Austroalpine Liassic Ammonites from the Adnet Formation (Northern Calcareous Alps)

49 Pages·2012·4.64 MB·English

©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at
Jb. Geol. B.-A. ISSN 0016-7800 Band 136 S.163-211 Wien, Juli 1993
Austroalpine Liassic Ammonites
from the Adnet Formation
(Northern Calcareous Alps)
By CHRISTIAN MEISTER & FLORIAN BÖHM *)
With 14Text-Figures and 9 Plates
Oslerreich
Salzburg
Oberöslerreich
Nördliche Kalkalpen
Lias
Ammoniten
Oslerreichische Karle 1.50.000 Biostratigraphie
BI/1l1er94,95,96,126 Palaeogeographie
Contents
Zusammenfassung 163
Abstract. ................................................................................................................ 164
Resume 164
1. Introduction 164
2. Geographical and Geological Framework 164
3. Lithological Description and Qualitative/Quantitative Ammonite Distributions 165
3.1. Schmiedwirt Quarry 165
3.2. Breitenberg Quarry 166
3.3. Rotkogel Outcrop 166
3.4. Rötelstein Outcrop 168
4. Systematic Palaeontology 169
PhylioceratinaARKELL 1950 171
LytoceratinaHYATT1889 174
Ammonitina HYATT1889 175
5. Biostratigraphical Framework 184
5.1. Sinemurian 184
5.1.1. Early Sinemurian 184
5.1.2. Late Sinemurian 184
5.2. Pliensbachian 184
5.2.1. Early Pliensbachian (Carixian) 184
5.2.2. Late Pliensbachian (Domerian) 186
5.3. Toarcian 186
6. Faunal Composition and Palaeogeographical Remarks 189
7. Conclusion 190
Acknowledgements 190
References 208
Oberostalpine Liasammoniten aus der Adnetformation
(Nördlichen Kalkalpen)
Zusammenfassung
Das Oberostalpin spielt eine Schlüsselrolle für das Verständnis der Verteilungsmuster der jurassischen Ammonitenfaunen und für die Fixierung
genauer biostratigraphischer Korrelationen zwischen Tethyaler und Euroborealer Faunenprovinz.
Eine mittelliassische Ammonitenfauna (mehr als 500 Exemplare), die wir an vier Lokalitäten der Nördlichen Kalkalpen (Salzburg und südliches
Dachsteingebiet) aus Adneter Schichten aufsammelten, erlaubt es uns, 21 Horizonte oder biostratigraphische Faunen-Horizonte aufzustellen. Diese
fügen sich zum Teil indie nordwesteuropäische Standardzonierung, zum Teil indieder Tethysregionen. DieSemicostatum-, Obtusum- und Raricosta-
tum-Zone des Sinemur und die Jamesoni-, Ibex-, Davoei- und Margaritatus-Zone des Pliensbach sind durch die Ammonitenfaunen belegt.
Die Faunenzusammensetzung läßt deutlich den Tethys-Charakter des Oberostalpins erkennen. Daneben sind aber infast allen Horizonten eurobo-
reale Einflüsse erkennbar, besonders durch das Auftreten von Asleroceras aft. confusum und aff. sieilare, Tragophylloceras, Plalypleuroceras, Uplonia,
Acanlhopleuroceras, Liparoceratidae, Prodaclylioceras davoei,Amaltheidae, P(Malleiceras) und P(Fieldingiceras).
*) Author's addresses: Dr.C.MEISTER,Museum d'Histoire Naturelle deGeneve, Department of Palaeontology and Geology, 1RtedeMalagnou, cp434,
CH-1211 Geneva 6; Dr.F.BÖHM, Paläontologisches Institut, Universität Erlangen, Loewenichstrasse 28, 0-8520 Erlangen.
163
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Abstract
The Upper Austroalpine occupies akey position for the understanding of the patterns of Liassic ammonite distribution and for making precise
correlation between the Tethyan and Euroboreal realms.
Thecollection of aMiddle Liassic ammonite fauna (more than 500 specimens) from four localities inthe Northern Calcareous Alps (Salzburg area
and southern Dachstein) allows us to establish aset of 21 horizons or biostratigraphicallevels for the Adnet Formation. These units are readily
correlated with parts of the NWEuropean standard zonation and parts of the zonation used for the Tethyan realm. The Semicostatum, Obtusum and
Raricostatum zones for the Sinemurian stage andthe Jamesoni, Ibex, Davoei and Margaritatus zones for the Pliensbachian stage areidentified inthe
formation with ammonites.
The faunal composition clearly indicates the Tethyan affinities of the Upper Austroalpine; but this alpine unit is constantly subject to Euroboreal
influences, marked especially bythe presence ofAsterocerasaff. confusumand aff. stel/are, Tragophylloceras, Platypleuroceras, Uptonia, Acanthopleuroceras,
Liparoceratidae, Prodactylioceras davoei, Amaltheidae, P(Matteiceras), P(Fieldingiceras).
Les ammonites austroalpines de la Formation d'Adnet au Lias
(Alpes calcaires septentrionales)
Resume
L'Austroalpin superieur occupe une position clef pour lacomprehension des "modes" dedistributions des ammonites et pour I'etablissement de
correlations precises entre lesdomaines tethysien eteuroboreal. Larecolte d'une faune d'ammonites duLias moyen (plus de500exemplaires) dans 4
localites desAlpes Calcaires du Nord (region deSalzburg et partie meridionale des Dachstein) nous apermis demettre enevidence uneserie de21
horizons ou niveaux pour laformation d'Adnet. Ces unites biostratigraphiques s'integrent en partie dans la zonation standard du nord-ouest de
l'Europe etenpartie dans lazonation utilisee pour Iedomaine Tethysien. Ainsi leszones äSemicostatum, Obtusum et Raricostatum pour IeSinemu-
rien etleszones äJamesoni, Ibex, Davoei etMargaritatus pour IePliensbachien sont attesteespar lapresence d'ammonites.
Lescompositions fauniques montrent clairement lesaffinites tethysiennes del'Austroalpin superieur. Toutefois cette unite alpine reste constam-
ment soumise aux influences euroboreales avec enparticulier Asteroceras afl. confusum etafl. stellare, Tragophylloceras, Platypleuroceras, Uptonia, Acan-
thopleuroceras, Liparoceratidae, Prodactylioceras davoei, Amaltheidae, P(Matteiceras) etP(Fieldingiceras).
1. Introduction tary permo-mesozoic cover of the Upper Austroalpine pa-
laeozoic basement. Today the Northern Calcareous Alps
The Liassic of the Upper Austroalpine unit of Austria are in an allochtonous position, sheared off from their for-
was the subject of numerous palaeontological studies mer basement and transported to the north by the Creta-
mainly during the second half of the 19th and the begin- ceous orogeny. Internally the Northern Calcareous Alps
ning of the 20th century. Most of the famous outcrops for themselves are apile of allochthonous nappes.
Liassic ammonites are situated in the Salzburg area
Two of our sections (Schmiedwirt and Breitenberg) lie
(Salzkammergut, Tennengau) at the Adnet quarries, and
within the Osterhornscholle, which is part of the Tirolic
the Schafberg north of Lake Wolfgang (HAUER,1853, 1854
nappes. The third (Rotkogel) belongs to the Höllenge-
a,b, 1856; SUESS & MOJSISOVICS, 1868; WÄHNER,
birgsdecke, another part of this nappe complex. The sedi-
1882-1898, 1886, 1903; GEYER,1893; ROSENBERG,1909;
ments ofTirolic nappes were situated on the middle part of
PIA, 1914; BLIND, 1963; SIEBER,1961, 1975; WENDT, 1971;
the Northern Calcareous Alps before their tectonic dis-
SCHÄFFER& STEIGER,1986). The southern Dachstein area
placement. The fourth section (Rötelstein) is part of the
with another less famous but very prolific site for ammo-
southern Juvavic nappes (Hallstätter Schollen). The Ju-
nites, was the subject of the works ofTRAUTH (1925), TOLL-
vavic nappes are supposed to have formed the southern
MANN(1960) and HIRSCHBERG&JACOBSHAGEN(1965).
rim of the carbonate platform of the Northern Calcareous
Although the Lias of Austria is very fossiliferous, sur-
Alps during the Upper Triassic (LIEN, 1987). There may
prisingly the biostratigraphy in this "key" region has never
have been an ocean or at least an area of thinned crust
been accurately studied to understand the patterns of am-
further south of this zone (Text-Figs. 1and 14).
monite distribution and to make precise correlations be-
During the Liassic period the distance between the
tween the Tethyan and Euroboreal realms sensu DOM-
Tirolic and Juvavic realms may have been about 100 km
MERGUES& MEISTER(1991, p. 267). This is partly explained
(SPENGLER,1956), while the distance today is only about
by the often condensed levels and the scarcity of conti-
30 km north-south. Little is known about the area be-
nous sections. In our work on Sinemurian and Pliensba-
tween the Northern Calcareous Alps and the European
chian ammonites we seek to give adetailed stratigraphical
continent during the Liassic because most of the rocks of
description of four sections from different tectonic units of
this region are metamorphosed or overthrust by the Aus-
the Northern Calcareous Alps in order to improve the bio-
troalpine nappes. Most probably there was no oceanic
stratigraphical framework of this period of time. From our
crust there at that time but a rather shallow (up to a few
data it is also possible to make some remarks on the pa-
hundred meters) sea and possibly some land.
laeo- and biogeographical situation of the Upper Aus-
troalpine unit. The Adnet Formation, the source of most of our sam-
ples, is characterized by red, micritic, partly nodular
limestones. It is of Liassic age and is partly comparable to
the Ammonitico Rosso Inferiore of the Southern Alps. At
2. Geographical the Rötelstein, we also found some ammonites inthe grey
and Geological Framework Fleckenmergel. Inthis study we concentrate on the Upper
Sinemurian and Pliensbachian parts of the sections, ex-
The four sections studied are situated inthe middle part cluding the earlier Liassic and Toarcian, which may be the
of the Northern Calcareous Alps, which are the sedimen- subject of future work.
164
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Gmunden
•
Reichraming
Nappe
Osterhorn
Block
Hagen- Tennengebirge
Block
Rötelstein
DBaJuvarle and
Tiro1le Nappes
Northern Su_vie
• Nappes
20 km ITTTIlTTl Southern Su_vie
WlllllJ (Hallstatt) Nappes
Upper Austroalpine Southalpine
~~-~H-e1lvetie ~~ Pmeersmozoo-ic [[]]] mPeersmozoo-ic V.•............•.1APllpuitd.oicnes
_ Penninie 1:-:-1 Pelaeozole t-:-~Basement.
Basement.
~Lo_r
~ CrY"talline
~ Aust.roalplne
50 km
Text-Fig. 1.
Themiddle part ofthe Northern Calcareous Alps (upper) with the locations ofthe sections studied (above).
Tectonic overview ofthe Eastern Alps (below).
Thesections belong tothree different tectonic units. Schmiedwirt, Breitenberg andRotkogellie within different parts oftheTirolic nappes, Rötelstein
belongs tothe Juvavic Hallstatt Nappe (after TOLLMANN1,976a; GWINNER,1978).
3. Lithological Description
and Qualitative/Quantitative Ammonite Distributions
3.1. Schmiedwirt Quarry part of the famous quarries of Adnet, the type section of
the Adnet Formation. The quarry is still occasionally used.
The Schmiedwirt quarry lies near the old Wiestal Road, Quarrying is a very old tradition in the Wiestal. It can be
5 km NW of Hallein, SE of Salzburg, between the traced back at least to the 15th century (KIESLINGER,
Schmiedwirt and the Bischoff inn (Text-Fig. 2). The site is 1964).
165
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Text-Fig.2.
LocationmapoftheSchmiedwirtquarrynearAdnet.
about 500 msoutheast of the summit of the Breitenberg at
-
approximately 1150 m (Text-Fig. 4). The Breitenberg oc-
München
curence belongs to the same tectonic unit as that in the
Schmiedwirt quarry. Both belong to the Osterhornscholle,
which is part of the Tirolic nappe complex. As at Schmied-
wirt the beds are flat lying, and there is only minor tectonic
disturbance. There are two publications on the quarry:
one by SUESS& MOJSISOVlcs (1868), who gave a detailed
description of the stratigraphic sequence, but concen-
trated on the Upper Triassic strata. The other was by BUND
(1963) who worked on the Lower Liassic ammonites. There
are two geological maps by PLÖCHINGER(1973, 1982) cov-
ering this area. The lithology and succession (Text-Fig. 5)
of the section are very similar to those of Schmiedwirt
quarry. Again most of the rocks are red micritic limestones
Little detailed geological or palaeontological work had of the Adnet Formation. They are less nodular but more
been done in this quarry before. HAUER(1856) in his work marly than at Schmiedwirt. For our study only the upper
on Liassic cephalopods described some samples from part of the section is of interest. It starts with two thick
the Schmiedwirt quarry. SIEBER (1961) also mentioned beds of crinoidal micrite. There follows asuccession up to
some ammonites from this site in a short note. There are 20 cm thick of slightly nodular, thin-bedded limestones
also geological maps by SCHLAGER& SCHLAGER(1960, with marl interbeds. On top there is a packet of slightly
1 : 10.000) and PLÖCHINGER(1987,1 :50.000). brecciated, nodular, platy limestone, poor in microfossils
The thickness of the outcropping section isabout 20 m. and free of ammonites.lt iscapped bya pebbly marl with a
It comprises most of the Early Jurassic. Below the quarry mixed fauna of Carixian to Domerian ammonites (layer 18).
floor there are afew meters of bedded, grey, cherty lime- The section ends with a massive nodular breccia bed half
stones, which crop out along the way to the quarry. They ameter thick. There are no exposures above it, so it is not
are probably of Hettangian age. They are capped by alay- clear how much is missing between the breccia and the
erof reddish crinoidallimestone, ayellow-red hard ground radiolarite of probable Upper Jurassic age, which is ex-
and agrey breccia with clasts of grey limestones contain- posed at the top of the Breitenberg quarry.
ing yellow chert nodules (base of section inText-Fig. 3).
Above the breccia, reddish-grey, thin-bedded limestone
without chert (layer 2 in Text-Fig. 3) passes into red lime-
stone of the Adnet Formation with thin marly layers. Incon- 3.3. Rotkogel Outcrop
trast to the Adnet Formation at its type section in aquarry
at Adnet (3 km to the southeast) there are few beds with The Rotkogel outcroup east of Bad lschl (Text-Fig. 4)ex-
nodular fabric. The Adnet Formation at the Schmiedwirt poses on its eastern side a section of Middle Liassic red
quarry is a biodetrital wackestone with crinoidal debris Adnet Formation. It rests on a thick series of grey marly
(enriched in some layers), ostracods and foraminifera. limestones of Sinemurian age (Fleckenmergel, [SCHÄFFER
Within the uppermost five meters of the section the rock & STEIGER,1986]). There is a geological map (SCHÄFFER,
becomes more marly and nodular (layers 8-10, 12and 13 1982), but apart from brief account of SCHÄFFER&STEIGER
inText-Fig. 3)with lenses of fine-grained breccia. The top (1986) not much has been written about this locality and
bed is a breccia up to 1.5 m thick (layer 14 in Text-Fig. 3), its stratigraphy.
which lies unconformably on the underlying strata. Above The tectonic situation inth.isarea is much more complex
are afew centimeters of red marl and about half ameter of than in the Osterhornscholle. The Rotkogel block rests on
manganiferous red limestone, badly exposed, which may the Tirolic Höllengebirgsdecke, which is the eastern
be referred to the Upper Liassic or possibly Middle Juras- equivalent of the Osterhornscholle. It is not quite certain
sic. The radiolarite, which is attributed to the Oxfordian in that this block is really a part of that nappe. If so, then its
this region, is only found in boulders just above the man- palaeogeographic position was probably similar to that of
ganiferous limestone. the Osterhornscholle during the Liassic.
All of the sediments forming this section were deposited The section starts with reddish marly bedded lime-
on aslightly inclined slope at awater depth of afew hund- stones (Text-Fig. 6). The marl content is even higher than
red meters, below storm wave base and the photic zone. at the Breitenberg. In the upper part of the section three
breccia beds occur, separated by deep red marls and mar-
ly limestones, which became of to their high content of
fine crinoidal debris have a "sandy" appearance. This is a
very typical facies for the middle and higher Liassic of the
Tirolic nappes. It has been also found in the Osterhorn-
3.2. Breitenberg Quarry
scholle, where it has been called "Adneter Mergel" (Adnet
The outcrop at the Breitenberg, as at Schmiedwirt, is in Marls) by TOLLMANN(1976 b) and thoroughly described by
an old quarry, but this one is abandoned. The quarry lies PLÖCHINGER(1975), who called it "Saubachschichten" .
166
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Text-Fig.3.
Schmiedwirt quarry.
SCHMIEDWIRT Lithological profile andam-
monite ranges.
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Angulallcaras sp,(1),Apoderocaras (Mlltocaras) juv. (1),Juraphyllitas ~ Marly limestone
Pnaarrdtsllch(1ic)a,rJausra8pthrylallllotacsostlalbtuamrtus(5lu),mTernospllsdo(c1e)r,aPs.(aZfle.togcr,azraltste)llg(r2.)z,etas (2), ~- Calcareous marl
Mataderoceras af!.gr.mutlcum (1)Inthescrees. !!!!
Layers ofmarl
167
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1 km
1 km Rettenbaoh
Rotkogel
t t
Breitenberg Schön-Alm
10 km Loser
t
Text-Fig.4.
Location mapfor the Breitenberg (upper part left) andthe Rotkogel (upper right).
Thelower part shows anoverview ofthearea.
3.4. Rötelstein Outcrop strata. It is one of the best sections for Fleckenmergel of
the whole eastern Calcareous Alps. Only the base and top
Unlike the other three sections the Rötelstein (otherwise
of the section are cut out by thrusts, and there are some
"Rettenstein") is part of the so-called Hallstätter Schollen
minor displacements by blockfaulting.
or Southern Juvavic Nappes (Text-Fig. 1). These formed
the southernmost part of the Northern Calcareous Alps The outcrops are situated at the upper end of the
during the Triassic and probably Liassic. So the Rötelstein Weitenhausgraben between steep cliffs of Triassic dolo-
was in a different palaeogeographical position as com- mites below and Upper Jurassic limestone above. They
pared to the other localities (TOLLMANN,1981). There may can be reached from the west by following small paths
have been some emerged ridges between the Juvavic (this through alarge thicket of dwarf pines (Text-Fig. 7).
site) and the Tirolic (first three sites) depositional areas Previous publications include ageological map (GANSS,
during the Liassic, but this isnot very probable. The whole KÜMEL& SPENGLER,1954) with a detailed description of
area was probably adeep neritic or shallow bathyal sea at the section, and the papers of TOLLMANN(1960) and of
this time. HIRSCHBERG& JACOBSHAGEN(1965), discussing the strati-
Even though the tectonics are severe in this region, the graphy and listing numerous ammonites from the section.
outcrops show a rather undisturbed section of Liassic HIRSCHBERG& JACOBSHAGENsuggested a mixing of Ca-
168
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BREITENBERG
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Galaticeras sp. (1)Inthe screes
Text-Fig.5.
Breitenberg quarry.
Lithological profile andammonite ranges.
rixian and Domerian ammonites in the upper part of in microfacies between the grey and the red marls. The red
the section, which according to our results is not the marly limestone ends with a yellow-stained hardground.
case. Above there are about two meters of red marls with layers
The sequence starts with a thick succession of grey rich in shells of the small bivalve Bositra. They are capped
marls (Flecken mergel) of Lower to Middle Liassic age by radiolarite, locally developed as a breccia with meter-
(Text-Fig. 8). At the top they become slightly more calcar- sized blocks of grey shallow-water limestone. The breccia
eous and then pass abruptly into about 10 mof red, slight- may be of tectonic origin. The Upper Jurassic Plassenkalk
ly nodular marls and marly limestones, containing a very forms the cliffs above the section separated from by a
rich ammonite fauna. There is no obvious difference fault.
4. Systematic Palaeontology
Inthis paper we will not repeat descriptions and discus- DOMMERGUES& MEISTER, 1987 alb, 1989 alb, 1990 alb;
sions that are thoroughly detailed in previous works. For MEISTER& Loup, 1989; BLAU& MEISTER,1991. For certain
the well-known taxa we refer the reader to these studies: taxa we only give brief remarks and in most cases the taxo-
DOMMERGUESet aI., 1985, 1990; MEISTER, 1986, 1989; nomy is consistent with all these published studies.
169
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1 1
Text-Fig.6.
Rotkogellithological profile andammonite ranges.
.
Dachstein
......•.
"
.Rötelstein
Text-Fig. 7.
Location mapfor the Rötelstein.
Thedashed areasbelow andabove theoutcrops arecliffs.
170
©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at
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Text-Fig. B.
Rötelstein.
Lithological profile andammonite ranges.
Moreover because of our often badly preserved and Typ e spee ies: Ammonites heterophyllus SOWERBY1820.
fragmentary material, it is unfortunately impossible to ob- Rem ark: The proliferation of taxa of the long-ranging
tain good and acute measurements for the ammonites. Phylloceratidae provides a problem for each stage or
substage (Upper Sinemurian, Car[xian and Domerian).
Indeed many forms with the same morphology have dif-
ferent taxonomic names only because of their different
Suborder: Phylloceratina ARKELL 1950 ages. This results in a stress of biostratigraphical con-
Superfamily: Phyllocerataceae ZITTEL 1884 siderations rather than on morphological criteria for this
long-ranging family.
Family: Phylloceratidae ZITTEL 1884
Often the morphological variability that can be ob-
Subfamily: Phylloceratinae ZITTEL 1884 served over a long time range does not exceed the in-
Genus: Phylloceras SUESS 1865 traspecific variability that we can find in the Ammoni-
171
©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at
tina. So for us Zetoceras, Calliphylloceras and Calaiceras are L0caI ran ge: Obtusum zone (confusum horizon) .... Gib-
considered at best as sub-genera of Phylloceras and each bosus subzone (ragazzoni horizon).
of them probably is characterized by few species. In-
deed the main morphological differences are for Calli-
phylloceras the presence or persistence of constrictions,
for Zetoceras the compressed whorl section and for Calai- Subgenus: Calliphylloceras SPATH1927
cerasthe broad whorl section. Moreover they are often in
association in the same beds. Ty pes pecies: Phylloceras disputabile ZITTEL1869.
Inother terms, the genera or subgenera of the Phylloce-
ratidae would probably better be considered as spe-
cies, and the numerous "species" names of Phylloceras, P. (Calliphylloceras) bicicolae(MENEGHINI1874)
Zetoceras, Calliphylloceras, Calaiceras show only the morpho-
Plate 1, Figs. 2,5
logical intraspecific variability.
*1874 Phylloceras Bicicolae MENEGHINI,p. 106.
1989 Calliphylloceras bicicolae (MENEGHINI).- MEISTER,PI. 2, Fig. 3,
4with synonymy.
1991 Phylloceras (Calliphylloceras) bicicolae (MENEGHINI). - BLAU &
MEISTER,PI. 1, Fig. 6-9; PI. 2, Fig. 1.
Phylloceras gr. frondosum (REYNES 1868)
This regularly strongly constricted ammonite is rather
Plate 2, Figs. 1,2
common in our outcrops. Again the proliferation of taxa
*1868 Ammonites frondosus REYNES,PI. 5, Fig. 1. does not give agood image of the "reality". In our opinion
Ammonites Hebertinus REYNES,PI. 2, Fig. 3. P(C.)bicicolae includes nearly all of the "species" as descri-
1884 Phylloceras Meneghinii GEMMELLARO,PI. 2, Fig. 13-17. bed by BRAGA& RIVAS(1987). Only perhaps P (C.) dubium
1986 Phylloceras frondosum (REYNES).- GAKOVIC,PI. 1, Fig. 1.
(FUCINI),a more constricted form or P (C.) stoppani (MENEG-
Phylloceras hebertinum (REYNES).- GAKOVIC,PI. 1, Fig. 2.
1989 Phylloceras frondosum(REYNES). - MEISTER,PI. 2, Fig. 1,2 with HINI),a more compressed one, could be a different spe-
synonymy. cies.
Phylloceras hebertinum (REYNES). - MEISTER, PI. 2, Fig. 5, 7 L0caI ran ge: Luridum subzone (Reynesocoeloceras hori-
with synonymy.
zon) ....Gibbosus subzone (algovianum horizon).
We regroup, into this species, quite "globose" and more
compressed morphologies of typical Phylloceras (Text-
Fig. 9 C, D), often in association in the same beds (MEI-
STER,1989). This may be the expression of a sexual di- Subgenus: Zetoceras KovAcs 1939
morphism, indeed during the Pliensbachian, very often we
have the association of both morphologies . Ty pes pecies: Ammonites zetesd'ORBIGNY 1850.
.w
n ,Q
.Q
,n. ,n
.n
(J
Text-Fig.9.
Whorlsections(x 33).
A = P. (letoceras) gr.zetes(d'ORBI-
GNV)B, = Galaticerassp.indet.,CD
n = Phylloceras gr. frondosum (REV-
NES), E,F = P. (Calaiceras) calais
(MENEGHINIG),= "Metaderoceras"
venustulus (DUMORTlER), H = Proto-
..n grammoceras gr. dilectum (FUCINI),
. . I,J = Protogrammoceras gr. volubile
./\. (FUCINI-) panlanelli (FUCINI)K, =
"Protogrammoceras" gr. costicilla-
tum (FUCINI),L = Asteroceras gr.
slellare (SOWERBV), M,N,0=Astero-
cerasaft. confusum SPATH, P=Aste-
rocerasgr.quadragonatum (HVATT).
172