ASTERACEAE FROM WOOL MILL SITES IN SOUTH CAROLINA, INCLUDING NEW RECORDS FOR NORTH AMERICA

2004·3.7 MB·English

by  Guy L Nesom

ASTERACEAEFROMWOOLMILLSITESIN
SOUTHCAROLINA,INCLUDINGNEWRECORDS
EORNORTHAMERICA
GuyL.Nesom
BotanicalResearchInstituteofTexas
509PecanStreet
FortWorth,Texas76102-4060,U.S.A.
ABSTRACT
Documentationisgivenfor73taxaolAstcraceaecollectedin1957-1960fromwoolmillsitesinBer-
keleyCountyandFlorenceCountySouthCarolina.Twenty-eight(28)ofthesetaxaarefirstreports
forthefloraotSouthCarolina;seventeen(17)othersrepresentthefirstdocumentationtorearlier
reportsforthestate.Six(6)arefirstre|iortsforNorthAmerica:Chcvreuliasarmfntosa.,CHumnhaeta
ai'i^erttinu,andPanphaleaheterophyllii(nati\'etoSouthAmerica),andPeripieuraarida.Sluartina
bamata,andVittaJiniasulcata(nativetoAustralia). Calotiscuneijoiia(nativetoAustralia)isre-
portedforNorthAmericaforthesecondtime.Fifty-nine(39)ofthe73taxaarenotnativetoSouth
Carohna:oithese.29arenativetocontinentsotherthanNorthAmerica,while30arenativetoNorth
Americabutfromregionsoutsideotthestate.
RESUMEN
Seofrecedocumentacionde73taxadeAsteraceaecolecradosenelintervalof957-1960enloscondados
deBerkeleyyFlorenceCountyCarolinadelSurVeintiocho(28)deestostaxasonprimerascitaspara
lafloradeCarolinadelSur;otrosdiezysieie(17)representanlosprimerostestigosparacitasprevias
enelestado.Seis(6)sonprimerascitasparaNorteAmerica:Chevreuliasarmentosa,Gamochaela
argeniina,andPanphaleaheterophylla(nativadeSurAmerica),yPeripieuraarida,Stuartinahamata,
yVittadiniasulcata(nativadeAustralia).Calotiscuneijoiia(nativadeAustralia)secitaparaNorte
Americaporsegundavez.Cincuenta\'nue\'c(59jdclos73taxanosonnatu'osdeCarolinadelSur;dc
estos,29sonnativosdeotroscontmentesdilerentesdeNorteAmerica,mientrasque30sonnativos
deNorteAmericaperoderegionesfueradelestado.
Intheyears1957through1960,aspartoftheFloraoftheCarolinasprojectand
workingfromtheUniversityofNorthCarohna-ChapelHillherbarium,Harry
AhlesandcompanionJohnHaesloopmaderepeatedcollectingtripstotwo'wool
combingmills'inSouthCarolina.OneofthesewasinJamestowninBerkeley
Co.,theothernorthofJohnsonvilleinFlorenceCo.;bothsitesarelocatedon
thecoastalplain,40-50kilometersfromthecoastandabout60kilometersfrom
eachother.Someofthecollectionswereidentifiedandincorporatedintothe
generalcollectionsatNCUseveraldecadesago;othersweremountedbutwere
notidentifiedorinsertedintothegeneralcollectionsuntil2001.Onlyafewof
thesewereincludedintheaccountoftheFloraoltheCarolinas(Radfordetal,
1968).Somehavebeenthebasisofrecentreports(withoutdocumentation)for
distributionrecords(Kartesz1999andothers).
SIDA21(2):1215-1223.2004
1216 BRIT.ORG/SIDA21(2)
Thewoolcombingmillsreceivedrawfleeceandprocessedittowardpro-
ductionofcleanwoolforspinning.Earlystagesintheprocessaremechanical
shaking(removesdirtandsomeplantmatter),"scourmg"(removeschemical
substancesandsand),and"combing"(removesfinerplantmaterialaswellas
shorter,weakerfibers).WastefromtheseprocessesattheSouthCarolinamills
wasthesourceofpropagulesforspeciescollectedbyAhlesandHaesloop.De-
liveryoftherawwoolprobablywasbyrailroad,becausebothofthesitesare
alongtheCSXRailroad,whichrunsroughlyparalleltothecoast.Bothmills
apparentlyarestillinoperation.
Seventy-threetaxa(73)ofAsteraceaein54generaarerepresentedinthecol-
lectionsidentifiedhere.Twenty-eight(28)ofthesearefirstreportsforthefloraof
SouthCarolina;seventeen(17)othersrepresentthetirstdocumentationforear-
lierreportsforthestate.Six(6)arefirstreportsforNorthAmerica:Chevreulia
sarmentosa,Gamochaetaargentina,andPanphalcaheterophyllaarenativeto
SouthAmerica;Peripkuraarida,Stuartinahamata,andVittadiniasulcataare
nativetoAustralia.Calotiscuneifolia(nativetoAustralia)isreportedforNorth
Americaforthesecondtime.Fifty-nine(59)ofthespeciesarenotnativetoSouth
Carolina;ofthese,29arenativetocontinentsotherthanNorthAmerica,while
30arenativetoNorthAmericabutfromregionsoutsideolthestate.
ItwouldbeusefultoreinvestigatethestatusofthesetaxainSouthCaro-
lina,inordertodeterminewhichhavepersistedorincreasedtheirdistribution.
Anumberofthetaxareportedheremayhavethepotentialtospreadandbe-
comedamagingweeds,ortheymaysimplybewaifs(scnsuNesom2000).
CollectionsbyAhlesandHaesloopweremadein1957(April,May,July,Sep-
tember,October),1958(April,IMay,June,July,September),1959(May,July),and
1960(May,July).Fromeachofthetwowoolmillsites,collectionswereprovided
withessentiallythesamelabeldata,exceptforthecollectionnumberanddate.
SOUTH CAROLINA. Berkeley Co.:wastegroundaroundSanteeWool
CombingMill,JamestownonS.C.f^te.45,[Apr-Oct1957-1960fH.E.Ah/csjcol-
lectionnumber]withJ.G.Haesloop.
SOUTHCAROLINA.FlorenceCo.:wastegroundaroundtheWellmanWool
CombingMill,northofJohnsonvilleonS.C.Rte.41,lApr-Oct1957-1960],H.E.
Ahfcstcollectionnumber]withJ.G.Haesloop.
PLANTLIST
Thegeneraandspeciesarearrangedalphabetically.Ifmorethanonecollection
wasmadeforasinglespecies,thecitationsarearrangedchronologically.Vouch-
ersarelocatedatNCU,unlessotherwiseindicated.
AchilleamillefoliumL. U.S.A.;previouslyknownfromSouthCaro-
BerkeleyCo.:13Jun]958,Ahles42939 lina(Radfordetal,1968).
BerkeleyCo.:27May1959,Ahles52717 AmblyolepissetigeraDC
NativetowesternAsia,widelynaturalizedinthe BerkeleyCo.:8Apr1957,Ahles22614
NESOM,ASTERACEAEFROMWOOLMILLSITESINSOUTHCAROLINA 1217
NativetoTexasandMexico;firstreportforSouth BidensbipinnataL
Carolina. BerkeleyCo.:20May1957,Ahles25836(NCU,
ANBeamrtbli<prveroelesevtyioioaCNuoos.ar:lrtyt2he7kmAniMmosaeiwyrinif1ocfl9ar5io9aam,nLAd.Showlueitsdhe5lC2ya8rd0oi5lsitnraib(uRtaedd-; NBaetrikMSveeMelxeUity)coCoot;.hp:er1e4svoiMuoatuhsyleyr1n9k5n8a,ondwAhnleeafssrto4em0r4nS8o8Uu.tS.hA.Caarnod-
fordetal.1968). lina(Radfordetal.1968).
ABBBNaeeemtrrrpikkkChAhveeeeailelllrraaeeeeotyyycb.loahiCCCtmyooon...haar:::eia322bssn082yodduOSSTrKtceeeaahtppn-rcnctu111eeen999nss5c55tzs778ur,e,,la(eAloAA1;hhhi9llUl9dr.eee9eeSsss)p.s,A3o34.8r9(5dt2aD51oesC891cd556f)uafrNomrueetSantsottuetadhs FBNFFFallllitoooodirrrrekUiveeee.znnennnnSescccco.dteeeeAwpo.iinCCCCnMlooooa....teofn::::hsxrdea322i1o8204cmsLao.oOMMSSufceaaaotetpnyyuhwdtw1111he99oC9955set5587Ctn7h,,8aet,/e/rr\r^AroA/anh/lhilili/ls/aeenteeAssnsaasmtd34e42e(89r0s6Rs/i;/46ao5c34d9upa396ft9r,oherenvdaaitsoueturtesarllalny-.
AnthemiscotulaL. 1968).
BFBFNaeelltrrooikkrrveeeeellnneecctyyeeoCCCCEoooou....::::ro2221p0718eMMJMuaaaalnyyyd}9111A6999s055i5,a797,,,,AwhAAAlihhhedlllseeeelsss5y3252752n65a876t6299u770ralizedin BBNeairtdkiefvenrelosemtypooCSotlo.hyu:elt5ehepJaiuCslsatre]Bor9lln5ai7kn,Uea.AS(h.RAl.ae;dsfp3or0red8v0ieo9tusall.y19k6n8o).wn
theU.S.A.;previouslyknownfromSouth BidensfrondosaL
Carolina(Radfordetal.1968). BerkeleyCo.:22Sep1958,Ahles49204
ArctiumminusBernh. NativeandwidespreadinNorthAmerica;previ-
BerkeleyCo.:11Jul\96Q,Ahles53824 ouslyknownfromSouthCarolina(Radford
NativetoEurope,widelynaturalizedintheU.S.A.; etal.1968),
previouslyknownfromSouthCarolina(Rad- CaiotiscuneifollaR.Br
fordetal.1968). BerkeleyCo.:8Apr1957,Ahles22537
ABerrtkeNeamltieisyvieCaot.ao:n1en1ausJatuelrLn\9E60u,roAhpleesan5d38A4s8ia,natural- BBeerrkkNeeallteeiyyveCCoot..o::A11u4s3tJrMaualniya],19r95e58p8,o,ArAhthleledessi4n24t90h35e56,4U.4S2.A9.7o3nly
fiozredSoinutvharCiaoruoslisntaa.tesoftheUSA;firstreport ffoorrMSaosustahcChaursosleitnat.s(Sorrie1992);firstreport
ABBBNeeerarrrttkkkieCeeevmalllenieeeasyyydtiaoCCCa;ooof...wb:::ieir22e1ss107nttneJMMriureaalsnpyyoW\ir9a11lt6n99l0dd55f,.79or,,Avnha/ASol^nhorehlu/tsbeethiss5hee325rn8C25nna178ir469soU27l.iSn.aA,.and CBNaeatrrikGdlrveiaueulnllaeuitfyzsobecyCdnopooa.yKir:sancttr8nhtseoUAecr,cspaSnzter,tApA(,ef11h;rr99aie5r9cld9e7au),p,sas/ot^nrd/aLdtiot./eeecwdssue2smfo2tofe5ren5wrSt0enoesuAtdsteihrah,enCrneaaa.rtnoud--
ABBNaeertrrtikkeCveemaellineestyyaoidCCaEaoou..v::ruao22lnp55gdeaMMrtaiaahnsyyedL1]eAv99asa6i6sra00t,,,evwArAuihnhldllgeeeUassl.rSy55.iA33ns.a44;t00up32rraelviizoeudsliyn CBBeearrrkkS=teeRllhCeeeaauyymrtt.CCuoo)hs..a::Nmb21uya73smelJtMaauiannncyaut1s1u995s(58B9o,L,i,AssAshuh.lble&sesps.R45be2uo28te79.f7)9/4cLu(a5drea(tB.o(iDss.ysJn..
k19n6o8)w.nfromSouthCarolina(Radfordetal, BBeerrkkKeeellileeyy1CC9oo9..7::)2257MMaayy]1995690,,AAhhlleess5523835742
NativetosouthernEuropeandnorthernAfrica,
1
1218 BRIT.ORG/SIDA21{2)
naturalizedinscatteredstates.mostlyalong NativeandwidespreadinNorthAmerica;previ-
thePacificcoast;reportedforSouthCaro- ouslyknownfromSouthCarolina(Radford
linabyKartesz(1999),documentedhere. etal.1968).
CentaureaamericanaNutt CoreopsisverticiilataL
BerkeleyCo.:27May1959,Ahles52776 BerkeleyCo.:13Jun1938.Ahles42900
BerkeleyCo.:27May1959,Ahles52843 NativetothesouthernU.S.A.;previouslyknown
BerkeleyCo.:25May1960,Ahles53492 fromSouthCarolina(Radfordetal.1968).
NatifmvoeernSptroeiumdtahrhieClrayer.otolitnhaebcyenKtarratlesUz.S(.1A.9;99r)e,pdoorctue-d CBNaeotrtikuvelelaetyaouCsoAt.u:rsat8lriaAslpir(aSi]ae9nb5ed7r,)NAhHeloweoskZ.2e2(a.5l9a2nd;natural-
CentaureamelitensisL izedinvariouswesternU.S.A.statesand
BerkeleyCo.:8Apr]957,Ahles22624 Florida;firstreportforSouthCarolina.
NBBaeetrrikkoCnSveeatrhelltihoeeunentyyrnroqaeuCClssroioito.s.saz:)u:ttete1d1hs(34;1ei9JnrrM8unet0anph)Eyeo,u]rwd9rt1eo5oe9s8cdp5t,8euef,AmorahreAnnlhnSdelUots.enuSes4.ot2dAr4h9.t0lhahC5e9ane4rrrde2o.nlv(iaAdnfreraiti.ocbuaLys,. DBNiaettrtikZUrirven.eeielSapcve.ltohayAoari.rnaCtit(doeohG.g,ude:lrsaef2aMan8rrvesedodeoSgmionieCtloavpe&nelasrinr1C,fris9roaoi5roun(n7nnLsei,c.q)aalo/unuW^ti(/dGhsiPri/etrernergseegi1s3ulo9tAo5tno9uc5,ne1sa9ltrn;ir51aMta9tiil9ueti7srca)a,hi,lenNilatzlenhewd&de
CentaureasolstitialisL Tucker1997;Preston1997;Tuckcr1995);first
BerkeleyCo.:25MayI960,Ahles53487 reportforSouthCarolina.Inviewoftheother
BerkeleyCo.:11 Jul ]960,Ahles53846(det.L, speciesreported herefromAustralia,it
Shinners) seemslikelythatthewoolmillplantsof
NativetowesternAsiaandtheMediterranean Dittnchiagraveolensalsooriginatedfrom
regionofEurope,naturalizedandweedyin thereratherthanfromthenativeregionof
mostoftheU.S.A.;reportedforSouthCaro- thespecies.
linabyHillandHorn(1997). Dracopisamplexicaulis(Vahl)Cass.
Chaetopappaasteroides(Nutt) DC var. BerkeleyCo.:20May1957,Ahles25840
asteroides BerkeleyCo.:25May1960,/^/i/es53369
BerkeleyCo.:8Apr]957,Ahles22551 Nativeprimarilytothesouth<entralU.S.A.;re-
NativetonorthernMexicoandthesouth-cen- portedforSouthCarolinabyKartesz(1999),
tralU.S.A.;firstreportforSouthCarolina. documentedhere.
Chevreuliasarmentosa(Pers.)Blake Dyssodiapapposa(Vent.)A.S.Hitchc.
BerkeleyCo.:14May1958,Ahles40509 BerkeleyCo.:8Apr]937,Ahles22620
NativetoSouthAmerica(Brazil,Paraguay,Uru- BerkeleyCo.:20May1957,Ahles25892
guay,Bolivia,Argentina);firstreportforthe BerkeleyCo.:14May1958,Ahles40483
U.S.A.andSouthCarolina. BerkeleyCo.:14May1958,Ahles40553
CBeornkyezleaybCoo.n:ar1i4eMnasiys1(9L.5)8,CrAohnleqs.40568 NatifviresttoretphoretsfoourtShowuetshteCranrolainnda.centralU.S.A.;
BerkeleyCo.:21Jul]958,Ahles46996 EupatoriumdubiumWilld
NativetoSouthAmerica,naturalizedinmost BerkeleyCo.:15Jul1959,Ahles52950
southernstatesoftheU.S.A.;previously NativetoAtlanticcoaststatesoftheU.S.A.,from
knownfromSouthCarolina(f^adfordetal. SouthCarolinanorthward;prevlouslyknown
1968). fromSouthCarolina(Radfordetal.1968).
CoreopsistinctoriaNuttvartinctoria EvaxmulticaulisDC.
BerkeleyCo.:27May1959,Ahles52825 BerkeleyCo.:8Apr]9S7,Ahles22591
BerkeleyCo.:11Jul]960,Ahles53822 BerkeleyCo.:14May1958,Ahles4051
1 )
NESOM,ASTERACEAEFROMWOOLMILLSITESINSOUTHCAROLINA 1219
NativetoMexicoandvariousstatesofthesouth- GamochaetaargentinaCabr
ernU.S.A.;reportedforSouthCarolinaby BerkeleyCo.:8Apr1957,/^/i/es22594
Cronquist(1980),documentedhere. NativetoSouthAmerica(ArgentinaandUru-
EvaxproliferaNutt.exDC. guay);firstreportfortheU.S.A.andSouth
BerkeleyCo.:14May1958,/\h/es405/2,405/3 Carolina.
BerkeleyCo.:27May]959,Ahles52737 Gamochaetacalviceps(Fern)Cabr
NativetoMexicoandvariousstatesoftheGreat BerkeleyCo.:27May1959,/iWes52842
PlainsandsouthernU.S.A.;firstreportfor PerhapsnativetothesoutheasternUSA,appar-
SouthCarolina, entlynaturalizedworldwide;recognizedby
FBaecrekleilseyreCot.u:s2a7(MLaamy.)]S9c5h9u,lAthzl-eBisp5.2731 tbhyisNneasmoema(n20d04c)i.rcumscriptionintheU.S.A.
BerkeleyCo.:27May1959,Ahles52858 Gamochaetapensylvanica(WiId)Cabr.
NativetoSouthAmericaandnaturalizedin BerkeleyCo.:8Apr1957,/\/i/es22590
statesoftheGulfcoastandsouthernAtlan- ProbablynativetoSouthAmerica(Nesom2004),
ticcoast;previouslyknownfromSouthCaro- naturalized inthesouthern U.S.A.and
lina(Radfordetal.1968). throughouttheworld;previouslyknown
Flaveriatrinervia(Spreng.)Mohr fromSouthCarolina(Radfordetal.1968).
BerkeleyCo.:28Sep19-57,/I/i/e535584 Gamochaetasimplicicaulis(WilldexSpreng
BerkeleyCo.:21Jul]9S8,Ahles47035 Cabr.
BerkeleyCo.:22Sep1958,/^/i/es49/72 BerkeleyCo.:5Jul1957,/^Wes50825
BerkeleyCo.:22Sep1958,Ahles49215 BerkeleyCo.:21Jul1958,/\/i/es47038
NativetonorthwesternMexicoandthesouth- BerkeleyCo.:15Jul1959,Ahles52959
westernUnitedStates,naturalizedinafew BerkeleyCo.:11Jul]960,Ahles53783
statesofeasternU.S.A,;firstreportforSouth NativetoSouthAmerica,naturalizedinafew
Carolina. statesofAtlanticgulfcoastU.S.A.;previously
Gail(lHaorodki.a)Bp.uLk.hTuerlnlear Fougvar.drummondii kNneoswonmf(r1o99m9)S.outhCarolina,firstreportedby
BerkeleyCo.:8Apr1957,/^/i/es22528 GrindelialanceolataNutt.varlanceolata
BerkeleyCo.:20May1957,/l/-|/es25852 BerkeleyCo.:13Jun1958,Ahles42920
BerkeleyCo.:14May]958,Ahles40460 Nativetothesouth-centralU.S.A.;firstreportfor
BerkeleyCo.:13Jun1958,Ahles42902 SouthCarolina.Theplantissterile,butthe
BerkeleyCo.:13Jun1958,Ahles42950 identificationisprobablycorrect.Theplant
BerkeleyCo.:13Jun1958,/^h/e542969 hasrelativelybroadleaveslikeplantsfrom
BerkeleyCo.:25May1960,Ahles53367 eastTexas.
Nativeprimarilytothesouth-centralU.S.A.but Gutierreziasarothrae(Pursh)Britt.&Rusby
fnoartuSroaultizhedCairnolvianraioubsyoKtahretresszta(t1e9s9;9r)e,pdoorctued- BNeartkievleetyoCoM.e:x2i8cSoeapn1d95th7,e/\w/ie/sest3e5r2n69U.S.A.;first
mentedhere.Thecollectionsweredeter- reportforSouthCarolina.
minedbyannotationas\/ai.drummondiiby
B.L.Turner&T.J.Watsonin1995. Gutierreziatexana(DC.)Torr.&A.Grayvar.
texana
GBFelaromkroeelncechyeaCCeoo.t.::a88aAAnpptrril11l99a55n77a,,(AAUhhrllbee)ss22A22n5d62e29r8b. BNBeaertrkikeveleleetyyoCCMoo..e::x2i12c1oJSuealpn1d916T90e5,x8a,AshA,lhenlasets5u43r98al21i9z3edorpos-
ProbablynativetotheCaribbeanregionand siblynativeinafewotherstatesofcentral
ecprienrrchUuaSmpAss;cSrfoiirupsttthiroeAncmoegirnniictzahe,edpeUbr.ySh.taAhp.issbsnyoautmNheeesaasontmd- HelUe.nS.iAu.;mfirasmtarerpuormt(fRorafS)oHu.thRoCcakrovlairnab.adium
(2004). (A.GrayexS.Wats.)Waterfall
1 1
1220 BRIT.ORG/SIDA21(2)
BerkeleyCo.:20May1957,Ahles25829 NativetothesouthwesternU.S.A.;fir5treportfor
BerkeleyCo.:20May1957,Ahles25834 SouthCarolina.
BBeerrkkeelleeyyCCoo..::1143JMuany1199S588,,AAhhlleess4420943768 Hyptowceheadeeriis(Hboroaks.iSliiAemn.si)sBa(kLeesrs)Griseb var.
BNeartkiSmevoeleunetyttoheCodT.Ce:ahxr2eao7rsel.Mianaanydb1yO95k9Kl,aarAhthoelmseasz;5(21r79e29p97o)r,teddocfuo-r NBBBaeeetrrrikkkveeeellleeetyyyoCCCoooS...o:::u228t87hApSMAreampye1r91]i5997c55,a19,,A,hAnAlhaheltlseuesr2sa235l65i225z73e93d30instates
HeleniumelegansDCvarelegans alongtheGulfcoastandsouthernAtlantic;
BerkeleyCo.:25May]960,Ahles53357 previouslyknownfromSouthCarolina,first
NativetoTexas,Oklahoma,andLouisiana;re- reportedbyShinners(1966)fromawoolmill
portedforSouthCarolinabyKartesz(1999), collection.
documentedhere. HypochaerisglabraL
Helenium microcephalum DC. var BerkeleyCo.:14May1958,Ahles40469
microcephalum BerkeleyCo.:13Jun1958,Ahles42954
BerkeleyCo.:20May1957,Ahles25844(NCU, BerkeleyCo.:27May1959,/\h/es52758
SMU) NativetoSouthAmerica,naturalizedinstates
BerkeleyCo.:5Jul]957,Ahles30803 alongtheGulfcoastandsouthernAtlantic
BerkeleyCo.:21Jul]958,Ahles47002 andvariousothers;previouslyknownfrom
BerkeleyCo.:27May1959,/^Wes527/9 SouthCarolina(Radfordetal.1968).
BNFBFelaelrotrokrikreeveelnelnececyeyetoCCCCooooT....e::::x12a21115s6,JJMuMuOlaalky1yl]9a961160h990,o66,A00m,hA,alhA,AelhhsellNs5eees3s57w355583301M456e0(6xN50iCcUo,,SMaUn)d IBPBveerarrokkbapeearnllbeeenlvyyyuiaCCnoooau..tLs::ilv23ye80tOkSocenttpohew119n9c55e7n7,ft,rrA/ao^hl/lmiea/esnsSd3o58u5e6ta109sh4t6eCranrUo.lSi.An.a;
CKoarltoersazdo(1;99r9e)p,odrtoecdumfeorntSeoduthherCea.rolinaby (Anonymous1997),documentedhere.
BHeerlkiealnetyhCuos.:a2n0nMuauys1L.957,Ahles25862 NBIvaeatrikavexelieltyloaCrwoie.ss:tP1eu3rrsJnhuNnor\t9h58A,mAehrliecsa;4f2i9r6st1reportfor
FBelrokreelnecyeCCoo..::1223JSuenp1]995S88.,AAhhlleess4429918475 SouthCarolina.
BBBeeerrrkkkeeellleeeyyyCCCooo...:::221157JMMuaalyy1911699065,09,,Ah7/l\^/ehis//ees5s35575274/05767 BIBveearrkkxeeallneetyyhiCCoof..o::i2i17a4MMNuaatyyt]199S598,,AAhhlleess542085033
BerkeleyCo.:11JulI960,Ahles53820 NativetomoststatesoftheU.S.A.exceptthe
NativeornaturalizedacrosstheentireU.S.A., Southeast;firstreportforSouthCarolina.
previouslyknownfromSouthCarolina(l^ad- Krigiavirginica(L.)Willd.
fordetal.1968), BerkeleyCo.:8Apr1957,/\/i/ei22627
Heliomerismultiflora(Nutt)Blakevar.multi- IlorcnceCo.:8Apr1957,Ahles22546
flora(syn=Viguieramultiflora{Nun.)Blake) FlorenceCo.:9Apr1958,Ahles38274
BerkeleyCo.:14May1958,Ahles4054 NativetotheeasternU.S.A.;previouslyknown
BerkeleyCo.:27May1959,Ahles52778 fromSouthCarolina(Radfordetal.1968),
NativetothewesternUSAandMexico;firstre- LactucagraminifoliaMichx.
portforSouthCarolina. BerkeleyCo.:13Jun1958,Ahles42968
HymenoxysodorataDC Nativetocoastalstatesofthesoutheastern
BerkeleyCo,:8Apr\957,Ahles22512 U.S.A.;previouslyknownfromSouthCaro-
BerkeleyCo.:8Apr1957,/^/i/es22605 lina(Radfordetal.1968).
5
NESOM,ASTERACEAEFROMWOOLMILLSITESINSOUTHCAROLINA 1221
PanphaleaheterophyllaLess(identficaiionby U.S.A.;previouslyknownfromSouthCaro-
JohnPruski) lina(Anonymous1997),documentedhere.
BerkeleyCo.:8Apr1957,/\/i/e522585 SolivasessilisRuiz&Pavon (syn=Soliva
NativetoSouthAmerica(Brazil, Uruguay, Ar- pterosperma(Juss.)Less.)
gCeanrtoilnian)a;f(iPrrsutskriep&orNtefsorotmhe20U0.4S)..A.andSouth NBeartkievleeytoCo.S:o2u7thMaAyme1r9i5c9a,,Ahnlaetsur5a2l7i1z8edinthe
PartheniumhysterophorusL. southeasternU.S.A.;previouslyknownfrom
BerkeleyCo.:20May1957,Ahles25850 SouthCarolina(Radfordetal.1968).
BerkeleyCo.:5Jul]957,Ahles308W SonchusoleraceusL.
BBeerrkkeelleeyyCCoo..::2125JSuelp!199595,8,Ah/^lfet/ses54299/660/ NBaetrikveeletyoCEou.r:o8peA,pnror1t9h5e7r,n/\a/in/desw2e2s6t/ernAsia,and
ApparentlynativetotheWestIndiesandadja- northernAfrica,widelynaturalizedinNorth
centNorthAmericaandSouthAmerica, ArTierica;previouslyknownfromSouthCaro-
tnraatlurUa.lSi.zAe.dainndvaerlisoeuwshesrtaet;esfiorsftsroeuptohr-tcefno-r lina(Radfordetal.1968).
SouthCarolina. StuartinahamataPhilipson
Peripleuraarida(Burbidge)Nesom(syn= BNeartkiveeletyoCAou.s:t8ralAipar;fi1r9s5t7,reApholretsf2o2r5t9h3eU.S.A.and
VittadiniaaridaBurbidge) SouthCarolina.
BerkNealteiyveCot.o:A1u4stMraalyia1;9f5i8r,s/t\r/ie/peosrt40f5or4/theU.S.A. Symphyotrichumdivaricatum(Nutt.)Nesom
andSouthCarolina. (syn=AstefsubulatusMichx.var.Iigulatus
Shinners)
Pseudognaphaliumstramineum(Kunth) BerkeleyCo.:30Oct]957,Ahles38169
Anderb. NativetonorthernMexicoandthesouth-cen-
BerkeleyCo.:11Jul1960,/^/i/es53825 tralU.S.A.;firstreportforSouthCarolina.
NativetoMexicoandthewesternU.S.A.,natu-
ralizedinVirginia,NorthCarolina,andSouth TagetesminutaL.
Carolina;previouslyknownfromSouthCaro- BerkeleyCo.:20May1957,Ahles25845
lina(Radfordetal.1968). BerkeleyCo.:28Sep1957,Ahles35609
Ratibidacolumnifera(Nutt.)Woot.&Standi. NBaetrikveeletyoCoS.o:u2t2hSAemper1i9c5a8,,/iin/it/ehse49U./S8.SA.natural-
BBeerrkkeelleeyyCCoo..::2104MMaayy11995S87,,AAhhlleess4205486380 oizuesdlymakinnolwyninfrAtolmanStoicutchoaCsatraollistnaate(sR;apdrfeovrid-
BerkeleyCo.:27May19S9,Ahles52712 etal.1968).
BBBBBFNeeaeeelrrtrrrokkikkkrveeeeeeelllllneeeeectyyyyyeoCCCCCCooMoooo......e::::::x22222i1577551coMMMMMJuaaaaaalyyyyyn1d9111\1699999t066556h,00990e,,,,,/\cAAAAA/eihhhhh/nllllleteeeeesrsssssa5l355555723323U583483.0579S67.11100A.,widely TNBeearttkirlHKevaioalennoneeeksytaa.uors)rCi;oiTf.sef:oixlr8asiltsaiA,nrpee(rNaspreyo1inrw9fl5o=7lfM,ioeHraAxyhSi(mlocHeeuoosn,to2ohkO2x.kC5)yal1sra3Gohrllioiennmeeaaan.r,eifaovlnairda
naturalized;reportedforSouthCarolinaby Thymophyllatenuiloba (DC.) Small var.
Kartesz(1999),documentedhere. tenuiloba)(syn-^Dyssodiatenuiloba(DC.)
RudbeckiahirtaLvarangustifolia(IV.Moore) B.L.Robins,var.tenu//o6a)
Perdue BerkeleyCo.:8Apr1957,Ahles22580
BBBNeeearrrtkkkieeevllleeeeyyytoCCCooot...:::he221073coJMMauaasnyyta1l11999555s879t,,,aAtAAhehhlslleeesossf4252s529o881u636t30heastern NatiCGrvureelpofotnroqctuoMiafessoxtrtisSc1too9au8tate0nhs;dA(CeiTa.aerg.xboalaFsilm,noaarn,.iadbtaeuarlanolndigzMei1ds9s8ii8ns)s;aifpfipreisw,t
9
1222 BRIT.ORG/5I0A21(2)
Uropappuslindleyi(DC.)Nutt. VittadiniasulcataN.Burbidge
FlorenceCo.:26May1960,Ahles53543 BerkeleyCo.:8Apr1957,Ahles22538
NativetothesouthwesternU.S.A.;firstreportfor NativetoAustralia;firstreportfortheU.S.A.and
SouthCarolina. SouthCarolina.
Verbesinaencelioides(Cav.)var.encelioides XanthiumspinosumL
BerkeleyCo.:20May1957,Ahles25548 BerkeleyCo.:8Apr]957,Ahles22608
FlorenceCo.:28May}957,Ahles2671 BerkeleyCo.:20May1957,Ahles25872
BerkeleyCo.:5Jul1957,Ahles30812 FlorenceCo.:28May1957,Ahles26689
BerkeleyCo.:14May1958,Ahles40486,40554 BerkeleyCo.:30Oct1957,Ahles38199
FlorenceCo.:21Jul]958,Ahles46950 BerkeleyCo.:13Jun1958,Ahles42924
BerkeleyCo.:22Sep]958,Ahles49180 NativetoSouthAmerica,widelynaturalizedin
BerkeleyCo.:27May1959,Ahles52810 NorthAmerica;reportedforSouthCarolina
BerkeleyCo.:15Jul1959,Ahles52945 byKartesz(1999),documentedhere.
NFFalltooirrveeennccteeoCCMoo..e::x2i16c1oJMuaalyn1d9]6t906h,0eA,hwAleheslstees5r35n7365U24.S6.A.,natu^ XBearnkMteilhleliyeurCm)oT.o:srt5rr.J&uulmA.a]Gr95ri7au,ymAhLlesva3r08c4a8nadense(P
lmrUi.aenSla.ni,Azt.e;rededpiprnoheresvretievoe.eudrsalblyysktCanotoelwsennonffatrnhoem(s1S9o7ou4tu)ht,ehadsCotaceruro-n- NatiCavsaeroatloiwNneoaerd(t;RhadpAfrmoeervrdiioceutaslaaly.nd1k9nw68oi)dw.enlyfdriosmtriSbouuttehd
AC1<NOWI.HDC,Mf:NTS
1amgratelultorhelptromthestaffofherbariumNCVandforcommentson
themanuscriptbyLindaLee(USCIU.JohnPruskt(MO)providedtheidentifi-
cationofPanphaleahcLcrophylla.
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