigh- 00°46’E), Boulohou (08°46’N 00°40’E), bouring eastern Ghana, its occurrence is best & Tchitchako (08°34’N 00°37’E, includingastretch treated as unconfirmed (Dowsett-Lemaire along the Koue River, 08°30’N 00°37’E, at the Dowsett 2007), although a specimen was collect- Togo/Ghana border), and Bounako (09°10’N ed just south of the park, at Bismarckburg, by 00°53’E). We also camped beside the Kamassi Reichenow in 1891 (Cheke &Walsh 1996). River east of Boulohou (08°47’N 00°43’E) and This brings the total number of bird species near Mt. Kpeya (08°48’N 00°49’E), both well recorded from the park to 242, or 39% of the & within the park boundaries. As time was limited, species documented for Togo (Cheke Walsh wewere unable tovisit the southern section ofthe 1996).This is indicative ofthe relative qualityand park (between 08°30’N and 08°20’N), which is importance of the park to the conservation of more extensivelyforested. Togo’s birds andshouldserveas impetus to its fur- Surveys consisted of walking slowly through ther, and increased, protection. Twenty-four representative habitat (i.e., those areas representa- species restricted to the Sudan-Guinea Savanna tive of the majority of the park), stopping at biome were recorded in the park, along with an locations of high bird activity, and recording all equal number from the Guinea-Congo Forests species detectedvisuallyoraurally. Wepaidspecial biome (Appendices 1 and 2). It should be noted attention to globally threatened or rare species. that the adjacent, proposed Kyabobo National Several nocturnal surveys were conducted by lis- Park in Ghana, holds nearly three times the tening from fixed points and/or bywalking trails. Guinea-Congolian forest species of Fazao- Borrow & Demey (2001, 2004), whose Malfakassa National Park (Dowsett-Lemaire & nomenclature, taxonomyandsequenceis followed Dowsett 2007). here, was used for field identification, Chappuis As dataconcerningmigrants inWesternAfrica & & (2000) to identify vocalisations, and Cheke are frequently lacking (R. J. Dowsett F. Walsh (1996) for information on range and status Dowsett-Lemaire pers. comm.), Appendix 1 also inTogo. includes the dates of first detection for both Palearctic and intra-African migrants (following Results and Discussion Borrow & Demey2001, 2004) observedat Fazao- During approximately sixweeks offield work, we MalfakassaNational Park. Dates givenare onlyfor recorded 199 bird species, one ofwhich, White- migrants that are not considered annual residents browed Forest Flycatcher Fraseria cinerascens is in the area ofthe park (Borrow & Demey 2001, , newforTogo.ThesearelistedinAppendix 1,with 2004). It should be noted, however, that in some their relative abundance and biome in which they cases dates given might also be an affect ofwhen were observed, where applicable. A list of 43 specific habitats were surveyed during our work, species (some requiring confirmation [e.g., and are not necessarily indicative of a given MEPFT 2001]) previously recorded in the park species’ arrival or occurrence. but not encountered by us, formsAppendix 2. The present total should be considered provi- & Cheke Walsh (1996) cite four additional sional, as oursurveywas conducted at the onsetof species for the park that we have excluded from the dry season and did not cover the entire park. Appendix 2: Latham’s Forest Francolin Francolinus An increasing number of Palearctic and intra- lathami. Blue-headed Crested Flycatcher African migrantswere detected towards the endof Trochocercus nitens Dusky-blue Flycatcher our visit, in early December. Therefore, survey , Muscicapa comitata and Sooty Boubou Laniarius work further into the dry season (January-April) leucorhynchus. The latter three are based upon an and during the rainy season should yield some unpublished report (cited as ‘Minster Agriculture additional species. Likewise, for a more accurate Limited 1984’). Dusky-blue Flycatcher and Sooty overall species profile, surveys should certainly be Boubou have since been deemed doubtful or undertaken in the southern, more heavilyforested wrong, and Blue-headed Crested Flycatcher does portions ofthe park, where up to 40 not yet doc- & 204-BullABCVol15No2(2008) BirdsofFazao-MalfakassaNationalPark, Togo:Radley Campbell lamented forest species could potentially occur ident in Togo, previously known only from two & & (Dowsett-Lemaire Dowsett 2007). specimens (Cheke Walsh 1996). Also docu- mented in Kyobobo National Park, eastern Ghana & Notes on selected species (Dowsett-Lemaire Dowsett 2007). White-crested Tiger Heron Tigriornis leucolopha Abyssinian Ground Hornbill Bucorvusabyssinicus One flushed from cover along the Kamassi River Apair in wooded savanna c.10 km north ofFazao (atc.08°47’N00°43’E) on30November. Listedas on 10 December. Rare resident, in Togo now & i rare resident by Cheke Walsh (1996), with almost entirely restricted to Fazao-Malfakassa and & anly one recent record, oftwo individuals, along Keran National Parks (Cheke Walsh 1996). the Koue River, in the Fazao Mountains, in 1988. Also known from Kyobobo National Park, eastern Double-toothed Barbet Lybius bidentatusand & Ghana (Dowsett-Lemaire Dowsett 2007). Bearded Barbet Lybiusdubius One L. bidentatus calling in a wooded, semi- BlackStork Ciconia nigra cultivated areaalongasmall stream near the Koue One flushed from cover along the Loukoulou River, at the south-western boundary ofthe park, River, in wooded savanna south-east of Mt. on 24 November. One L. dubius observed calling ECpeya, on 12 December. Considered a rare in open woodland on the north bank ofthe Mo Palearctic migrant to Togo, where generally River, at the north-eastern boundary ofthe park, recorded from protected areas in the northern on 3 December. Unlike reports of these two & & savannas ofthe country (Cheke Walsh 1996). species by Dowsett-Lemaire Dowsett (2007) at Kyobobo National Park, we did not observe them GlossyIbis Plegadisfalcinellus in close proximity to one another, but at the One in flight along a river near Bounako on 3 northern (L. dubius) and southern L bidentatus) ( . DecemberandanotheralongtheLoukoulouRiver extremities ofour survey area. lear Mt. Kpeya on 11 December. Listed as a rare & Palearctic migrant to Togo by Cheke Walsh Western OlivaceousWarblerHippolais opaca (1996). One seen inwoodedhabitat near Boulohouon 16 November. Earliest record of this uncommon & Bateleur Terathopius ecaudatus Palearctic migrant in Togo: Cheke Walsh \nadultmaleandajuvenileobservedinthenorth (1996) give 23 November as the earliest date. af the park, several km south of Bounako, on 8 December. Numbers are declining in many areas White-browed Forest FlycatcherFraseria & (Cheke Walsh 1996). cinerascens One perchinglowin tangledvegetation in apatch Cassin’s HawkEagle Spizaetusafricanus of closed-canopy forest along the Sekou River One seen perched and in flight over the heavily (Fig. 5), near Souroukou (08°45’N 00°40’E), on m vooded slopes above Boulohou on 12 November; 11 November. Observedfrom 6 forc.8 minutes mother over forest along a marshyareasouth-east and identified byits slate-greyupperparts and tail, afTchitchako on 23 November. Identified by its whitish underparts with a pale grey wash and alackish upperparts and white underparts with blackish scalloping on the breast, blackish fore- alack markings on its breast-sides, axillaries and crown, lores and ear-coverts, and a clean white thighs. A rare resident in Togo (Cheke & Walsh supraloral spot extending above the eye. Not pre- & 1996). viously recorded in Togo. Dowsett-Lemaire Dowsett (2007) list this species as a rare resident Black-shouldered Nightjar Caprimulgus in nearby Kyabobo National Park in eastern nigriscapularis Ghana, part ofwhich shares a border with Fazao- One callinginwoodland alongthe Kamassi River, Malfakassa. ~astofBoulohou, ontheeveningof29November. \.t least two heard at forest edge near Mt. Kpeya during the nights of10-12 December. A rare res- BirdsofFazao-MalfakassaNationalPark, Togo:Radley&Campbell BullABCVol15No2(2008)-205 Gambaga FlycatcherMuscicapagambagae on the grey-brown head and upperparts, and One hawking insects in a small cultivated area whitish underparts with very faintly streaked within open woodland habitat a few km from upper breast (Spotted Flycatcher M. striata has a Fazao, on 1 November. Identification was based more heavily streaked breast). An uncommon res- & 206-BullABCVol15No2(2008) BirdsofFazao-MalfakassaNationalPark, Togo:Radley Campbell ident of Northern Guinea Savanna (Cheke & (1996).Alsoknownfrom KyoboboNational Park, Walsh 1996), our observation requires eastern Ghana (Dowsett-Lemaire & Dowsett confirmation. 2007). Lead-coloured FlycatcherMyioparusplumbeus Eurasian Golden Oriole Oriolus oriolus One at the edge ofriparian forest cA km north of One in woodland broken by cultivation a few km Fazao on 31 October. Cheke &Walsh (1996) list west of Fazao on 6 November. The black wings this species as a scarce resident in Togo. However, were clearlyobserved. Onlythree previous records & Dowsett-Lemaire Dowsett (2007) report it to ofthis Palearctic migrant to Togo are mentioned bethecommonestflycatcherin KyoboboNational by Cheke &Walsh (1996). Park, eastern Ghana. Exclamatory ParadiseWhydah Vidua interjecta BrubruNilausafer Single males in breedingplumageseenatFazao on Two singing in savanna several km south of 29 October, atTchitchako on 27 November, and Bounako on 6 December and one foraging in at Bounako on 3 December. Identified by the rel- wooded savanna south of Mt. Kpeya on 13 atively dark maroon nape and the broad, rather December. Listed as a very scarce resident with blunt-ended tail streamers (as broad as, or broader only three recent records by Cheke & Walsh than, the bird’s bodyin flight). Red-winged Pytilia Pytiliaphoenicoptera the species’ host, was found , Captions to photos on oppositepage in the park during our survey. Status in Togo uncertain but probably an uncommon resident Figure 1. Savannawoodlandanddryforest, Fazao- (Cheke &Walsh 1996).Togo ParadiseWhydah V MalfakassaNational Park, November2005 (P. M. togoensismightalso occurin theparkas thespecies Radley) it parasitises, Yellow-winged Pytilia Pytilia Savanearboreeetforetseche, Parc National deFazao- Malfakassa, novembre2005 (P. M. Radley) hypogrammica, is present. Figure 2. Dryforestnorth ofTchitchako, Fazao- Acknowledgements MRaadllfeayk)assaNational Park, November2005 (P. M. Our research was funded through the Critical Ecosystem Partnership Fund (CEPF) and the Foretsecheau norddeTchitchako, ParcNational de Primate Action Fund (PAF) of Conservation Fazao-Malfakassa, novembre2005 (P. M. Radley) International, Washington DC, USA, and Figure3. Riparian habitatalongthe KpazaRiver, Fazao- Conservation des Especes et Populations Animales MalfakassaNational Park, 23 November2005 (P. M. (CEPA), France. Many thanks to the Franz Weber Radley) Foundation, especially to Frederic Marchand (local Vegetation riverainelelongdelaKpaza, ParcNational de director ofFazao-Malfakassa National Park) and to Fazao-Malfakassa, 23 novembre2005 (P. M. Radley) the ‘Direction de la faune et de la chasse’ for granti- Figure4. Cultivatedwoodland nearFazao, Fazao- ng permits to work in the park and proffering MalfakassaNational Park, 1 November2005 (P. M. logistical support. Robert Cheke, Ron Demey, and Radley) Bob Dowsett and Franqoise Dowsett-Lemaire gave Zonearboreeetcultiveepres deFazao, ParcNational de valuable input and advice pertaining to our findings Fazao-Malfakassa, 1 novembre2005 (P. M. Radley) and results. Figure 5. HabitatinwhichWhite-browedForest FlycatcherFraseriacinerascenswasfound, Fazao- References wMaatlefrafkaallssca.2NaktmiosnoaulthP-aeraks:t(oAf)BRoiuplaroihaonuftorheasttfaeendds into Borrow, N. & Demey, R. 2001.A Guideto theBirdsof theSekouRiver; (B) RiparianforestalongtheSekou WesternAfrica. Princeton, NJ: Princeton University River, 18 November2005 (P. M. Radley) Press. Habitatdanslequelle Gobemoucheasourcils blancs Borrow, N.& Demey, R. 2004. Birds ofWesternAfrica. Fraseriacinerascensaete trouve, ParcNationaldeFazao- Princeton, NJ: Princeton UniversityPress. Malfakassa: (A) Foretriveraineetchutes environ2 km Chappuis, C. 2000. African Bird Sounds. Vol. 2: West sud-estde Boulohoudont1’eausejettedans lariviere andCentralAfrica. 11 CDs. Paris: Societed’Etudes Sekou ; (B) ForetriverainelelongdelaSekou, 18 Ornithologiques de France &London, UK: British novembre2005 (P. M. Radley) Library. & BirdsofFazao-MalfakassaNationalPark, Togo:Radley Campbell BullABCVol15No2(2008)-207 Cheke, R.A. &Walsh,J. F. 1996. TheBirdsofTogo:An Ministere de l’Environnement et de la Protection AnnotatedCheck-list. BOUChecklistNo. 14.Tring: ForestiereduTogo (MEPFT) 2001.Rapportd’activ- British Ornithologists’ Union. ites sur les oiseaux migrateurs du palearctique Cheke, R. A. 2001. Togo. In Fishpool, L. D. C. & occidental (OMPO) et les zones humides du Togo. Evans, M. I. (eds.) Important BirdAreas in Africa Lome : Direction de la Faune et de la Chasse, andAssociatedIslands:PrioritySitesforConservation. RepubliqueTogolaise. Newbury: Pisces Publications & Cambridge, UK: aCNMIDivision ofFish & Wildlife PO Box 10007, BirdLife International. MP , Dowsett-Lemaire, F. & Dowsett, R. J. 2007. The avi- pSraaitpiannc,[email protected], USA. E-mail: fauna of the proposed Kyabobo National Park in eastern Ghana. Malimhus29: 61-88. bMax-Planck Institute for Evolutionary Anthropology, Fry, C. H., Keith, S. & Urban, E. K. (eds.) 2000. The Deutscher Platz 6, 04103 Leipzig, Germany. E-mail: Birds ofAfrica. Vol. 6. London, UK: Academic [email protected] Press. Received 6 June 2006; revision accepted 27 January Jenik, J. 1984. Coastal upwelling and distributional 2008. pattern of West African vegetation. Preslia 56: 193-204. Appendix1.BirdspeciesrecordedinFazao-MalfakassaNationalPark,27October-13December2006. Annexe1.Especesd’oiseauxobserveesdansleParcnationaldeFazao-Malfakassa,27octobre-13decembre2006. Family/Species Biome Abundance Migrant/Date Ardeidae Tigriornisleucolopha White-crestedTigerHeron Onoreahuppeblanche GCF R Bubulcusibis CattleEgret Herongarde-boeufs F AM,27Oct Butoridesstriata Green-backedHeron Heronstrie F Egrettagarzetta LittleEgret Aigrettegarzette U Ardeapurpurea PurpleHeron Heronpourpre R Ardeacinerea GreyHeron Heroncendre F Scopidae Scopusumbretta Hamerkop Ombretteafricaine C Ciconiidae Ciconianigra BlackStork Cigognenoire R PM,11 Dec Ciconiaciconia WhiteStork Cigogneblanche R PM,5Nov Threskiornithidae Plegadisfalcinellus GlossyIbis Ibisfalcinelle R PM,3Dec Accipitridae Milvusmigrans BlackKite Milannoir C AM,27Oct Gypohieraxangolensis Palm-nutVulture Palmisteafricain F Necrosyrtesmonachus HoodedVulture Vautourcharognard U Gypsafricanus AfricanWhite-backedVulture Vautourafricain R Terathopiusecaudatus Bateleur Bateleurdessavanes R Polyboroidestypus AfricanHarrierHawk Gymnogened'Afrique F Micronisusgabar GabarGoshawk Autourgabar R Accipitertachiro AfricanGoshawk Autourtachiro R Accipiterbadius Shikra Eperviershikra F Accipitererythropus Red-thighedSparrowhawk EpervierdeHartlaub GCF R Accipitermelanoleucus BlackSparrowhawk Autournoir R Kaupifalcomonogrammicus LizardBuzzard Autourunibande F Buteoauguralis Red-neckedBuzzard Bused'Afrique F Spizaetusafricanus Cassin’sHawkEagle AigledeCassin GCF U Polemaetusbellicosus MartialEagle Aiglemartial U Falconidae Falcoalopex FoxKestrel Crecerellerenard SGS U Falcochicquera Red-neckedFalcon Fauconchicquera R Falcobiarmicus LannerFalcon Fauconlanier U Falcoperegrinus PeregrineFalcon Fauconpelerin R Phasianidae Ptilopachuspetrosus StonePartridge Poulederoche C & 208-BullABCVol15No2(2008) BirdsofFazao-MalfakassaNationalPark, Togo:Radley Campbell Francolinusahantensis AhantaFrancolin Francolind’Ahanta GCF F Francolinusbicalcaratus Double-spurredFrancolin Francolinadoubleeperon F Numididae Numidameleagris HelmetedGuineafowl PintadedeNumidie U Rallidae Amauromisflavirostra BlackCrake Raleabeejaune R Burhinidae Burhinussenegalensis SenegalThick-knee OedicnemeduSenegal R Charadriidae Vanellusspinosus Spur-wingedLapwing Vanneauaeperons R Scolopacidae Tringaochropus GreenSandpiper Chevalierculblanc C PM,23Nov Actitishypoleucos CommonSandpiper Chevalierguignette C PM,23Nov Columbidae Treroncalm AfricanGreenPigeon Colombarafrontnu C Turturtympanistria TambourineDove Tourtelettetambourette R Turturafer Blue-spottedWoodDove Tourteletteamethystine F Turturabyssinicus Black-billedWoodDove Tourteletted’Abyssinie R Streptopeliasemitorquata Red-eyedDove Tourterelleacollier F Streptopeliavinacea VinaceousDove Tourterellevineuse C Psittacidae Poicephalussenegalus SenegalParrot Perroquetyouyou SGS F Psittaculakrameri Rose-ringedParakeet Perrucheacollier U Musophagidae Tauracopersa GreenTuraco Touracovert GCF C Musophagaviolacea VioletTuraco Touracoviolet SGS U Criniferpiscator WesternGreyPlantain-eater Touracogris F Cuculidae Chrysococcyxklaas Klaas’sCuckoo CoucoudeKlaas U AM,30Nov Ceuthmocharesaereus Yellowbill Malcohaabeejaune R Centropussenegalensis SenegalCoucal CoucalduSenegal C Tytonidae Tytoalba BamOwl Effraiedesclochers U Strigidae Otussenegalensis AfricanScopsOwl Petit-dueafricain F Bubocinerascens GreyishEagleOwl Grand-dueduSahel F Glaucidiumperlatum Pearl-spottedOwlet Chevechetteperlee R Strixwoodfordii AfricanWoodOwl Chouetteafricaine F Caprimulgidae Caprimulgusclimacurus Long-tailedNightjar Engouleventalonguequeue U Caprimulgusnigriscapularis Black-shoulderedNightjar Engouleventaepaulettenoires GCF R Apodidae Telacanthuraussheri MottledSpinetail Martinetd’Ussher F Cypsiurusparvus AfricanPalmSwift Martinetdespalmes F Apusaffinis LittleSwift Martinetdesmaisons F Tachymarptismelba AlpineSwift Martinetaventreblanc R AM,12Dec Trogonidae Apalodermanarina Narina'sTrogon Trogonnarina R Alcedinidae Halcyonleucocephala Grey-headedKingfisher Martin-chasseuratetegrise F AM,31Oct Halcyonmalimbica Blue-breastedKingfisher Martin-chasseurapoitrinebleue F Halcyonchelicuti StripedKingfisher Martin-chasseurstrie F Alcedocristata MalachiteKingfisher Martin-pecheurhuppe U Alcedoquadribrachys Shining-blueKingfisher Martin-pecheurazure U Megacerylemaxima GiantKingfisher Martin-pecheurgeant R Meropidae Meropspusillus LittleBee-eater Guepiernain R Meropsbulocki Red-throatedBee-eater Guepieragorgerouge SGS F Meropsalbicollis White-throatedBee-eater Guepieragorgeblanche R AM,28Nov Meropsapiaster EuropeanBee-eater Guepierd’Europe R PM,22Nov Meropsnubicus NorthernCarmineBee-eater Guepierecarlate R AM,13Nov BirdsofFazao-MalfakassaNationalPark, Togo:Radley&Campbell BullABCVol15No2(2008)-209 Coraciasnaevius Rufous-crownedRoller Rolliervarie U AM,8Dec Coraciascyanogaster Blue-belliedRoller Rollieraventrebleu SGS R Eurystomusglaucurus Broad-billedRoller Rolleviolet U AM,6Nov Phoeniculidae Phoeniculuspurpureus GreenWood-hoopoe Irrisormoqueur R Rhinopomastusaterrimus BlackWood-hoopoe Irrisornoir R Bucerotidae Bucorvusabyssinicus AbyssinianGroundHornbill Bucorved’Abyssinie R Tockusfasciatus AfricanPiedHornbill Calaolongibande GCF C Tockusnasutus AfricanGreyHornbill Calaoabeenoir C Bycanistesfistulator PipingHornbill Calaosiffleur GCF u Capitonidae Pogoniulusbilineatus Yellow-rumpedTinkerbird Barbionacroupionjaune c Pogoniuluschrysoconus Yellow-frontedTinkerbird Barbionafrontjaune c Lybiusvieilloti Vieillot'sBarbet BarbicandeVieillot u Lybiusbidentatus Double-toothedBarbet Barbicanbidente u Lybiusdubius BeardedBarbet Barbicanapoitrinerouge SGS u Indicatoridae Indicatorindicator GreaterHoneyguide GrandIndicateur R Indicatorminor LesserHoneyguide PetitIndicateur R Picidae Campetherapunctuligera Fine-spottedWoodpecker Picatachesnoires R Campetheracailliautii Green-backedWoodpecker PicdeCailliaut U Dendropicosfuscescens CardinalWoodpecker Piccardinal R Dendropicosgoertae GreyWoodpecker Picgoertan U Dendropicosobsoletus Brown-backedWoodpecker Picadosbrun R Alaudidae Pinarocoryserythropygia Rufous-rumpedLark Alouetteaqueuerousse R AM,13Dec Hirundinidae Psalidoprocneobscura FantiSaw-wing Hirondellefanti GCF U AM,31Oct Hirundoabyssinica LesserStripedSwallow Hirondelleagorgestriee R AM,5Dec Hirundodaurica Red-rumpedSwallow Hirondellerousseline R AM,26Nov Hirundofuligula RockMartin Hirondelleisabelline U Hirundosmithii Wire-tailedSwallow Hirondellealongsbrins U Hirundolucida Red-chestedSwallow HirondelledeGuinee R Hirundorustica BarnSwallow Hirondellerustique U PM,28Oct Delichonurbicum CommonHouseMartin Hirondelledefenetre U PM,4Nov Motacillidae Motacillaflava YellowWagtail Bergeronnetteprintaniere U PM,23Nov Anthussp. Pipitsp. Pipitsp. R Campephagidae Coracinapectoralis White-breastedCuckooshrike Echenilleuraventreblanc R Pycnonotidae Andropaduswens LittleGreenbul Bulbulverdatre R Baeopogonindicator HoneyguideGreenbul Bulbulaqueueblanche GCF U Pyrrhurusscandens Leaflove Bulbulaqueuerousse GCF C Pycnonotusbarbatus CommonBulbul Bulbuldesjardins C Nicatorchloris WesternNicator Bulbulnicator GCF R Turdidae Cossyphaniveicapilla Snowy-crownedRobinChat Cossypheacalotteneigeuse R Cercomelafamiliaris FamiliarChat Traquetfamilier U Myrmecocichlaalbifrons White-frontedBlackChat Traquetafrontblanc SGS R Turduspelios AfricanThrush Merleafricain F Sylviidae Melocichlamentalis AfricanMoustachedWarbler Melocichleamoustaches R Acrocephalusscirpaceus EurasianReedWarbler Rousserolleeffarvatte U PM,28Nov Acrocephalusarundinaceus GreatReedWarbler Rousserolleturdo'ide U PM,29Nov Hippolaisopaca WesternOlivaceousWarbler Hypolaispale R PM,16Nov Hippolaispolyglotta MelodiousWarbler Hypolai'spolyglotte F PM,15Nov Cisticolacantans SingingCisticola Cisticolechanteuse U Cisticolalateralis WhistlingCisticola Cisticolesiffleuse U Cisticolaaberrans Rock-lovingCisticola Cisticoleparesseuse R 210-BullABCVol15No2(2008) BirdsofFazao-MalfakassaNationalPark, Togo:Radley&Campbell Cisticolabrachypterus Short-wingedCisticola Cisticoleaailescourtes R Priniasubflava Tawny-flankedPrinia Priniamodeste C Camaropterabrachyura Grey-backedCamaroptera Camaroptereatetegrise C Eremomelapusilla SenegalEremomela Eremomeleadosvert SGS C Sylviettabrachyura NorthernCrombec Crombecsitelle R Sylviettavirens GreenCrombec Crombecvert GCF U Phylloscopustrochilus WillowWarbler Pouillotfitis R PM,6Dec Hypergerusatriceps OrioleWarbler Noircaploriot SGS R Muscicapidae Fraseriacinerascens White-browedForestFlycatcher Gobemoucheasourcilsblancs GCF R Melaenomisedolioides NorthernBlackFlycatcher Gobemouchedrongo U 1Muscicapagambagae GambagaFlycatcher GobemouchedeGambaga SGS R Myioparusplumbeus Lead-colouredFlycatcher Gobemouchemesange R Ficedulahypoleuca EuropeanPiedFlycatcher Gobemouchenoir C PM,11Nov Monarchidae Elminialongicauda AfricanBlueFlycatcher Tchitrecbleu U Terpsiphoneviridis AfricanParadiseFlycatcher Tchitrecd’Afrique F Platysteiridae Megabyasflammulatus ShrikeFlycatcher Biasecorcheur GCF U Platysteiracyanea CommonWattle-eye Priritacollier U Batissenegalensis SenegalBatis PriritduSenegal u Timaliidae llladopsisfulvescens Brownllladopsis Akalatbrun GCF R Turdoidesplebejus BrownBabbler Crateropebrun U Turdoidesreinwardtii BlackcapBabbler Crateropeatetenoire SGS R Phyllanthusatripennis CapuchinBabbler Phyllanthecapucin GCF R Paridae Parusguineensis White-shoulderedBlackTit Mesangeaepaulettes U Nectariniidae Cyanomitraverticalis Green-headedSunbird Souimangaateteverte u Cyanomitraolivacea OliveSunbird Souimangaolivatre R Chalcomitraadelberti Buff-throatedSunbird Souimangaagorgerousse GCF R Chalcomitrasenegalensis Scarlet-chestedSunbird Souimangaapoitrinerouge U Hedydipnacollaris CollaredSunbird Souimangaacollier C Hedydipnaplatura PygmySunbird Souimangapygmee u AM,30Nov Cinnyrisvenustus VariableSunbird Souimangaaventrejaune c Cinnyrissuperbus SuperbSunbird Souimangasuperbe GCF u Cinnyriscoccinigastrus SplendidSunbird Souimangaeclatant SGS R Cinnyriscupreus CopperSunbird Souimangacuivre R Zosteropidae Zosteropssenegalensis YellowWhite-eye Zosteropsjaune R Laniidae Corvinellacorvina Yellow-billedShrike Corvinelleabeejaune SGS R Malaconotidae Malaconotussulfureopectus Sulphur-breastedBushshrike Gladiateursoufre R Antichromusminutus MarshTchagra Tchagradesmarais F Tchagrasenegalus Black-crownedTchagra Tchagraatetenoire F Dryoscopusgambensis NorthernPuffback CubladeGambie F Laniariusaethiopicus TropicalBoubou Gonolekd’Abyssinie F Nilausafer Brubru Brubruafricain U Prionopidae Prionopsplumatus WhiteHelmetshrike Bagadaiscasque F Oriolidae Oriolusnigripennis Black-wingedOriole Loriotaailesnoires GCF U Oriolusauratus AfricanGoldenOriole Loriotdore F AM,31Oct Oriolusoriolus EurasianGoldenOriole Loriotd’Europe R PM,5Nov Dicruridae Dicrurusludwigii Square-tailedDrongo DrongodeLudwig F Dicrurusadsimilis Fork-tailedDrongo Drongobrillant F Corvidae Ptilostomusafer Piapiac Piapiacafricain SGS R Sturnidae Lamprotornispurpureus PurpleGlossyStarling Choucadorpourpre SGS R BirdsofFazao-MalfakassaNationalPark, Togo:Radley&Campbell BullABCVol15No2(2008)-211 Lamprotornischalcurus Bronze-tailedGlossyStarling Choucadoraqueueviolette SGS R Lamprotornischloropterus LesserBlue-earedStarling ChoucadordeSwainson F Lamprotornissplendidus SplendidGlossyStarling Choucadorsplendide R Passeridae Petroniadentata BushPetronia PetitMoineau SGS F AM,3Nov Ploceidae Ploceusheuglini Heuglin’sMaskedWeaver Tisserinmasque SGS U Ploceuscucullatus VillageWeaver Tisseringendarme C Ploceusnigricollis Black-neckedWeaver Tisserinacounoir R Anaplectesrubriceps Red-headedWeaver Tisserinecarlate R Queleaerythrops Red-headedQuelea Travailleurateterouge U AM,4Nov Euplecteshordeaceus Black-wingedBishop Euplectemonseigneur U Euplectesfranciscanus NorthernRedBishop Euplectefranciscain U Euplectesmacroura Yellow-mantledWidowbird Euplecteadosd’or R Estrildidae Nesochariscapistrata Grey-headedOliveback Dos-vertajouesblanches SGS R Pytiliahypogrammica Yellow-wingedPytilia Beaumarquetaailesjaunes SGS R Pytiliaphoenicoptera Red-wingedPytilia Beaumarquetaurore SGS R Lagonostictasenegala Red-billedFirefinch AmaranteduSenegal F Lagonostictarara Black-belliedFirefinch Amaranteaventrenoir SGS F Lagonostictalarvata Black-facedFirefinch Amarantemasque SGS R Estrildamelpoda Orange-cheekedWaxbill Astrildajouesoranges F Spermestescucullatus BronzeMannikin Capucinnonnette C Viduidae Viduamacroura Pin-tailedWhydah Veuvedominicaine u Viduainterjecta ExclamatoryParadiseWhydah Veuved’Uelle SGS u Fringillidae Serinusmozambicus Yellow-frontedCanary SerinduMozambique F Emberizidae Emberizatahapisi Cinnamon-breastedRockBunting Bruantcannelle F AM,29Oct Emberizacabanisi Cabanis’sBunting BruantdeCabanis U 1=Requiresconfirmation Biome Abundance SGS=restrictedtotheSudan-GuineaSavannabiome(24species) C=Common:observeddaily,eithersinglyorinnumbers GCF=restrictedtotheGuinea-CongoForestsbiome(20species) F=Fairlycommon:observedonmostdays Migrant/Date U=Uncommon:irregularlyobservedandnotonthemajorityofdays PM=PalearcticMigrant R=Rare:rarelyobserved,oneortworecordsofsingleindividuals. AM=Intra-AfricanMigrant Appendix2.AdditionalbirdspeciespreviouslyreportedfromFazao-MalfakassaNationalPark. Annexe2.Especesd’oiseauxrapporteesauparavantpourleParcnationaldeFazao-Malfakassapard’autreschercheurs. Family/Spec/es Biome Reference Ardeidae Nycticoraxnycticorax Black-crownedNightHeron Bihoreaugris MEPFT2001 Ciconiidae Ciconiaepiscopus Woolly-neckedStork Cigogneepiscopate MEPFT2001 Threskiornithidae Bostrychiahagedash HadadaIbis Ibishagedash Cheke&Walsh1996 Anatidae Dendrocygnabicolor FulvousWhistlingDuck Dendrocygnefauve MEPFT2001 Dendrocygnaviduata White-facedWhistlingDuck Dendrocygneveuf MEPFT2001 Pandionidae Pandionhaliaetus Osprey Balbusardpecheur MEPFT2001 Accipitridae Neophronpercnopterus EgyptianVulture Percnoptered’Egypte Cheke&Walsh1996 Pernisapivorus EuropeanHoneyBuzzard Bondreeapivore R.Chekeinlitt.2006 Aquilarapax TawnyEagle Aigleravisseur Cheke&Walsh1996 Hieraaetuspennatus BootedEagle Aiglebotte R.Chekeinlitt.2006 Hieraaetusayresii Ayres’sHawkEagle Aigled’Ayres Cheke&Walsh1996 Stephanoaetuscoronatus CrownedEagle Aiglecouronne R.Chekeinlitt.2006 & 212-BullABCVol15No2(2008) BirdsofFazao-MalfakassaNationalPark, Togo:Radley Campbell

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