DENVER MUSEUM OF NATURE & SCIENCE REPORTS NUMBER 3, JULY 2, 2016 Program and Abstracts 20th International Congress of Arachnology July 2–9, 2016, Colorado School of Mines, Golden, Colorado Edited by Preface Paula E. Cushing1 Welcome to the 20th International Congress of Arachnology! This congress is jointly hosted by the International Society of Arachnology, the American Arachnological Society, and the Denver Museum of Nature & Science. A total of 375 participants are in attendance representing 39 different countries. An additional 36 accompanying participants are taking advantage of Colorado’s scenery and charms. Of the registered participants 154 are students. This book contains the schedule of events, the scientific program, and the abstracts. In total, there will be five keynote addresses, nine formal sym- posia, 240 oral presentations (including symposium talks), and 129 poster presentations. The abstracts are arranged in alphabetical order by the surname of the presenting author. Above each abstract is an indication of the type of talk (i.e., keynote address, symposium talk, oral presentation, poster presentation. Student presentations that are part of the student competition are indicated with asterisks (*) in the abstract list and in the schedule. Speakers whose names are under- lined in the program are session moderators charged with keeping the session on time. Thanks to the following foundations and organizations for providing support for this meeting: ISA, AAS, DMNS, Kenneth King Foundation, Laudier Histology, Schlinger Foundation, Golden Chamber 1Department of Zoology of Commerce, Siri Publications, BioQuip, Cricket Science. Denver Museum of Nature & Science 2001 Colorado Boulevard On behalf of the Organizing Committee, Denver, Colorado 80205-5798, U.S.A. Welcome to Golden – Where the West Lives! [email protected] Paula E. Cushing Cushing 20th International Congress of Arachnology CONTENTS Preface................................................................................................... 1 Emergency Contact Information & Wireless ................................................. 3 Organizers and Committee Chairs .............................................................. 3 Proceedings of the 20th International Congress of Arachnology (2016) and guest access to New Zealand Journal of Zoology ........................... 4 General Information ................................................................................ 5 Oral Presentations ....................................................................... 5 Poster Presentations ..................................................................... 5 Student Awards ............................................................................ 5 Venue and Town ........................................................................... 5 Golden Ticket ............................................................................... 5 Vendors & Special Book Signing Event............................................... 5 Group & Committee Meetings ......................................................... 5 Arachnid Collecting....................................................................... 5 Reprint Table ............................................................................... 6 Silent Auction .............................................................................. 6 Alcohol Rules & Laws .................................................................... 6 Marijuana Laws ........................................................................... 6 Smoking in Golden ....................................................................... 6 Tubing in Clear Creek .................................................................... 6 Hiking on South Table Mountain ..................................................... 6 Social Events & Drink Tickets ......................................................... 6 Russian Party .............................................................................. 7 Check-out Items for Dorm Residents ................................................ 7 Promotion for the Next ICA and AAS meetings ................................... 7 Daily Schedule ........................................................................................ 8 IUCN Workshop Thursday, June 30–Saturday, July 2 ........................... 8 Registration and Welcome, Saturday, July 2 ...................................... 9 Sunday, July 3 .............................................................................. 10 Monday, July 4 ............................................................................. 13 Tuesday, July 5 ............................................................................. 16 Wednesday, July 6, Mid-Congress Excursions ..................................... 18 Thursday, July 7 ........................................................................... 19 Friday, July 8 ............................................................................... 21 Poster Session I, Sunday, July 3: List of Participants ...................................... 23 Poster Session II, Tuesday, July 5: List of Participants .................................... 27 Arachnid FilmFest Program ....................................................................... 31 Accompanying People Agenda .................................................................... 31 Post-Congress Excursion Agenda ................................................................ 32 Abstracts (Alphabetical by Surname of Presenter) ........................................ 33 Attendees................................................................................................ 209 2 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 3 Cushing 20th International Congress of Arachnology EMERGENCY CONTACT INFORMATION & Student Competition Committee Chairs: WIRELESS Richard Bradley, The Ohio State University, USA Yael Lubin, Ben Gurion University, Israel Emergency Contact Information: Committee members: Police / Ambulance Emergency dial 911 J. Andrew Roberts, The Ohio State University – Newark, Meeting Host, Paula Cushing: USA (student travel grant) Mobile Phone: 303-246-8382 Sarah Stellwagen, postdoc Army Research Laboratory, Home Phone: 303-273-5707 USA (student travel grant) Courtney Scheskie, DMNS Office Manager and Assistant Rainer Foelix, Aarau, Switzerland (Laudier Grant) Meeting Coordinator: Peter Michalik, Ernst-Moritz-Arndt-Universität Greifswald, Mobile Phone: 864-642-7468 Germany (Laudier Grant) Patrick Casto, DMNS Volunteer and ICA Meeting volunteer: Elizabeth Jakobs, University of Massachusetts Amherst Mobile Phone: 303-596-6382 (non-student travel grant) Wireless: Maggie Hodge, Louisiana School for Math, Science, and When on campus, open your browser. Choose CSM Guest the Arts, USA (non-student travel grant) (not CSM Wireless). Enter your personal email address. Al Cady, Miami University, USA (student research grant) This will serve as your password to connect to the internet. Bruce Cutler, University of Kansas, USA (student research grant) Eileen Hebets, University of Nebraska, USA (HLMFAR) ORGANIZERS AND COMMITTEE CHAIRS Paul Selden, University of Kansas, USA (HLMFAR) Todd Blackledge, University of Akron, USA (HLMFAR) Organizing Committee: Thanks also to numerous colleagues serving as Paula E. Cushing, Denver Museum of Nature & Science judges for the Student Awards! Courtney Scheskie, Denver Museum of Nature & Science Patrick Casto, Denver Museum of Nature & Science Post-Congress Excursion Guides: Paula Cushing, DMNS, USA Program Committee: Paul and Maura Selden, University of Kansas, USA Linda Rayor, Cornell University Courtney Scheskie, DMNS, USA Ingi Agnarsson, University of Vermont Thanks also to officers of the ISA and AAS for their ICA Website Management: assistance. Peter Michalik, Ernst-Moritz-Arndt-Universität Greifswald, Thanks to the following DMNS staff and volunteers for Germany helping with the Congress: Ryan Bartlett, Kate Baughn, Heiko Metzner, psbrands GmbH, Germany Gary Baxley, Jack Brookhart, Lynne Carpenter, Tim Chaney, Travel Grant Committee Chairs: Mark Christopher, Cat Davis, Jessica Ellis, Debra Freeman, Mike Draney, University of Wisconsin-Green Bay, USA David Footle, Mike Getty, Megan Gish, Karen Hall, Martin Heck, (AAS Student Travel Grant) Kathy Honda, Jennifer M. Jennings, Patsy Kelly, Eric Knutson, Damien Laudier, Laudier Histology, USA (Laudier Student Frank Krell, Efrain Leal, John Love, Jason Lucas, Gretchen K. Grants for Histological Research) Moritz-Hale, Tammy Ann Morris, Vicki Moyer, Gini Philipp, Karen Cangialosi, Keene State College, USA (Kenneth Rosine Ribelin, Nancy Roberts, Hilary Sachs, Steve Schilling, King and Schlinger Foundation Travel Grants for Nina Shilodon, Margo Stephenson, Jeff Stephenson, Chad Non-Students) Swiercinsky, Rick Teichler, Rebekah Thompson, Jill and John Weigel, Julie Whitman-Zai, Dennis Wojcicki. 2 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 3 Cushing 20th International Congress of Arachnology PROCEEDINGS OF THE 20TH INTERNA- a manuscript for submission. Congress manuscripts that TIONAL CONGRESS OF ARACHNOLOGY do not closely follow the Instructions to Authors may not (2016) be considered for publication. JOURNAL OF ARACHNOLOGY Authors who are not fluent in English are urged to have a native English speaker read, edit, and help revise The proceedings of the 20th International Congress of the manuscript prior to submission. Arachnology (ICA) will be published in 2017 in volume 45, number 2, of the Journal of Arachnology. Review. All Congress manuscripts will be reviewed in the The editors of the journal are pleased to invite same manner and with the same requirements for scien- submissions for the proceedings issue (details below). tific quality and content as are all other submissions to Note that, in addition to the usual research/data the Journal of Arachnology. papers, we also invite symposium organizers or participants to submit review papers keyed to the NEW ZEALAND JOURNAL OF ZOOLOGY topics in their symposia. New Zealand Journal of Zoology has graciously granted Open Access. All accepted ICA articles will be published free access to attendees of the ICA from 1–31 July 2016. as “immediate open access” papers, meaning that they To access this journal, go to: www.tandfonline.com/r/ will be openly downloadable from the JoA website upon NZ-Zoology publication, at no charge either to authors or to readers and without the need for a password. Submissions. All submissions to the Journal of Arachnology are now made online at http://www.edito- rialmanager.com/arachno/default.aspx When submitting a manuscript for the proceedings, be sure to choose “ICA Paper” when asked to “Choose Article Type.” Deadline. October 1, 2016. Manuscripts submitted after the October 1st deadline will be considered for later issues of the Journal. Format. Regular Congress manuscripts should be no more than 15 manuscript pages in length (double spaced, 12-pt font), not including figures and tables. For manuscripts longer than 15 pages, authors must contact the Managing Editor prior to submission. Review papers may be longer than 15 pages at the discretion of the Editor-in- Chief/Managing Editor. Instructions to Authors are available at http://www. americanarachnology.org/JOA.html#instructions and should be followed closely when writing and assembling 4 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 5 Cushing 20th International Congress of Arachnology GENERAL INFORMATION Golden Ticket. The Golden Ticket in your welcome package is accepted in restaurants and stores with Oral Presentations. Each speaker (except plenary/ a Golden Ticket in the window. Do not scratch off the keynote speakers) will have 15 minutes total including magic window until you are ready to pay your bill. questions and discussion. We have two to three concur- Vendors & Special Book Signing Event. We are rent sessions. Speakers will be asked to step off the stage pleased to welcome BioQuip and Cricket Science as at the 15 minute mark in order for us to stay on schedule our guest vendors. Vendor tables will be in the Grand and allow audience members to attend talks in concur- Lobby of the Green Center July 3rd, 4th, and 5th. On July rent sessions. 4th from 5:30 – 7:00 p.m. the following book authors Please prepare your presentation in MS Office will be at the BioQuip table to sign their books: Richard PowerPoint for Windows (preferably) and download it Bradley – Common Spiders of North America; Rainer to a USB memory stick. Presentation volunteers will be Foelix – Biology of Spiders; Heather Proctor – Mites: available to assist presenters to download presentations Ecology, Evolution and Behaviour; Justin Schmidt – no later than the break prior to their session. Earlier is The Sting of the Wild; and Richard Vetter – The Brown better. Recluse Spider. If you have your own copy of any Posters. Poster boards are situated in the Grand Lobby of these tomes, bring them to the meeting to get the of the Green Center. Posters should be no more than author’s signature and if you do not, BioQuip will have 4 ft x 4 ft (1.2 m x 1.2 m) in size. Poster presenters some copies of all these books available for purchase. may hang posters during the afternoon of July 3rd. Stop by the BioQuip table before heading over to the 4th Poster Session I presenters should be standing by their of July barbeque! posters no later than 7 p.m. July 3rd. Poster Session II Group & Committee Meetings. Please refer to the presenters should be standing by their posters no later schedule for the times and locations of the following than 7 p.m. July 5th. meetings: IUCN pre-congress workshop (June 30th– Student Awards. Awards for best student oral presen- July 2nd); Student Competition Judges meetings (July tations and best student poster presentations will be 3rd and 8th); dinner for those interested in morphology, given during the banquet the evening of July 8th at the histology, and fine structure organized by Damien Denver Museum of Nature & Science (DMNS). For both Laudier (July 3rd); AAS Executive Committee meeting oral and poster presentations, awards will be given in (July 3rd); students & early career social event (July the categories of Systematics, Ecology, & Biogeogra- 3rd); Journal of Arachnology Editors’ meeting (July phy; Morphology, Physiology, & Silk; and Behavior & 5th); AAS Business Meeting and promotion of the 2017 Ecology. AAS meeting in Mexico (July 7th); ISA Business Meeting and promotion of the 2019 ICA meeting in New Zealand Venue and Town. Maps of the Colorado School of Mines (July 8th). (CSM) campus and the town of Golden can be found in your welcome material. Most events will be held on the Arachnid Collecting. Permits are required to collect CSM campus except where noted in the program. The arachnids in County, State, and National Parks. No organizers encourage you to explore downtown Golden, permits are required to collect arachnids in National enjoy a stroll along the banks of Clear Creek, and sample Forests. Collecting is permitted in the high elevation the many restaurants and bars in town. During your free areas (Loveland Pass and Berthoud Pass) where mid- time, wander through the Pioneer “village” along Clear congress excursion participants will travel but is not Creek, see the exhibits at the Foothills Art Center, or go on permitted at Rocky Mountain Arsenal where those a Coors Brewery Tour. excursion participants will travel to see bison and 4 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 5 Cushing 20th International Congress of Arachnology wildlife. Meeting organizers will have some collect- in any way (smoking, eating, or vaping) is not allowed ing supplies, including alcohol, available during the in public places; use of pot on federal land (national excursions. parks, national forests) is still illegal; hotel owners can dictate rules for their own facility. Reprint Table. A reprint table will be set up in the Grand Lobby of the Green Center during the meeting. Reprints Smoking in Golden. The City of Golden passed a are free; however, donations are appreciated. Suggested smoke-free law in 2015 that prohibits smoking or donations are 25 cents US for regular reprints and 50 vaping on downtown area sidewalks and public rights cents - $2 for monographs. A donation jar will be on the of way on both sides of Washington Avenue between 10th reprint table. and 14th Streets, as well as both sides of 12th and 13th Streets between Arapahoe and Ford Streets, including Silent Auction. On July 7th, the Vendor Tables will be customer seating/service areas located on those public replaced with displays and bidding sheets of items for rights of way. sale during the silent auction. These items will remain on display in the Grand Lobby from 10 a.m. July 7th Tubing in Clear Creek. We will have a couple of until 1 p.m. July 8th. At 1 p.m., the silent auction items tubes and life jackets to borrow (for a $5 deposit) at will be packed up and moved to the Denver Museum of the Registration / Information Desk in the Grand Lobby Nature & Science. Silent Auction ends promptly at 6:00 for anyone interested in tubing on Clear Creek. You can p.m. at the DMNS. also rent tubes and life jackets in the Student Recre- ation Center. Remember that mountain streams are fast Alcohol Rules & Laws. Alcohol (the kind you drink) is moving and rocky. Please don’t drown (way too much prohibited in all University residence halls and adjacent paperwork!). public spaces. Open containers of alcohol are not permit- ted on public streets. Hiking on South Table Mountain. The hike up South Table Mountain (the mountain that the solifuge on the Marijuana Laws. The possession, use, manufacture, logo is standing on) takes about 30–40 minutes (one- or distribution of illicit drugs (including marijuana) in way). It is a fairly easy hike. You can access the trail by all University residence halls, public spaces, buildings walking down 19th Street towards the mountain. Beware and other facilities, is prohibited. No marijuana stores of rattlesnakes – they are abundant on the mountain are in Golden. If you want to take advantage of Colo- but they are not particularly aggressive and will rattle rado’s lenient pot laws, you will have to go to Denver in warning (usually). Please leave the rattlesnakes or areas outside Golden to find pot shops. And please alone and don’t get bitten (see parenthetical comment be aware that, according to Colorado law: you must be under tubing). 21 to buy, have, or use retail marijuana; it is a felony for anyone to give, sell or share marijuana with anyone Social Events & Drink Tickets. The four alcohol under 21; you must present a valid ID proving you are tickets received by each participant can be used for one 21 or older; Colorado residents can buy and possess alcoholic beverage at the socials on July 2nd, 3rd, 4th, or up to 1 ounce of marijuana at a time; adults with an 5th. Non-alcoholic beverages are free. The alcohol tickets out-of-state ID can purchase up to a quarter-ounce at may not be used for alcoholic beverages at the Russian a time; some stores may only allow cash purchase (no Party, at the Butterfly Pavilion, or at the DMNS banquet. credit cards); only licensed retailers can sell marijuana There will be cash bars at the Butterfly Pavilion and at products; adults 21 or older can give up to 1 ounce of the DMNS. If you use up all your red drink tickets we will marijuana to another adult but cannot sell it; having have a volunteer drink vendor on hand to sell you more more than one ounce of marijuana in your possession tickets ($5 US each) that you can exchange for a drink can result in legal charges and fines; using marijuana on July 2nd, 3rd, 4th, or 5th. 6 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 7 Cushing 20th International Congress of Arachnology The following ICA social events are open to all Check-out Items for Dorm Residents. Dorm resi- participants and accompanying people. You do not need dents may borrow pots, pans, dishes, and utensils for a special ticket to attend the following events: July 2nd use in the dorm kitchens from the Registration/Infor- Welcome Social, July 3rd Poster Session Social, July 4th mation table in Green Center Lobby. Please borrow only BBQ and Fireworks, July 5th Poster Session Social, July 6th what you need during your stay and return items by the Russian Party (but see below), July 7th Butterfly Pavilion end of the week. Social, July 8th End-of-Congress Banquet at the Denver Promotion for the next ICA and AAS Meetings. Cor Museum of Nature & Science. Vink, host of the next ICA meeting in 2019, will give a Russian Party. In keeping with ICA and Arachnologi- talk about that meeting at the end of the ICA Business cal Congress traditions, we are sponsoring a Russian Meeting on July 8th in Ricketson Auditorium. The ICA Party. There will be a DJ and a dance floor! We ask that Business meeting will be from 17:00–18:00. The 2017 all participants bring an alcoholic beverage, a non- AAS meeting will be hosted by Fernando Alvarez Padilla alcoholic beverage, or food to share. If you come empty in Merida Yucatan, Mexico. Fernando will promote his handed, then please provide a $5 donation to offset meeting during the first 15 minutes of the AAS Business costs. You are absolutely not allowed to bring any food meeting to be held just before the Russian Party. infused with marijuana. 6 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 7 Cushing 20th International Congress of Arachnology DAILY SCHEDULE Thursday, June 30 IUCN Workshop Group 1: Golden Visitor's Center Conference Room, 1010 Washington Ave. 9:00 Pedro Cardoso Introduction 10:00 Training 11:00 12:00 Lunch (included) 13:00 14:00 13:00–17:00 Training Continued 15:00 16:00 Friday, July 1 IUCN Workshop Group 1 & 2: Golden Visitor's Center Conference Room, 1010 Washington Ave. 9:00 10:00 9:00–12:00 Training - Group 1 11:00 12:00 Lunch (included) 13:00 14:00 13:00–17:00 Redlisting species - Groups 1 & 2 15:00 16:00 Saturday July 2 IUCN Workshop Group 1 & 2: Golden Visitor's Center Conference Room, 1010 Washington Ave. 9:00 10:00 9:00–12:00 Redlisting Continued 11:00 12:00 Lunch (included) 13:00 14:00 13:00–17:00 Redlisting & Summary 15:00 16:00 17:00 Welcome Social, Grand Lobby Green Center, Colorado School of Mines Campus 8 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 9 Cushing 20th International Congress of Arachnology Saturday, July 2 Registration and Welcome Social: Grand Lobby, Green Center, 1500 Illinois Street, Golden, Colorado on the Colorado School of Mines Campus 9:00 10:00 11:00 12:00 Registration and Check-In 13:00 14:00 15:00 16:00 17:00 18:00 17:00–20:00 Welcome Social 19:00 8 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 DENVER MUSEUM OF NATURE & SCIENCE REPORTS | No. 3, July 2, 2016 9
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