Available generic names for trilobites

221 Pages·2003·81.8 MB·English

by  P A Bell

AVAILABLE GENERIC NAMES FOR TRILOBITES
P.A. JELLANDJ.M. ADRAIN
Jell,RA.&Adrain,J.M.06202003:Availablegenericnamesfortrilobites. Memoirsofthe
QueenslandMuseum Brisbane. ISSN0079-8835.
A consolidated list of available generic names introduced since the beginning of the
binomial nomenclature system fortrilobites is presented forthefirst time. Each entry is
accompaniedbythe authoranddateofavailability,by thenameofthetype species,by a
lithostratigraphic orbiostratigraphic andgeographic reference forthe type species, by a
family assignment and by an age indication ofthe type species at the Period level (e.g.
MCAM, LDEV). A second listing ofthesenames istaxonomically arranged in families
withthefamilieslistedalphabetically,higherlevelclassificationbeingoutsidethescopeof
this work. Wealsoprovidea listofnamesthathaveapparently been applied to trilobites
butwhichremainnominantidawithinthe ICZNdefinition. Trilobites, genericnames,
checklist.
Peter A. Jell, Queensland Museum, PO Box 3300. South Brisbane, Queensland 4101,
Australia; Jonathan M. Adrain, Department ofGeoscience, 121 Trowbridge Hall, Univ-
ersityofIowa, Iowa Cit\\ Iowa 52242, USA; I August 2002.
Trilobite fossils attracted the attention of could find. This listwascopiedonan earlyspirit
humans in different parts ofthe world from the stencil machine to some 20 or more trilobite
verybeginning,probablyevenprehistoric times. workers aroundtheworld,principallythosewho
In the 1700s various Europeannatural historians would author the 1959 Treatise edition. Weller
began systematic study of living and fossil alsodrewonthis compilation forhisPresidential
organisms including trilobites. The name address to the Paleontological Society in 1951
Trilobita was introduced by Walch in 1771, at (Weller, 1952). However, his list was never
aboutthe sametimethatCarl Linnaeuswasdev- published and to ourknowledge no accompany-
eloping his system of binomial nomenclature. ingbibliographywasevercompiled. Inthe 1959
The early 800s witnessed a number ofmono- Treatisethebibliography isvery selectand large
1
graphic works on Palaeozoic fossils including numbers oftheauthorcitationsofgeneric names
numerous trilobites. Major works of Dalman are not included.
(H1a8w2l7)e,&BroCnogrndiaar(t18(4178)2,2)A,nWgaelhilnen(b1e8r5g1,(11882514)), underBwyatyhfeor19a8r0esv,iswihoennofeatrhleytprrileopbairtaetvioolnsumweeroef
and Banande (1846-1872) accelerated the syst- the Treatise a large increase in the number of
ematic study oftrilobites so that by 1893 when generic names to be dealtwith had occurred. To
Vogdesproducedthefirstannotatedbibliography assess the extent of the increase Harry Whit-
it was printed on over 400 pages. With sub- tingtonandSimonKellycompiledalistofgenera
sequentadditions(Vogdes, 1917, 1925)hiswork erected after publication of the first Treatise
provides the most comprehensive, ifnot always editionupto 1983. Thislistcontainedabout2500
themostaccurate, referencework forentranceto nameshavingbeencompiledfromtheZoological
trilobite studies ofthe 19th century. Record and Irom lists provided by various
In the 20^*^ century the volume ofpublic- individuals. Thislistwasdistributedtoprospect-
ations was such that the only comprehensive ive authors ofthe revisedtrilobitevolume ofthe
Treatise butnot otherwise published.
surveyoftrilobiteswasatthegenericlevel,inthe
Treatise on Invertebrate Paleontology (Moore, The 1959Treatisedealt with approximately
1959; Kaesler, 1997). Earlyinthepreparation of 1200trilobitegenericnames andthefirstrevised
the first edition of the trilobite volume of the volume of the trilobite (Part O) Treatise (1997)
Treatise J. Mar\'in Weller of the University of dealt with about 500 generic names although
Chicago compiled a list of trilobite generic Weller( 1952) had indicated 1436wereknown in
names that would need to be dealt with in the 1950. Thus the list presented below, with more
Treatise. Weller’s checklist included all names than 5000 generic names, represents a 430%
and variations including mispellings that he increase in availablegeneric names in the last 50
332 MEMOIRS OF THE QUEENSLAND MUSEUM
years. Weller's (1952, fig. 11) projection ofthe Most of these remain valid names for trilobite
rate of trilobite generic increase has not been genera but many have become invalid asjunior
realised; hecorrectlyforesawthatitwouldnotbe synonyms, junior homonyms, unjustified
possible. However, the aggregate number of emendations, unnecessary substitute names or
5000+farexcedeshiscalculatedmaximumcur\^e suppressed names but they all remain available
whichhadaprojection offewerthan3000 in the irrespective ofanysuch subsequent history.
year2000. Given Weller'sexperiencewearenot
so bold as la anticipate the future. However, the Alphabetical listing.
1990s saw a marked reduction in the numberof Each availablegenericname is followedby
new names introduced probably due to the the author (all upper case) and year of publ-
diminishingnumberofactivetaxonomists inthe ication. The type species (as originally cited)
fieldandpossiblyduealsotodecreasing areasof follows in square brackets. Next are data on the
Palaeozoicoutcrop remainingtobeinvestigated. typelocalityofthe species. Wherepossible
The cuiTent paper arose from independent ef- therockfonnationcontainingthetypelocalityis
forts by each of us. JMA had compiled Ord- quotedwiththeStateorProvinceand/orcountry.
ovicianand Siluriangenera insupportofstudies Ifthe formation is not known we try to provide
by Adrain et al. (1988) and Adrain & Westrop thebiozoneofthetypehorizonorfailingthatthe
(2000) and this list had been freely available best geographic data available. In dealing with
since 1998. PAJ assembled Cambrian generic largercountrieswequotethestateorprovincebut
names to define the extent of the Plychopar- inmanycasesonlythe countryisquoted. This is
ioideaandextendedthistopresentapaperonthe followed, after a semicolon, by the family
rateofappearanceofgenericnames. Thatpaper assignment(all in uppercase)whichcrossrefers
was delivered at the Third International Confer- tothe latersystematiclisting. Followingthe last
ence on trilobites and their relatives in Oxford, semicolon is an indication ofthe age ofthe type
April, 2001,where we agreed to join forces to horizonofthetypespecies. Thisis intheformof
produceacomprehensive list. Broadly speaking abbreviations ofthe Palaeozoicperiods namely
PAJisresponsiblefortheCambriancompilation. Cambrian (CAM), Ordovician (ORD), Silurian
JMA for the Ordovician and Silurian and JMA (SIL),Devonian(DEV)andPennian(PERM)or
and PAJ forthe Devonian to Pennian. subperiodsoftheCarboniferousi.e. the Mississ-
We hope that our effort herein will be a pirpepfiiaxned(bMyILS(Sl)oweorr),PMen(nmsidydllvea)noiraUn((upPpEerN)No)r
significant aidtoallsUidentsoftrilobitesintheir
considerations of whether or not to erect new combinations ofthese where appropriate.
genericnames. Alltoo frequentlywe seenames Atthe endofmanyentriesis ananotationin
passing to synonymy because the full literature square brackets indicating known synonymies.
search had not been undertaken or because the Thefirstthreelettersareanabbreviationofjunior
full extentofthe family concerned had notbeen (j.)orsenior(s.); objective(o.)orsubjective(s.);
scanned. We hope this publication will be an synonyms (s.) followed by the relative generic
invitationtobemorecomprehensiveinconsider- nameandthen theauthoritywerely on forthese
ation ofgeneric assignments and in discussions relationships. Although we have attempted to
ofaffinities. Moreover, we anticipate its use in quotetheearliestauthortorecognisesynonymies
other ways not the least of which may be itispossiblethatwemayinsomecasesun^ow-
quantitativebiodiversitystudies. ingly be quoting a secondary source. Some
METHOD OF ASSEMBLY synonymies proposed in the literature are not
quotedhereinbecausewedo notacceptthem. In
Wepresentbothobjectiveandsubjectivedata. other cases, where synonymies have been
The bibliography is objective whereas the syst- proposed and subsequentlydenied oraltered we
ematicarrangementisalmostentirelysubjective. have attempted to present our view ofthe most
In the alphabetical listing data are objective ex- probable currentsituation.
cept for family assignment and age. We have Inthecaseofnamesthatarejuniorobjective
tried, as far as possible, to consult the original synonyms (j.o.s.) orwerepreoccupied and have
literauire when compiling objective data. We beenreplacedthis is indicatedinsquarebrackets
have listed available names associated with at that name with relevant data supplied at the
trilobites, i.e. those names that entered the senior objective or replacement name. While
literature meeting the necessary' criteria of the thereareverymanyexamplesofmisspellingsof
International Code ofZoologicalNomenclature. generic names throughoutthehistoryoftrilobite
AVAILABLE GENERIC NAMES FORTRILOBITES 333
publications most of these are obvious typo- not have direct experience ourselves. We
graphical eiTors and those variant spellings are acknowledge that disagreements remain among
notincludedinourlists. However, insomecases trilobite workers in some groups and that our
we have identified thatthese errors have caused arrangement may not be universally accepted.
confusion or are simply not obvious in their However, by including all generic names we
meaning so we have includedthese examples in hope that future classifications will seek to deal
thelistindicatingtheerrorinsquarebrackets. more comprehensivelywith relatedgenera.
Bibliography We group the genera into families and
This list contains all references cited in the although we recognise the considerable effort
alphabetical list so that it includes papers in some workers have put into defining sub- and
whichnewgenericnameswereerected,inwhich superfamilies (particularly e.g. in the Proetidae,
specieswereerectedthatweresubsequentlydes- Redlichiidae, Ellipsocephalidae, or Asaphidae)
ignated as generic types and in which accepted we observe that general agreement has not been
synonymieswereproposed.Pleasenotethisdoes reached and in most cases such classifications
not constitute a complete list ofpapers on the have not been comprehensive in placing all
taxonomyoftrilobitesalthoughitcontainsavery relevant genera. We therefore, prefer to suggest
high percentage of such papers. Whereas broader groupings that can be assessed for
common bibliographic practice is to provide a intrafamilial classifications by more detailed
transliterationandatranslationoftitlesofpapers investigations.
in the Russian language we have provided only
the translation in the interests of economy of Nominanuda
space. We provide a list of generic names,
We have indicated papers published in apparently referring to trilobites, that remain, as
languages using otherthan the arable characters far as we are aware, nomina nuda as defined in
byenclosingtheirtitlesinsquarebrackets. While the International Code of Zoological Nomen-
transliteration of the names of Russian and clature. Thelistisprovidedasareferenceshould
Chineseauthors into Englishhasvariedwiththe these names appear in any future literature by
differenttranslationstandardsemployedw^ehave wayofvalidationor furtheruseasnomina nuda.
endeavoured to maintain spellings that are Thislistisnotasexhaustiveasthelistofavailable
recognisable and consistent thus giving the best names and we have not attempted to include all
chanceofrecognitionovertime. Forexamplewe literaturerelevantto these nomina nuda.
quote CHERNYSHEVA not TCHERNY-
SHEVA, EGOROVAnotYEGOROVA, XIANG ACKNOWLEDGEMENTS
not HSIANG ZHANG not CHANG, ZHU not
CHU etc. We have indicated most ofthese var-
iantsbyacrossreferenceinthelistofliterature. PAJisgratefultoJ.MarvinWeller,HarryWhit-
tington and Simon Kelly for the unpublished
Although there are a number ofinstances lists, mentioned above, which, along with the
whereauthorswiththesamefamilynameappear 1959 Treatise and the Zoological Record, were
RinICtHheTEbRib,lYioUgrAaNphyand(eZ.gH.OAUL)BEthReTirIa,ssoHcAiaHtNio,n tPhAeJintihtiaanlksbasiPsetfeorPtahlemeCramwb^rhioanencgoamgpeidlatiinona.n
with the year of publication removes any extensive mutually beneficial crosscheck ofthe
ncoanmfleisZioHnA. NHGowceovnefrus,ioinntishepocsassiebleofbethtewefeanmisliyx CUSa)m.brWieanarleisgtriantgefauglaitnosZthhainsgdaWteanbtaasneg,(STYatNyPaLn-a
separate authors and it is with this name alone Pegel, Carsten Brauckmann, Greg Edgecombe,
that an initial is included throughout the Peng Shanchi, Zhou Zhiyi, David Holloway,
alphabetical list. GerdGcyer,RichardForteyandothersw'howere
Paperswithmorethat2 authorsarereferred everreadyto answerourquestions aboutpartic-
to in the alphabetical list by the first author and ularproblemsandinmanycasesprovidedunsol-
others (e.g. GORYANSKY et ak, 1964). icited improvements to our compilation. PAJ is
thankful to Kathy Buckley, Meg Lloyd and
Systematic listing Victoria Harrison of the Queensland Museum
This listreflects ourownfindings in many andJMAisthankfultoTomHegna,Universityof
areas along with our assessments of other Iowa, all ofwhom ably sought out difficult-to-
workers opinions in groups with which we may obtain materials forus.
334 MEMOIRS OF THE QUEENSLAND MUSEUM
NOTES comparative statements. They fail to satisfy
Article 13.1.1,andshouldberegardedasnomina
1. Anumberofhomonymsareidentifiedinthe nuda.
compilation ofthis list and replacement names
are introduced where necessary. Generic names 5. InaccordancewthArticle23.9ofthe ICZN,
introduced are:- Eoasaphiscellus, Paracaly- Goniopleura is designated a twmen ohliiiim and
menemene. Paralorenzangella, Pterocephal- suppressed, and Maurotarion is considered a
opsiniis and Thgonyangaspis. nomenprotectum,andavalidname. Asfaraswe
areaware. Goniopleurahas neverbeenusedas a
2. According to Vogdes (1893) Rafinesque validnamesubsequenttoitsoriginalpublication.
1832 introduced Telesiops and Prowenites and
Maurotarion is inwidespreaduse asapresumed
indicated that in 1817 he had introduced validnameforthegroup(e.g..Adrain&Chatter-
Glomerites and that in 1821 he had introduced ton, 1995; Adrain & Edgecombe, 1996; Alberti,
Alloctops, Diphpsites, Tonioligus, Isociomesa 1969; Curtis & Lane, 1997; Edgecombe &
and Retusites. To our knowledge these names
Fortey, 2000; Holloway. 1994; Sandford, 2000).
have not been subsequently used and certainly
not in any modem sense. We are unable to 6. The synonymy ofHsuaspis and Estaingia
identify any specimens that can be attached to (Jell in Bengtson et ak, 1990) has been accepted
these names much less any identifiable type by a numberofsubsequentauthorstowhomPAJ
specimens. While these apparently remain apologises for identifying the senior synonym
availablenameswehavenotbeenabletoprovide withoutrefeiringtotheoriginalpublication. PAJ
anyoftheserialiseddataweareattachingtoother recently obtained acopy ofthe Science Journal,
names in the list. Academia Sinica volume 5 (W. Zhang in W.
3. Members ofthe Nektaspida have generally Zhang et al.. 1957) in which Hsuaspis was first
been regarded as trilobites or closely related to mentioned. That publication contains a line
drawinglabelledHsuaspissinensisandthename
tlrisitlionbgi.tesHoawnedveorn,tchlaatssbiafsiicsatairoenwiintchliundetdheignrtohuips ismentioned ina faunal list foranhorizon inthe
remainshighly contentious sowe donotattempt section being described. In a second publication
anyfamilyclassillcationatthisstageeventhough inthesameyear(W. Zhang, 1957)thenewname
one could confidently say that Naraoia belongs is mentioned on the coirelation chart with a
to theNaraoiidae. We exclude MaritimeUa and footnote indicating its type species. Neither of
Orieutella of Repina & Okuneva (1969) these 1957 mentions of the name satisfies the
athcecseeptairnegptsheeudoobfsoesrsvialtsi.onofRobison( 1984)that cIoCnZsNidcerrietderiaa fnoormaveanilanbiuldituym. Tihneenaacmheomfusthtebsee
4. Clarkson & Howells (1981) and Howells pLuubleitcaalt.io(n1s9.65T)h,uasyHesauraasfptiesrbPeoccoacmkeahvaadilearbelceteidn
(1982) considered several names proposed by its synonym Estaingia, which must be con-
Lainont (1978), including the encrinurid genus- sidered theseniorname.Availabilityof
MgarcoguupingcammeelslaA,risPiaor&haecgagligaa.chCiaal.gaacnhida,TnLpepiitac.hiaa,s Wsi.miZlhaarlnygdianteWs.fZrhoamnLguetetala.l..1(915976)5,)aasncdonmomtofnrloym
invalid. Clarkson Howells (1981) argued on quoted.
tthhee btahseins-coufrrteynpteemdaitteiroinalo,fctihteingICAZrtNi.cleHoIw3eAllosf Asecondapology from PAJ is forhaving mis-
w(i19th82a)daerqguuaetdethdaitagtnhoesneas.mesInwetrheenfootnsnueprpocratseed, sEesdtaitnhgeiiinnatie'obdyucQtpioinko(f19t7h5ea)faamnidlyi-ngardovueprtnanatmley
problems with subsequent identification oftype using the junior synonym Ichangiidae for this
specimens, and the fact that the designated type group of trilobites. Estangiinae is the senior
specimens may not be well enough preserved to family-group name and is elevated to Estaing-
permit recognition ofthe taxa, do not appear to iidae herein to replace Ichangiidae.
havebearingontheavailabilityofthenames,but 7. Anchiopella was introduced by Reed
rather indicate that they should be regarded as (1907b) for a subgroup ofD. anchiops that he
nomifia dubia. For Howells’s (1982) arguments referred to as the cristagalli subgroup. He
about diagnoses, Anstobeggia. Leiichia. Para- confirmedthisashistypespeciesdesignationina
calgachia, and Trippia areeach accompaniedby later paper. Lesperance & Bourque 0971) out-
anexplicitdiagnosisinLament(1978)andhence linedindetailthedifficulthistoryoPAnchiopella
satisfy Article 13 ofthe ICZN. Calgachia and ?ir\(JAnchiopsisandconcludedthatbothareavail-
Macguigiiiella, however, lackadiagnosis orany able names. We agree with their conclusions.
AVAILABLE GENERIC NAMES FORTRILOBITES 335
Thus, in the absence ofgoodreason to suppress with Szechuanella wherein Paraacidaspis was
Anchiopella,Cooper’s(1982)actionofassigning used in association with an available species
cristagalli to Bainella means that Bainella be- name even though the quoted type species,
comes a junior subjective synonym of Anch- hunanica, remained a nomen nudum until
iopella. Poletaevamade it available in 1963. Thus when
8. Shergold et al. (1988) detailed the nomen- Shergoldetal.(2000)attributedParaacidaspisto
clatural history of Szechuanella. It was intro- Poletaeva, 1963 with the type species hummica
duced, withtype speciesszecJiuanensis^ asano- alsodating from Poletaeva. 1963 they employed
men nudum by Lu (1959). Zhang & Fan (1960) adifferentlogictothatwhichhadbeenappliedto
described a new species, rectangida^ and as- Szechuanella. Theseareexactlyanalogouscases
signed it to Szechuanella while quoting Lu’s and Paraacidaspis must be considered to date
szechuanensisas the type species even though it from Poletaeva, 1960 with P. sibirica the type
remained a nomen nudum until made available speciesbymonotypy. IfanythingParaacidaspis
tbhyatLtuheinac1t9o62f.ZhSahnegrg&olFdaenlwals.m(g19S8z8e)chcuoannseildleareidn (haIs960a)btehtatnerdcoleasimSzetochauvaaniellablialiitnyZihnanPgole&taFeavna
associationwithanavailablespeciesgroupname (1960) because Poletaeva expressly diagnosed
made reciangula the type species by monotypy. the genus whereas Zhang & Fan diagnosed the
This was accepted by Peng (1990) who cons- species. However, the lattercanbe takentobe a
PSidzaeerrcaehsduiaeSnc.ehlsulzaaencebhleulcaaanLmeIneUsa,ivsa1i9Ll7ua7,b.leT19ht6uh2rsowutoeghabZgehrleaoennggth&atto gaveanielraibcledisapgencoiseiss-gbreocuaputsaexoitnacpopnltiaeisnetdotthheereoinnl.y
Fan (1960) with its type species rectangula by 9. InaccordancewithArticle23.9oftheICZN,
monotypy. Paraszechuanella LIU, in ZHOU et Dicranactis is designated a nomenoblitum (and
al., 1977 has R latilimbata as type species and suppressed) andScahnscutellum is considered a
QOxAmmszechuanensis Lu, 1962. nomenprotectum(andvalid).Dicranactishasnot
The history of Paraacidaspis^ detailed by beenused inthemodernliterature(Snajdr, 1984)
Shergoldetal.(2000),hasthenameintroducedas and Scabriscutellum is in widespread use as a
anomennudum in 1957andconsideredanomen presumed valid name for the group (Alberti,
nudumagaininPoletaeva's I960paper(Bruton, 1971, 1981b, 1983;Archinal, 1994;Basse, 1996;
1987). However, Poletaeva made available P. Feist, 1974; Liitke, 1965; Maksimova, 1960,
sibiricain 1960andthuscreatedadirectanalogy 1979).
336 MEMOIRS OF THE QUEENSLAND MUSEUM
ALPHABETICALLIST
Aayemenaytcheia LIEBERMAN, 1994 [Dechenella Acantholenus MATTHEW, 1898 [Leptoplastus
paragi'anulata ORMISTON, 1967] Blue Fjiord spiniger MATTHEW, 1889] New Brunswick,
Fm,Nunavut,Canada; PROHTIDAE; MDEV. Canada; ACROCEPHALITIDAE; UCAM [j.s.s.
Abadiella HUPE. 1953a [bourgini] Amouslek Fm, ofAcrocephalites, fideOPIK, 1967].
Morocco; ABADIELLIDAE; LCAM. Acanihometopus LU & QIAN, 1983a [obesus]
Abagnostus YANG & LIU in YANG el al., 1991 Changshan Fm, Liaoning, China; SOLENO-
[circularis] Baguamiao Fm, N Sichuan. China; PLEURIDAE: UCAM.
PERONOPSTDAE; MCAM |j.s.s. of Lisogor- AcanthomicniaccaHUPE, 1953a[Micmaccawalcotti
agnostus, fide SHERGOLD&LAURIE. 1997]. MATTHEW,1899b] Conception Bay. New-
Abakania POLETAEVA, 1973 [superba] Sanash- foundland. Canada; CHENGKOUIIDAE;
W
lykgol Horizon, Savan, Russia; CORY- LCAM.
NEXOCHIDAE;LCAM.' Acanthoparypha WHITTINGTON & EVITT, 1954
Abakanopleura SIVOV in EGOROVA et al, 1955 [perforata] Edinburg Fm, Virginia, USA;
[kenensis] Arbat Fm, \V Sayan. Russia; CHEIRURIDAE; UORD.
SOLENOPLEURIDAE; UCAM. Acanthophillipsia YUAN etal., 1992 [guiyangensis]
Abakolia BOGNIBOVA in WCHERNYSHEVA, 1971 Maokou Fin, Guizhou, China; PROETIDAE;
[paiica] Abakol Suite. Altay Sayan, Russia; LPERM.
WEYMOUTHIIDAE; MCAM. Acanthopleurella GROOM, 1902 [grindrodi] Bronsil
AbduHnaspis ERGALIEV. 1980 [concinnus] Tri- ShFm, England; SHUMARDIIDAE; LORD.
siilcagnosiiis trisiilcus Zone, Kazakhstan; Acanthopyge HAWLE & CORDA, 1847 [leuchten-
PARABOLINOIDIDAE;LORD. bergii] Koneprusy Lst, Czech Republic; LICH-
AbharellaWITTKE. 1984[magnocula]MilaFm,Iran; IDAE;LDEV
ANOMOCARIDAE;MUCAM. Acastava RICHTER & RICHTER, 1954 [Crvphaeus
Abia POLETAEVA in GONCHAROVA ct al.. 1972 atava W.E. SCHMIDT, 1907] Middle Heirdorf
[ritae] Salair, Russia; PTEROCEPHALIIDAE; Group, Sicgerland, Germany; ACASTIDAE;
MCAM
[j.s.sofLabiostrina,fideJELL,herein]. LDEV.
Acadagnostus KOBAYASHl, 1939a [Agnostus acad- Acaste GOLDFUSS, 1843 [Calymene? downingiae
icus HARTT in DAWSON. 1868] Eccaparad- MURCHISON, 1839] Much Wenlock Lst Fm,
oxides eteminicus Zone. New Brunswick, England; ACASTIDAE;LSIL.
Canada; PERONOPSIDAE; MCAM [j.s.s. of Acastella REED. 1925 [Phacops (Acaste) macro-
Peronopsis. fide ROBISON. 1995]. centrus]WhiteclifieFm.England;ACASTIDAE;
AcadolenusSDZUY. 1967 [decoais] EAsturia,Spain; USIL.
ELLIPSOCEPHALIDAE; MCAM. Acastellina RICHTER & RICHTER. 1954 [Acaste
Acaduparadoxidcs §NAJDR.1957b [Paradoxides (Acastella) nolens RICHTER & RICHTER,
sacheri BARRANDE, 1852] Jince Fm, Czech 1952] Heisdorfer Fm, Eifel, Germany;
Republic; PARADOXIDIDAE:MCAM. ACASTIDAE; LDEV
Acanthagnostus QLAN, 1982 [longispinus] Kushan Acastina REED, 1927 [unnecessary replacement and
Fm, Shandong, China; CLAVAGNOSTIDAE; thusj.o.s. ofAcaste],
UCAM [j.s.s.ofCIavagnostus,fideSHERGOLD AcastocephalaSHERGOLD. 1966 [Phacops(Acaste)
etal.. 1990]. downingiae var. macrops SALTER, 1864a]
Acanthaloma CONRAD, 1840 [suppressed by dec- Coalbrookdale Fm, England; ACASTIDAE;
ision of ICZN, for discussion see WHIT- LSIL.
TINGTON, 1952c] [j.s.s. of Leonaspis, fide AcastoidesDELO, 1935 [Acastehenni R. RICHTER,
WHITTINGTON,
1952c]. 1909]WetteldorfSst,RhenishMassif,Gennany;
Acanthalomina PRANTL & PRIBYL, 1949a ACASTIDAE; LDEV.
(Odonlopleura ininuta BARRANDE, 1846a] AcastopygeTOMC2YKOWAinTOMCZYKOWA&
Kopanina Fm. Czech Republic; ODONTO- WITWICKA, 1974 [shergoldi] Upper Podlasie
PLEURIDAE, USIL. Beds, Swarzewo. Poland: ACASTIDAE: USIL.
Acanthocephalus QIU in QIU et al., 1983 [longi- Acernaspis CAMPBELL, 1967 [Phacops orestes
PspRiOnAusS]APHHsIuScChIuDaAngE,; MFCm,AMJ.iangsu, China; BPIHLALCIONPGISD,A1E8;60aL]SIJLu.piterFm,Quebec,Canada;
Acanthocneinis HAWLE & CORDA, 1847 [ver- AcerocareANGELIN, 1854[ecome]Sandby,Sweden;
rucosa] Jince Fm. Czech Republic; OLENIDAE;UCAM.
SOLENOPLEURIDAE; MCAM [j.s.s. of Sao, Acerocarina POULSEN, 1952 [Cyclognathus micro-
fideSNAJDR. 1958a]. pygus LINNARSSON, 1875] Sweden; OLEN-
Acanthogramnia HAWLE & CORDA, 1847 [spec- IDAE; UCAM [replacement name for Cyclo-
PiSoNLsAaE]JUDRJRiIn,DceA19E5F8;ma,].MCCzeAcMh R[ejp.us.bsl.ico;fSSOaoL,ENfiOd-e AceroBgpnaralotkehhutasus]sh.,KKOaLzaOkBhsOtVaAn;,P1R9O8E8T[IacDeAr]E;AkLkSaInLs.Lst.
AVAILABLE GENERIC NAMES FOR TRILOBITES 337
Achatella DELO, 1935 [Dalmanites achates BIL- NW Siberia, Russia; ACROCEPHALITIDAE;
LINGS. 1860a] Cobourg Fm, Ontario; UCAM.
PTERYGOMETOPIDAE;LORD. Acrocephalites WALLERIUS, 1895 [Calymene
Acheilops ULRICH in BRIDGE, 1931 [dilatus] stenometopa ANGELIN, 1851] Vastergotland,
EminenceDol, Missouri, USA; CATILLICEPH- Sweden; ACROCEPHALITIDAE; UCAM.
ALIDAE; UCAM. AcroccphalopsPOULSEN, 1927 [gibber] CapeWood
Acheilus CLARK. 1924 [levisensis] Levis Fm, Fm.NWGreenland;BOLASPIDID.AE;MCAM.
Quebec.Canada; KINGSTONIIDAE; UCAM. AcrodirotesOPIK. 1967 [fastosa] Georgina Lst, Que-
Acheilus RAYMOND, 1924 [preocc., replaced by ensland, Australia; ACROCEPHALITIDAE;
UCAM.
Peracheilus].
AchlysopsisFRITZ, 1968[liokata]PiocheSh,Nevada, Acrohybus RAYMOND, 1937 [argutus] Rockledge
USA; PTYCHOPARIIDAE; MCAM. Cgte, Vennont, USA; HUNGAIIDAE; UCAM
Aciculolenus PALMER. 1965 [peculiaris] Elvinia [j.s.s.ofHungaia, fideJELL,herein].
Zone,Nevada, USA;OLENIDAE; UCAM. AcrolichasFOERSTE, 1919b[LichascucullusMEEK
Acidas[pliimdaetlal]aTPaOvgKiRysOkVySGKoAriYzAonti,n RSiObeZrOiaV,A,Rus1s9i6a3; U&SWA;ORLTIHCHEIND,AE1;86M8]OKRiDmm[sj.ws.is.cokfFAmm,phMiilsiscohuarsi,,
LICHAKEPHALIDAE; MCAM. fideWARBURG 1939].
Acidaspides LERMONTOVA, 1951a [precurrens] Acropyge QIAN. 1977 [multiscgmenta] Qinglong,
Boshche-Kulya, NE Kazakhstan; LICHA- Guizhou,China; PROETIDAE; UPERM.
KEPHALIDAE; UCAM. Actinobolus EICHWALD, 1858 [misspelling of
Acidaspidina LAZARENKO. 1960 [plana] Yenisey Actinolobus].
River, NW Siberia, Russia; LICHAKEPHAL- Actinolobus EICHWALD, 1858 [preocc., not replac-
IDAE; UCAM. ed] [j.s.s.ofIllaenus.fideJAANUSSON, 1954].
Acidaspis MURCHISON, 1839 [brightii] Much Actinopeltis HAWLE & CORDA, 1847 [caroli-
Wcnlock Lst Fm, England; ODONTOPLEUR- alexandri] Kraluv Dvur Fm, Czech Republic;
CHEIRURIDAE;UORD.
IDAE; LSIL.
AcidiphorusRAYMOND, 1925 [spinifer]TablePoint Actinopeltis POULSEN, 1946 [preocc., replaced by
Fm. Newfoundland, Canada; BATHYURIDAE; Grincllaspis].
MORD. Aculeodiscus SNAJDR, 1950 [bohemicus] Jince Fm,
Acidiscus RASETTI, 1966 [birdi] Acimetopus hiloh- Czech Republic; EODISCIDAE; MCAM. [j.s.s.
atus Faunulc, New York, USA; WEYMOUTH- ofDawsonia, fideJELL, 1997]
IIDAE; LCAM. Acutaspis REPINA in REPINA et ah, 1976 [facilis]
AciduascuisdusOuPs]IKC,una1n97t9Bu[sPhtyLcihmaegsntoosnteu,sQ(uAeceinsdluasnuds,) AShLeIvDliAnEs;kFLmC,ASMibe[rji.as.,s.RuosfsiTar;iaEngLuLlaIsPpiSsO,CJEEPLHL-,
Australia; PTYCHAGNOSTIDAE; MCAM 1997].
2[j0.s0.0s.ofPtvchagnostus,fidePENG&ROBISON, AcutaKtaazgankohssttuasn;ERAGGANLOISETVI,D1A9E8;0 [UacCutAatMus][jS.sa.ksi.ano,f
Acimebtiolpoub].sarRuAsSETFTaIu,nul19e6,6 [NbielowbatuYso]rAkc,imeUtSopAu;s AcutiAcgrnyopshtoupss, fCiRdeORNOIBEIRSO&N,FE1I9S8T8,].2000 [Phacops
WEYMOUTHIIDAE; LCAM. (Trimerocephalus) acuticeps KAYSER, 1889]
Acmarhachis RESSER. 1938b [typicalis] Nolichucky AOdPoIrDfAELs;t,UDREheVn.ish Massif, Gennany; PHAC-
AAccroenctFLthsmoet,p.uysSAgwlAeeaNdbHGeaAnmEa;WL,LICNUEOS.R&AY;1NC85AEO1GRXN[DOaOACcSu,HtTIa1InD8gD4uAAl7EuEis;;n]VMAUOnCCGdAArDMMaEr..Su,m AAcduatsitkKmoaaecnntcssoapapshs,uaesnlUsuSHimAsA;MHWBINERTLCA&LHCEEHHLRYAL.MH,ENT1,19O941P1499]I8D5[HtAae[EslCek;hoeetliPylrpEoiRpcLMyusmgt.],e
1925 [eiTorforAcanthopyge]. Elmside Fm. New South Wales, Australia;
Acrocephalaspina ERGALIEV. 1980 [insueta] PHACqPIDAE; LDEV.
Aksayan, Kazakhstan;EULOMIDAE; UCAM. Adelogonus OPIK, 1967 [solus] O'Hara Sh, Queens-
Acrocephalaspis IVSHIN. 1956 [fidus] Kujandinsk land, Australia; ??DAMESELLIDAE; UCAM.
IHoDrAiEz;onU,CceAnMtr.al Kcrzakhstan; SOLENOPLEUR- Aedotes OPIK, 1967 [instans] Mungerebar Lst,
Queensland, Australia; LEIOSTEGIIDAE;
Acrocephalclla ROZOVA, 1963 [granulosa] Ngan- UCAM.
AasLaInTsIkDHAoEr;izUonC.ASMi.beria, Russia; ACROCEPH- Aeglina BARRANDE, 1850 [replacement name for
Egle] [j.o.s. ofCyclopyge].
Acrocephalina TROEDSSON. 1937 [amiata] Tor- Aegrotocatellus ADRAIN & EDGECOMBE, 1995
suqlagh Fm, Xinjiang, China; DOKIMO- [jaggcri] Douro Fm, Nunavut, Canada;
CEPHALIDAE; UCAM. ENCRINURIDAE; USIL.
AcrocEepNhKalOin&ellNaIKMI.FROOROMVA,NE19N68KO[AcirnocLeAphZaAliRn-a Aegun[apsrpoisxiHmAa]JRSUdLzLuIvNeAllian-AReEgPuInNaAspeitsah,Zo1n9e7,5
borealicaLAZARENKO, 1960]KulyumbinaFm, Turkestan; DOLICHOMETOPIDAE; MCAM.
338 MEMOIRS OF THE QUEENSLAND MUSEUM
AAeeotnhieadiBoUnRidMcEIZSHTOERU,&184J3UIji-no.s.QoIfUPreotetuals.],. 1983 AgrauClzoepcshisReRpUubZlIicC;KAP,TYC19H4O0PA[RreIssIeDriA]EJ;inMceCAFmM,
[fusiformis]Huani»nchkangFm,Zhejiang,China; [j.s.s. ofPtychoparia, fide SNAJDR, 1958a].
DIONIDIDAE; UORD. Agraulos HAWLE & CORDA, 1847 [Arion
AethiCaouQnItAy,NJi&angZsHu,OUCh,in1a:98U4N[CrEecRtTanAgIulNa;] MKuCnAsMha.n cCezteicchepRheapluubsliBc;ARARGARNADULEI,DA18E4;6aM]C.lAinMce[Ffimrs,t
AethochuangiaQIAN, 994 [nodula] ChangshanFm, replacementnameforArion].
Liaoning,China; L1EiOSTEGIIDAE; UCAM. Aguaraya RUSCONI, 1954b [acutispina] Mendoza,
AfghanBLcOucRmpDhFa.ilni.tAefsghWaOniLsFtaAnR;TA.CR19O7C0E[PaHngAiiLsItuTsI]DSAurEk;h AguilaAFrrmeg,lelnAatrigBneaEn;tNiIEnNaDD;EEITTDT:AOIC,lAOM1l9.l7D7A[Et;apiUaCi]ASMa.nta Rosita
Afghanocarc WOLFART. 1974b [angustegenatum] Agvrciiella IVSHIN. 1957 [inacropleura] E/-/>/aZone,
Surkh Bum, Afghanistan; ANOMOCARIDAE; central Kazakhstan; UNCERTAIN; LCAM.
MCAM. Aiaiaspis NAZAROV, 1973 [nativa] Ust-Maisk Fm,
Afrops ALBERTI, 1983 [larvifer] Ben Zireg, SW SMECASMib.eria, Russia; SOLENOPLEURIDAE;
AgalaAKtlaugszearkiLahI;sStPOaHGnAO;CROO.PLIE1D9N6AI1ED;[AcELaD;veEmVLo.sOuRs]DKe[jn.ysi.st.kaso,f .AidarUKeulCllaAyaML..ENREMOKaNzTakOhVstAa,n;19S5O1aL[EvNigOiPlaLnEs]URBoIsDhAchEe;-
AgasefClInloHkaroEucRiLhaNEh.YaRfSmiMdHmeOeEWNrV.iTAZO,ZHVoAA1nN9eG5,&0,E[1Cf9aH8lS5cEi]ab.RteaN]riYaPS.arHaREduoVsxAsiidaei;sn AikhalARiuTesOslilVaaA:,OS1OG9LI8E7EN[NpOKaPrOtLitEai]UnCRhOIuGDkAIukEEs;NkKMFOmC,A&SME.SSiKbeUrRia-,
AGRAULIDAE; MCAM. Ainasuella MAKSIMOVA, 1978a [Phacops ainasu-
Agaso COBBOLD in COBBOLD & POCOCK, 1934 ePnHsiAsCBOAPLIDAASEH;OVLAD.EV1968] central Kazakhstan;
A[rGusRhAtoUnLenIsDisAiE:P MfCorAcMhh.ammeri Grit, England; Ainoa MANNIL, 1958b [maeruensis] Keila Stage,
AgelaQIgunDeiAeaEn;OsPlMIaCKnA,d,M1.9A6u7st|rqaulaidar;atCumA]TIMuLnLgIeCreEbPaHrALLst-, AistagECfisnhdtoaeosnntAiguDass;RhCAaXHnIIENAFI,NnRiGh,UerR&eXIiinDnZ]jA.HiEaA;nNgU,G,OCRh1iD9n8a[5;j.s[.Asl.aGeoNvfiOHgoaSltTiusa-],
Agerina TJERNVIK. 1956 [erratica] Billingen Gp, IDAE'; UCAM.
Niirke. Sweden: LEIOSTEGIIDAE; LORD. Ajacicrepida W. ZHANG & JELL, 1987 [Arionellus
AgnosQtuaerednisslaOnPdI.K,Aus1t9r6a3lia[;amGplLiYnaPtTusA]GGNeOoSrgTiInDaALEst;, CajhaixnaW;A?LCCEORTATT,01P9Y05G]1CDhAaEngThMiaCAFmM,.Shandong,
UCAM. AjrikinaKRYSKOVinBOROVIKOV&KRYSKOV,
Agnostascus OPIK, 1967 [gravis] Mungerebar Lst. 1963 [bulakensis] southern Kazakhstan;
Queensland, Australia: AMMAGNOSTIDAE; ALSATASPIDIDAE; UCAM.
UCAM [j.s.s. ofProagnostus, fide SHERGOLD Akantharges PHLEGER, 1936 [Lichas gourdonni
ctaL, 1990]. BARROIS, 1886a| Schistes a trilobitcs de
Agnostoglossa OPIK. 1967 [bassa] Mungerebar Lst; Catherviellc. Pyrenees, France; LICHIDAE;
AMMAGNOSTIDAE; UCAM. [j.s.s. of MDEV
Ammagnostus. fideSHERGOLDetal., 1990] Akbashichia KOROBOV, 1989 [plana] Akbashinsky
Agno.stogoinis OPIK, 1967 [incognitus] O'Hara Sh, Fm, W Mongolia; REDLICHIINA FAMILY
Queensland. Australia; AGNOSTINA FAMILY UNCERTAIN; LCAM.
UNCERTAIN; UCAM. Akoldinioidia ZHOU & ZHANG, 1984 [pustulosa]
Agnostonymus OPIK, 1979 [Onymagnostus (Agn- Fengshan Fm.Hebei,China;SHUMARDIIDAE;
oslonymus) semiermis] Currant Bush Lst, UCAM [j.s.s.ofKoldinioidia,fideSHERGOLD,
Queensland. Australia; PTYCHAGNOSTIDAE, 1991.
MCAM [j.s.s. of Onymagnostus, fide SHER- AksapygeLISOGOR, 1977a[MacropygefoveataLIS-
GOLDelal.. 1990]. OGOR. 1970] Shabaklin Fm, Kazakhstan;
AgnostotesOPlK, 1963 [inconstans] PomegranateLst, CERATOPYGIDAE;UCAM[j.s.s.ofPromacro-
Queensland, Australia; DIPLAGNOSTIDAE; pyge, fide PENG, 1992].
UCAM. Aksayaspis G. ERGALIEV in ERGALIEV & ERG-
AgnostusBRONGNIART. 1822[Enlomostracitespisi- ALIEV, 2001 [convexus] Zhumabaisk Fm,
fomiis WAHLENBERG, 1821] Kinnckulle,Vas- Kazakhstan;CHEILOCEPHALIDAE; UCAM.
tergotland, Sweden; AGNOSTIDAE; UCAM. Aksuaspis ZHOU et al., 1998b [aksuensis] Qiulitag
Agrauloidcs HOWELL, 1937 Agraulos socialis Group, Xinjiang, China; BATHYURIDAE;
BILLINGS, 1874] Trinity B[ay, Newfoundland, LORD.
Canada; AGRAULIDAE; MCAM [j.s.s. of Aktugaielia ERGALIEV, 1980 [acuminulata] Tri-
Agraulos, fide HENNINGSMOEN in MOORE, siilcagnostus trisulciis Zone, Kazakhstan;
1959], REMOPLEURIDIDAE; LORD.
AVAILABLE GENERIC NAMES FORTRILOBITES 339
Alacephalus REPINA, 1960 [contorlus] Lower [j.s.s. of Jujuyaspis. fide NIKOLAISEN &
Kameshkov Fm, SW Siberia, Russia; EDEL- HENNINGSMOEN, 1985].
STEINASPIDIDAE; LCAM. Allobodochus OPIK, 1979 [Ptychagnostus (Goni-
Alanisia HUPE, 1953a [Camaraspis guillermoi agnostus)fumicolaOPIK, 1961] DevoncourtLst,
RICHTER&RICHTER, 1940]AlaniaSh,Spain; Queensland, Australia; PTYCHAGNOSTIDAE;
ESTAINGIIDAE; LCAM. MCAM.
Alaskadiscus S. ZHANG in W. ZHANG etal, 1980a AlloctopsRAFINESQUE, 1832 [seeNOTE 2].
[Neocobbolciia spinosa PALMER. 1968] Adams Ailoillaeniis LIU. 1982 [glaberous] Xinghuang,
Argillite,EAlaska;YUKONIIDAE; LCAM. Hunan,China; ILLAENIDAE; LORD.
Alaskalethe HAHN & HAHN. 1991 [mufneri] -Alloleiostegium ZHOU & ZHANG. 1978 [Leio-
Ladrones Lst. SE Alaska, USA; PROETIDAE; stegium (Alloleiostegium)punctatum] Yehli Fm,
PENT^. Hebei,China;LEIOSTEGIIDAE;LORD[j.s.s.of
Alataupleiira LISOGOR, 1977a [duoculiensis] Leiostegium, fideZHOU&J.ZHANG, 1984].
'Puyuksaisk Fm, Kazakhstan: ALSATASPID- Allolichas KRUEGER, 1992 [longispinus] Backstein
IDAE;LORD. Lst, Teschendorf. N Germany; LICHIDAE;
Alataurus REPINAin REPINAelal., 1964 [menneri] UORD.
Bazaikh Horizon, Kuznetsk Alatau, Russia; Alokistocarc LORENZ,1906 [Conocephaliles sub-
PALAEOLENIDAE; LCAM. coronatus HALL & WHITFIELD, 1877]
AlbansiaHOWELL, 1937 [pusilla] StAlbansSh,Ver- ALOKISTOCARIDAE; MCAM.
mont, USA; SOLENOPLEURIDAE; MCAM. Alokistocarella RESSER, 1938b [typicalis] Rutledge
Albertella WALCOTT. 1908 [helena] Gordon Sh, Fm, Virginia. USA: ALOKISTOCARIDAE;
Montana.USA;ZACANTHOIDIDAE;MCAM. MCAM.
AlbertellinaPALMERinPALMER&HALLEY, 1979 Alokistocaropsis REPINA in SHIRAKOVA & REP-
[aspinosa] Carrara Fm, Nevada, USA; ZAC- INA 1964 [elegans] Kuznetsk Alatau, Siberia,
ANTHOIDIDAE; MCAM. Russia;ALOKISTOCARIDAE; LCAM.
Albertclloidcs FRITZ. 1968 [mischi] Carrara Fm, Alomataspis OPIK, 1967 [enodis] Steamboat Sst,
Nevada,USA:ZACANTHOIDIDAE; MCAM. Queensland, Australia; PROASAPHISCID.AE;
Albertitoryphc ERBEN. 1966 [Astycoryphe cogneyi MCAM.
ALBERTI, 1964] Ghlira Lst, NW Morocco; Alreboaspis PRIBYL & VANEK in PRIBYL et al.,
TROPIDOCORYPHIDAE; LDEV 1985b [Borealaspis biformis LUDVIGSEN,
Alborsella KUSHAN. 1973 [stoecklini] Alborz Mtns, 1976] Whittaker Fm. Northwest Territories,
Iran; PTYCHASPIDIDAE; UCAM. Canada; CHEIRURIDAE; UORD [j.s.s. of
AlcaldopsARBIZU, 1979 [alcaldei]CantabrianMtns, Borealaspis, fideHESSIN, 1989].
Spain; ACASTIDAE; MDEV [j.s.s. of Alsataspis TURNER, 1940 [bakeri] Marathon Lst.
Heliopyge, fideMORZADEC, 1983]. Texas, USA; ALSATASPIDIDAE; LORD [).s.s.
Alcestc HAWLE& CORDA, 1847 [latissima] Kraluv ofSeleneccmc. fide WHITTINGTON. 1952].
Dvur Fm. Czech Republic; STYGINIDAE; Altaepeltis MAKSIMOVA. 1968 [Weberopeltis
UORD. bublitchenkoi MAKSIMOVA, 1960] Kur'msk
Alcymene RAMSKOLD et al., 1994 [Calymcnc neo- Fm. RudnyAltai,EKazakhstan; STYGINIDAE;
inlcrmedia RICHTER & RICHTER, 1954] MDEV.
HemscFm.Gotland,Sweden;CALYMENIDAE; Altaesajania MAKSIMOVA, 1978a [Phacopidella
USIL. primitiva MAKSIMOVA, 1960] Altay Sayan, S
Aldaiiaspis LERMONTOVA. 1940 [punctata] Para- Siberia. Russia; PHACOPIDAE;MDEV
doxicies forchhummeri Zone, Aldan reg^ion. Altaiaspis IVSHIN in KHALFIN, 1960 [altaiensis]
Yakutia; SOLENOPLEURIDAE; MCAM. Salair, Russia; EULOMIDAE; UCAM.
AldonaiaLERMONTOVA, 1940[omatajKetemeFm, Altajaspis ELKIN, 1965 [mirabilis] Yaagvrsk Fm,
Siberia,Russia: ALDONAIIDAE;LCAM. Altay, Russia; PROETIDAE; USIL.
Alekcinella IVSHIN, 1962 [delicata] Seletinsk AltikoliaHAJRULLINAin HAJRULLINA&YASK-
Horizon,Kazakhstan:DOKIMOCEPHALIDAE; OVICH, 1961 [posochovae] Mumbel Pass,
UCAM. Tadzikhistan; PTYCHOPARTIDAE; LCAM.
Alemtejoia KOBAYASHl. 1943a[Microdiscussouzai .Altiocculus SUNDBERG 1994 [dmmensis] Swasey
DELGADO, 1904] Spain; HEBEDISCIDAE, Lst, Utah, USA; ALOKISTOCARIDAE;
LCAM [j.s.s.ofDclgadella, fideJELL, 1997]. MCAM.
Aliger HOWELL, 1937 [preocc. replaced by Altiplanelaspis PRIBYL & VANEK, 1980 [palqu-
Aligeritesj. ensis] Kainella meiidionalis Biozone, Bolivia;
Aligerites HOWELL, 1942 [Aliger venustus HOW- UNCERTAIN; LORD.
ELL, 1937] St Albans Sh, Vermont. USA; Altitudella REPINA in REPfNA et al., 1974 [tenera]
UNCERTAIN; MCAM [replacement name for Sektensk Fm, Siberia, Russia; PTYCHOPARI-
Aliger]. IDAE; LCAM.
Alimbetaspis BALASHOVA, 1961 [kelleri] Akty- Alueva SDZUY, 1961 [undulata] Murcro, Spain;
ubinskregion, Kazakhstan; OLENIDAE; LORD ELLIPSOCEPHALIDAE; LCAM.
340 MEMOIRS OF THE QUEENSLAND MUSEUM
Alw>nulus TRIPP, 1967 [peregrinus] Upper Stinchar Sweden;LICHIDAE;UORD[replacementname
‘ Lst Scotland; ENCRINURIDAE; MORD. forParalichas].
Amblycranium ROSS, 1951 [variabile] Garden City Amphion PANDER, 1830 [preocc., replaced by
Fm, Idaho,USA; HYSTRICURIDAE; LORD. Pliomera].
Ambonolium RAYMOND. 1924 [liodenna] Gorge AmphitryonHAWLE&CORDA, 1847 [murchisoni]
Fm.Vermont,USA;ILLAENURIDAE;UCAM. Kraluv Dvur Fm. Czech Republic; REMO-
Ambonslium KOBAYASHl, 1941 [misspelling of PLEURIDIDAE;UORD.
Ambonolium]. Amphoriops H/\MMANN & LEONE, 1997 [Thale-
Amecephalina POULSEN, 1927 [mirabilis] Cape ops inflata HAMMANN, 1992] Cystoid Lst,
Wood Fm, Greenland; ALOKISTOCARIDAE; Arag6n, Spain; STYGINIDAE;UORD.
MCAM. Amphoton LORENZ, 1906 [steinmanni] Changhia
Ameccphalites EDDY & McCOLLUM, 1998 [sund- Fm,Shandong,China;DOLICHOMETOPIDAE;
bcrgi] Pioche Sh. Nevada, USA; PTYCHO- MCAM.
PARIIDAE;MCAM. Amphotonella KOBAYASHL 1942a [Dolicho-
Amecephaloides REPINA in REPINA et al., 1964 melopus alceste WALCOTT, 1905] Changhia
A[lsaotlaoun,tzReunsssiisa];ASLoOloKnItSsoTvOCHAoRriIzDoAn.E;KLuzCnAeMt.sk FMmC,AShMan[djo.sn.sg.,oCfhAinmap;hoDtOoLnI,CfHidOeMWE.TZOHPAIDNAGE&;
Amecephalus WALCOTT, 1924b [Ptychoparia JELL, 1987].
piochensis WALCOTT, 1886] Chisholm Fm, Ampullatocephalina LU & QIAN, 1983a [bifida]
Nevada,USA;ALOKISTOCARIDAE;MCAM. Changshan Fm, Liaoning, China; LEIOSTEGI-
Amechilus ROSS, 1951 [palaora] Garden City Fm, IDAE;UCAM.
Idaho,USA; UNCERTAIN: LORD. Ampulliglabella KOBAYASHl & HAMADA, 1984c
Americare LOCHMAN & HU. 1960 [Glyphaspis [kojimai] Kanokura Fm, Japan; PROETIDAE;
lelonensis RESSER, 1937b] Death Canyon Fm, PERM.
Wyoming, USA; ANOMOCARELLIDAE; Ampyx DALMAN, 1827 [Asaphus (Ampyx) nasutus
MCAM [j.s.s. ofGlyphaspis, fide MELZAK & DALMAN, 1827] Vastana, Sweden; RAPHIO-
WESTROP, 1994]. PHORIDAE;MORD.
Ameropiltonia BREZINSKI. 2000a [lauradanae] Ampyxeila DEAN, 1960 [Ampyx (Raphiophorus)
Compton Lst, Missouri, USA; PROETIDAE; edgelli REED. 1932] Onnian, England;
MISS. RAPHIOPHORIDAE; UORD.
Ameura WELLER. 1936 [Phillipsia (Griffithides) Ampvxina ULRICH, 1922 [bassleri] Thebes Sst,
sangamonensis MEEK & WORTHEN. 1865] Illinois,USA; RAPHIOPHORIDAE;UORD.
Lower Foil Scot Lst, Missouri; PROETIDAE; Ampyxinella KOROLEVA, 1959 [Ampyx rugosus
PENN. kOLOVA. 1936] Dzhebaglin Horizon,
Amgaspidella CHERNYSHEVA in KHALFIN, 1960 Dzhebagly Mountains, N Kazakhstan;
[elongata] Amgaian, Siberia, Russia; AMGAS- RAPHIOPHORIDAE;UORD.
PIDIDAE; MCAM. Ampyxinops W. ZHANG, 1979 [Ampyxina biloba
Amgaspis CHERNYSHEVA in CHERNYSHEVA et CHUGAEVA, 1958] Kopalin and Karakan
" aAl.M,GA19S5P6ID[mIeDdAiuEs;j MACmgAaMi.an, Siberia, Russia; PHoHrOizRoInDs,AECh;u-MIlOiRMDtns[,j.s.Ksa.zoafkhMsatlaonn;guRllAiPaH,IfOid-e
Amginia PEGEL, 1981 [cxpressa] Tankhaisk Fm, ZHOUetal., 1995].
Siberia.Russia: ANOMOCARIDAE;MCAM. Ampyxoides WHITTINGTON, 1965a [Ampyx
Amginoerbia CHERNYSHEVA in EGOROVAet al., semicostatusBILLINGS, 1865] TableCove Fm,
1976 IselectajElanskoe Fm, Lena River, Siberia, Newfoundland. Canada; RAPHIOPHORIDAE;
Russia; DINESIDAE; LCAM. MORD.
AmginouyiaCHERNYSHEVAetal., 1956[elegans]E Amquia RASETTI, 1946b [truncata] Levis Fm,
Yakutia, Russia;UTIIDAE;MCAM. Quebec,Canada;LLANOASPIDIDAE;UCAM.
Amiaspis LOCHMAN in LOCHMAN 8c DUNCAN, Amurticephalus SHAH et al., 1991 [elongatus]
1944 [erralica] Pili:rim Fm, Montana, USA; Nutunus Fm, Kashmir, India; AGRAULIDAE;
LONCHOCEPHAUrDAE;UCAM. MCAM [j.s.s. of Parachittidilla, fide JELL &
Amicus KOROLEVA, 1967 fmontanus] central HUGHES. 1997].
Kazakhstan;CYCLOPYGIDAE;UORD. Amzasskiella POLETAEVA in KHALFIN, 196W0
Ammagnostus OPIK, 1967 [psammius] O’Hara Sh, [mirabilis] Algainsk Fm, Gorny Shoria,
Queensland. Australia; AMMAGNOSTIDAE; Siberia, Russia;EULOMIDAE; LORD.
MUCAM. AnabaracepsREPINA, 1972[kharaulachiensis]jPara-
AmorphcUa ROZOVA. 1963 [modesta] Yurakiysky micmacca Zone. NE Yakutia; PARADOXID-
Horizon, Siberia, Russia; APHELASPIDIDAE; IDAE;MCAM.
UCAM. Anabaraspis LERMONTOVA, 1951b [splendens]
Amphilichas RAYMOND, 1905a [Platymetopus Keteme Fm, Siberia, Russia; PARADOXID-
lineatus ANGELIN, 1854] Boda Lst, Dalame, IDAE; MCAM.