Newsletter of the Colorado Native Plant Society dedicated to the appreciation and conservation of the Colorado native flora” . Volume 24 Number 5 November—December 2000 Generic Changes to the Scrophulariaceae affect the Rocky Mountain Flora Dr. Neil Snow, Biological Sciences, University of Northern Colorado A considerable nutnber of generic transfers from the incorrectly imply some kind ofprogress. Scrophulariaceae (figwort family) to the Orobanchaceae (broom- The second principle of phylogeneticsjs that only monophyletic rape family) and Plantaginaceae (plantain family) are now well (mo-no-fy-LEH-tik) groups should be giventaxonomic names. A supported by new research. Herein, I provide an overview ofthe monophyleticgroupisonethatincludesanancestraltaxonandall current philosophy underlying classification and the advent of (not just some) of its descendant taxa. In the past, taxonomic molecular sources of data in systematics, both of which are sig- names have been given to groups thatare notmonophyletic. One nificantly affecting the taxonomy of one of Colorado’s more example is Class Reptilia. Zoologists historically excluded birds 'irominentplantfamilies. fromReptilia,whichinsteadwere placedintheirownClassAves, ^'^nJlassification. The goal ofmodem taxonomy is to group organ- However, sincebirdsarealsodescendedfromthecommonances- ismsinsuchawaythattheclassificationreflectsevolutionaryhis- torofotherreptiles,itis incorrectunderphylogenetic systematics tory. The prevailing taxonomic school of thought among profes- to recognize Reptilia, without also including the birds. Stated sional taxonomists is “Phylogenetic Systematics,” or simply anotherway.ClassReptiliabecomesmonophyleticifthebirdsare — “Phylogenetics.”Thisphilosophy was firstdeveloped indetail by included this is the preferred phylogeneticclassification. the late German entomologist Willi Hennig. Although the initial ... systematics is a science and must reflect embrace of phylogenetics was slow and considerable debate ensued over its relative merits, phylogenetic systematics is now well-supported data firmlyestablishedasthemostscientificapproachtoclassification. Phylogenetic classification has twoover-riding principles. Moleculardata. Soonaftertheriseofphylogeneticsystematics,use The first is that taxa are grouped together only by evolutionarily ofmoleculardataexp^ded rapidly. CertaincharacteristicsofDNA derived characters (traits) that are sharedby twoormore species. found inthenucleus, mitochondria, andchloroplastscanbeusedas Forexample, the presence offlowerscannot be used to helpclas- taxonomicmarkerstoassistintheprocessofclassification,although sifymembersofthegenusCastillejawithinthepaintbrushes.This mitochondrial DNAis usedprimarilyforanimal classifications. isbecauseall angiospermshaveflowers,andflowersthereforeare Moleculardata have enabled some previous taxonomic questions not a derived character among members of Castilleja. However, to be answered satisfactorily. One example bears on relationships the presence offlowers is aderived characterrelativeto all ferns, tothe grassfamily.Anumberofcandidatefamilies had beenpro- whichreproduceonlyby spores. Inthis simpleexample,then,the posed as being mostclosely related tothe Poaceae. But none had presence of flowers could be used to group together species of “Scrophulariaceae”continues on page 2 Castilleja relative to ferns, since flowers are evolutionarily Contents derived relativeto spores. Theword systematists usefora shared derived character is “synapomorphy” (pronounced sin-AP-oh- About the Society .7 morf-ee). In contrast, an ancestral trait is a “sympleisiomorphy” Announcements ..... ....... .2, 3, 4, 6 (sim-PLEASE-ee-oh-morf-ee). These terms should be learned by Articles .1-2, 5 hose interested inthe moretechnical aspects of taxonomy, since Calendar .8 ^ney are used regularly by taxonomists. In contrast, “advanced” (for derived) and “primitive” (for ancestral) are falling out of Chapter News and Announcements ... .2, 6 favor, particularly since “advanced” has a historical tendency to Society News and Announcements .... .3-4 Page 2 Aquilegia Vol. 24 No. 5 “Scrophulariaceae*"continued frompage 1 existing floras recognize these groupings, future publications will been shown convincingly as more closely related than others recognize many ofthe newer concepts. Old habits, however, can using traditional sources of data, such as morphology, anatomy, be hard tobreak. - andchromosomaldata.Amolecularstudypublished several years Anumberofyears ago we tried toconvince my GrandmaAuerin^^ ago demonstrated convincingly that the Joinvilleaceae, which upstate New York to try a peanut butter andjelly sandwich. She consists ofonlyoneortwospecies widely scattered inthe Pacific stated that humans are creatures of habit, and a lifelong habit of region, is the sister family to the species-rich and ecologically hers was to politely decline peanut butter andjelly when offered. dominant grasses. Associating certain generawithparticularfamilies is alsohabitual Changes to the Scrophulariaceae. for botanists. Some may therefore resist the generic reconfigura- Historically, the Scrophulariaceae has tionsoftheScrophulariaceae,Orobanchaceae,andPlantaginaceae. been recognized as consisting of However, systematics is a science and must reflect well-support- some 220 genera, which are particu- ed data. Amore accurate understanding ofphylogenetic relation- larly common in temperate and tropi- shipshelpsus betterunderstandourfloristicheritage. Namesmat- cal mountains. Genera such as ter in some ways, but not in others. After all, the monkeyflower Castilleja, Mimulus, Pedicularis, adorningthemountain stream nearyourfeetduringalunchbreak Veronica, and several others are still looks spilfy, irrespective of its family placement or whether prominent in our area. Penstemon your sandwich contains peanut butterandJelly. even has its center ofdiversity in the mountainous areas of the western United States. However, the WESTERNSLOPEWORKSHOPS Scrophulariaceae was notmonophyletic in its previous classifica- ANNOUNCED tion. It probably comes as a surprise to many CoNPS members GayAustin, PlateauChapter that recentresearch has convincingly indicated the need to trans- fermost generacomprisingthe Scrophulariaceaeelsewhere. For 17 seasons, CoNPS workshops have bfought native plant In 1999, Drs. WalterJudd, ChristopherS, Campbell, ElizabethA. lovers togetherwithawell-informedinstructor. Theopportunity Kellogg, and Peter F. Stevens published Plant Systematics: A to receive one-on-one instruction and informative lectures has PhylogeneticApproach (SinauerAssociates, Inc.), which reveals made the workshop series one ofthe most popular Society pro- thefollowing generic changestothe Scrophulariaceae. This book grams. This season, the Plateau Chapterofthe Colorado Native is highlyrecommendedforthose interested inmore details about PlantSocietyis proudtoannounce several workshopstobecon^ phylogeneticsandtheimportanceofmoleculardata,aswell asthe ducted on the Western Slope. Attendees need only a love of specific reasons for these changes to the Scrophulariaceae, My plants and a desire to learn about them. To registerfor Western thanks to Dr. Richard Olmstead (University of Washington) and Slope workshops or for more information, please contact Gay Dr. Judd (University of Florida) for sharing information on this Austin athome (970) 641-6264orwork(970) 642-4406. recentclassification. How to Know the Families ofthe Western Slope Analysis ofboth molecularand non-moleculardata suggests that Leaden Dr. RobinBingham the following generafound in Coloradoformerly included in the Location: Delta Scrophulariaceae should be placedasfollows; Date: January 27,2001 Scrophulariaceae: VerbascumandScrophularia. Inits newer Dr. Robin Bingham, Botanist at Western State Col|ege, has and more restricted sense, the family now includes only the gearedthisworkshoptowardthebeginnerbotanistorthemember genera containing non-sagittate anthers, in which the anther who would like to review! As such, this workshop will focus on loculesareconfluentand openby a single distal slit. how to recognize the major plantfamilies ontheWesJein Slope. Orobanchaceae: Castilleja, Pedicularis, Agalinis, Dr. Bingham will alsodiscuss basictermsforplantidentification Orthocarpus, and Cordylanthus, The Orobanchaceae in its and how totell monocotsfromdicots. PleasecontactGayAustin newer sense includes the hemiparasitic genera formerly athome(970)641-6264forworkshopregistration, whichis$12. placedin the Scrophulariaceae. Grasses ofthe Western Slope Plantaginaceae: Antirrhinum, Besseya, Callitriche, Leaden Dr. DavidL. Buckner Ghionophila,Collinsia,Digitalis,Hippuris,Limosella,Linaria, Location: DeltaorGrandJunction Lindernia, Mimulus, Penstemon (including Leiostemon), Date: March 17, 2001 Rhinanthus, Veronica (including Pocilla), and Veronicastrum. The Plantaginaceae, thus, now includes the autotrophic Dr. Buckner, Consultant in Plant Ecology, Revegetation, and “scrophs.” The aquatic genera Callitriche and Hippuris are Restoration with ESCOAssociates, will leada basic grass iden- transfersfrom theCallitrichaceaeandHippuridaceae. tification class emphasizing grasses ofthe Western Slope. Tribe characteristics, specifically, will be covered in this workshop, Other changes in classification based on phylogenetic analysis which will begearedtowardbothbeginnersandadvancedmem- eventually will bemadeas morestudiesare published. Forexam- bers. Microscopes and specimens will be available. Please con- ple, somedata supportthemergingoftheAceraceae (maplefam- tact GayAustin at home (970) 641-6264for workshop registra- ily)intotheSapindaceae(soapbenyfamily).Althoughnoneofthe tion, which is $12. - Vol. 24 No. 5 Aquilegia Page 3 NEWS AND ANNOUNCEMENTS SOCIETY Minutes of the CoNPS Board Meetings September 23, 2000 Alice Guthrie, CONPS Secretary New Business: Officers Jill Handwerk (President), Dave Anderson (Vice President), Denise Culver (Treasurer), andAlice September22, 2000 Guthrie (Secretary) were approved by the Board, with the next Iteasnrer’s Report: Denise Culverdistributedthe budget report meeting scheduled for November 4, 2000 in Longmont Public year-to-date, as well as the profitand loss statement. Denise also Safety Building, Conference Room, 225 Kimbark. reviewed chapter subventions, calculated as 25 percent of mem- bershipduesforeachchaptermember. Help! Committee reports: Tllhairs Bob Clarke (Conservation), Sue TheBoardis requestingyourhelpwith thefollowing: Martin (Membership), and Rick Brune (Field Trips) are stepping Donation of a folding table, 5-6 feet long, for Society dis- done, with Velma Richards (Publications) cutting back on her plays. Please contactAlice Guthrie at (303) 651-3127. responsibilities. Eric Lane will take over as Chair of Volunteeropportunities: , Membership, while JeffDawson will chairPublications, work- ProGreenShow inDenver,January2-5, 2001.Volunteers ingwithVelma.TheChaifofPublicity is still vacantandthe sta- areneededtosetupandstafftheboothforeitherahalfday tus of the Field Studies Committee is under discussion. Leo orwhole day. ContactJill Handwerkat (970) 491-5857. BruederleoftheNewsletterCommitteereportedthattherewillbe Committee Chairs (andmepibers) are neededforthefol- five regular issues ofAquilegia in 2000, with the possibility ofa lowingcommittees: Conservation,FieldTrips,Legislative, special issuecomprisinganarticleentitled“SoilLichensandtheir Affairs,andPublicity. ContactJill Handwerkat(970)491 Application as Ecological Indicators in Sagebrush Habitats” con- 5857 ofAlice Guthrie at (303) 651-3127 for information tributed by RogerRosentreter. and to volunteer. New Business: The Rare Hant Book Fund was renamed the Bookreview:AquilegiaissolicitingareviewofTheLandscaping MyrnaP. SteinkampMemorialFundandwill be used similarly Revolution: Garden withMotherNature, NotAgainstHer by totheMarrFund.Interestgeneratedfromtheprinciplewill beded- Andyand SallyWasowski. Receiveafreecopyinreturnfora icatedtotheprotectionofColoradorareplants,as well asresearch 500-1000wordreview.ContactLeoP. Bruederleat(303)861- ^'y^dvancingourknowledgeoftheseplants.TheBoardvotedtosplit [email protected]. Communications into twocommittees, one overseeingAquilegia Informationneeded:Wewouldliketocompilealistoffairs publication and a second overseeing the CoNPS website, feo and shows atwhichtoeducatethe publicaboutCoNPS. If Bruederle will serve as Chairofthe NewsletterCommittee, while you know of local or county shows that would be appro- BobClarke will chairtheWeb Site Committee primarily oversee- priate, please call Alice Guthrie (303-651-3127) with the ingContent. SaraHill, whow^ commendedforherworkas web- name,contactinformation,and general datesofthevenuel master, will remain in that position. Jill Handwerk proposed a cooperative effort with Colorado State University to bring Dr. AQUILEGIA Deadline Approaches Gary Nabhan(Director,TheCenterforSustainableEnvironments, Submit on or before December 2pO0 1, University ofNorthernArizona) toFortCollins inordertodeliver a CSU Provost Lecture on EnvironmentalAchievement. The Please submit contributions forVol. 25 No. 1 ofAquilegia on or Board approved allocating $5(X) toward an honorarium for Dr. beforeDecember 1,2(KX). Includeauthor’snameandaddress,and Nabhan, who would also conduct a Society workshop, date and follow theformatfrom previous issues ofAquilegia as closely as timeto beannounced. Inadiscussionofthedefinition oftheterm possible. Previously published articles submitted for reprinting tnhaetifvoel,laoswiinngnaftriovmetphleanOtx,ftohredBoDiacitdioangarreye.d“tAoptpelmipeodratroilaysapcecceipets rseuqbumiirtetepdervmiiassei-omnaiflroormothnedeidsiktsor—ofMqrAigCinaplrepfuebrlaibclayt,ioonr.AIrBtiMcl—es thatoccurs naturally in an areaad therefore one that has not been are much appreciated. Please submit the contribution as a MS introduced by humans either accidentally or intentionally. Of WordorRTF(rich textformat) document. plants found in a particular place, the term is applied to those species that occur naturally in the region and at the site.” Gay The Importance of Native and Invasive Austin will revise this definition based on the ensuing discussion. Species in Land Management PeggyLyonpresentedFundingGuidelinestOTAdHocRequests, Society of Range ManagementAnnual Meeting which was approved. The Board subsequently approved a request for$500fromCityofBoulderOpenSpaceforrevisingandreprint- November 30 - December 1, 2000 ing the Russian olive brochure, on which the Society will be The Society for Range Management, USDA Forest Service, le'^knowledged. LisaTaskerannouncedtheEcologicalRestoration BLM,ChaffeeCounty,ColoradoSeedGrowersAssociation,and >^^Conference being sponsored by the Central Rockies Chapter of PawneeButtesSeedareputtingonaworkshopinFortCollinson the Society for Ecological Restoration, for which Lisa hopes the invasive speciesandthe use ofnativeplants infevegetation. For Board will considerlendingsupport. more information, pleasecontactDon Hijarat (970) 356-7002. Page 4 Aquilegia Vol. 24 No. 5 CeRSER NEWS Ecological Restoration Conference SOCIETY April 26-27, 2001 The Central Rockies Chapter of the Society for Ecological Request For Proposals Restoration announces a very exciting regional conference: ^‘Restoring the Rockies: Restoration and Conservation The John W. Marr Fund StrategiesintheWest,”tobeheldonThursdayandFriday,April OnMarch24, 1990,the BoardofDirectorsofthe Colorado Native 26th and 27th, 2001 in Keystone, Colorado. With multiple orga- Plant Society established the John W. nizationsthroughouttheRockiesnowpracticingrestorationwith- Marr Fundfor the award of small grants inadiversityofecosystems,ameetingtoexchangestrategiesand in support of research in plant biology. bring together a wide variety of individuals, groups, and coali- The Fund honors the late Dr. John Marr, tions shouldbeofser\dcetoall. Inadditiontoexchangeofideas, Professor at the University of Colorado the hope is that the conference will also help organizations form and the first President of the Colorado alliances so as to better ensure the long-term survival of our Native Plant Society. The Fund was diversehistoricalecosystemswithintheRockyMountainRegion. establishedtoencourage,throughthepro- Keystone Resortis only 75 miles from Denverandmoreconve- vision ofsmall grants-in-aid, research on niently locatedthantheFrontRangeforfolkscomingfrom west the biology and natural history of ofthecontinental divide. Theresortclaimsthelargest,most ver- Colorado native plants. Both field and satile meeting facilities in Colorado to say nothing of the sur- laboratorystudiesareeligibleforfunding. roundings and recreational activities. Rates should be very rea- Awards of $500 or less are typical and sonable,aslateApril islowseasonforKeystone. Don’tmissthis may represent the sole support of a pro- exciting opportunity to find out more about the practice ofeco- ject, may be part ofjoint funding agreed logical restoration in yourlocal ecosystems andhow itrelatesto upoiiwithothergrantingorganizations,or the biggergoal ofconserving ournative plantcommunities. maybeusedtosupplementotherfunding Submitting a PaperorPoster. obtmned bytheinvestigator(s). Acall for papers is in progress.Abstracts and posters pertaining Aquilegiachrysantha toall areasofrestorationarecurrently beingaccepted. Emphasis Artist:JanWingate will be placed on riparian and wetland areas, rangeland restora- P. Steinkamp Memorial Fund tion, restoration on private lands, restoration education (eg., On September22,2000, theBoardofDirectorsrenamed the Rare schoolyard habitats), restoration in rights-of-ways, and urban Plant Book Fund to honorDr. Myma P. Steinkamp. Myma was a restoration. Proposals are welcome from all disciplines, back- founding member of the Society, and serv^ many years both as grounds, and experience levels. Presentations should be Treasurer and Chair of the Membership Committee. She also designed to be 20 minutes long, with an additional five minutes served on the Conservation Committee, helping to establish the for question and answers. Longer time slots (up to 30 minutes) Society’s role in Colorado plant conservation and public lands may be available if space permits. Please make the necessary planning. She participated actively in the preparation of both edi- arrangements if your presentation does not fit the allotted 20 tions of the Society’s publication Rare Plants of Colorado. minutes. Duetotheanticipated diversity oftheaudience, papers Although this fund had been specifically designated for publica- shouldincludeabriefbackground. Forposterpresentations,pre- tionofRarePlantsofColorado,thescopeofthefundhasnowhas senters must provide their own free-standing bulletin boards. been broadened to include support for research on rare Colorado Abstracts and proposals should begin with the name and orga- native plants, through the provision of small grants-in-aid. Both nizational affiliation ofeach author with the presenting author’s field and laboratory studies are eligible for funding. Awards of name underlined. Also include the address, phone and e-mail (if $500orlessare expectedand may representthe sole support ofa available)ofthecorrespondingauthor,followedbyadescriptive project,maybepartofjointfundingagreeduponwithothergrant- title and a text summary notexceeding 300 words. Submission ing organizations, or may be used to supplement other funding deadline is December 1, 2000, although proposals may be obtained by the investigator(s) for research in Colorado. accepted beyond thatdate on aspace available basis. Electronic Contributionsto the memorial fund may bemade to: submissions as MS Word documents or RichText Files (rtf) are preferred and should be submitted to Jody Nelson at MymaR SteinkampMemorial Fund [email protected]. Ifelectronic submissionis notpossible, c/o Colorado Native Plant Society hard copyordisk should be mailed to: P.O.Box200 CentralRockiesChapteroftheSocietyforEcologicalRestoration FortCollins, CO80522 P.O. Box 17644 The Board of Directors is now soliciting proposals for both the Boulder, Colorado 803040644 Marr Fund and Steinkamp Memorial fund, for which the dead- Forthe most up to date information, please send e-mail to lisa- linesareJanuary 15,2001. Informationonguidelinesand require- [email protected]. Find alinktothe Central Rockies Chapter ments for proposals may be obtmned fiy contacting Board mem- through the international SER website www.ser.org. berEric Laneat(303) 239-4182. Vol. 24 No. 5 Aquilegia Page 5 MARR FUND: Reports from Recipients holistic Survey of South Central Colorado Status of the Green Sedge in Colorado Brian Elliott Shannon Kuchel During 1998 and 1999, an intense floristic survey was performed Carex viridula Michx. (Cyperaceae), the in south central Colorado. This study, supported in part by the green sedg^, is a short-lived, graminoid JohnMarrFundoftheColoradoNative PlantSociety, isthefocus perennial with adensely caespitose habit. ofmyMaster'sThesisattheUniversityofWyoming. Fieldworkis It is monoecious, with a single terminal now complete and has resulted in over 20,000 numbered collec- staminate spike and several sessile pistil- A tions, with detailed locationandecological dataforeach. setof late spikes. It typically occupies moist, specimens will be retained at the Rocky Mountain Herbarium, early successional sites characterized by with duplicates goingtootherpublic herbaria. fluctuatingandunpredictablewaterlevels. The study area includes the Wet Mountains, Sangre de Cristo These sites vaiy from calcareous, acidic, Range, Culebra Range, Spanish Peaks, and surrounding areas. sandy, or organic shorelines; runnels in Ranging from the edge of Colorado's eastern plains up to an limestone barriers; wet meadows; marsh- extensive alpine zone, the regionencompasses over5,000 square es; on borders of streams, ponds, and miles, with a rich diversity of vegetation types. The area has lakes;tofens. Carexviridulaisdistributed received relativelylittleattentionfrombotanists previouslyand is throughout northern and central North hardly mentioned in Weber and Wittman's Colorado Flora: America. It also extends to the southern EasternSlope. Rocky Mountain region in several dis- junct sites, including alpine wetlands in Thesurveydocumentedapproximately 1150taxa.Thisincludes44 Colorado, where it is rare. Although C. taxafrom the ColoradoNatural Heritage Program’s listofspecies viridula has been assigned a global rank- of spiecial concern. Notable collections include dwarf milkweed ing ofG5, indicating that itis demonstra- {Asclepias uncialis), Colorado larkspur {Delphinium alpestre), bly secure globally, it has received astate Artist;JanWingate Gray's Peak whitiowgrass {Draha grayana). Smith whitlowgrass rankingof SI, indicating that itis critically imperiled inColorado. .smithii),show}'prairieGentian {Eustomagrandiflorum),spiny The purpose of this research was to describe genetic diversity in ^ster {Herrickia horrida)., 12 populations of rock-loving populations of C. viridula from Colorado, and determine if these Neopanya {Neoparrya lithophila),alpine poppy {Papaverradica - populations exhibited reduced genetic diversity when compared to turn ssp. kluanensis), pictureieaf wintergreen (Pyrola picta), and otherpopulations ofthis speciesfrom NorthAmericaandEurope. Altai stitchwort {Stellaria irrigua). Also collected were In ordertoquantify geneticdiversity, small samples ofleaftissue Cymopterus constancei, were collected from 350 individuals representing seven extant newly described by Dr. Ron populations ofC. viridula in Coloradoand 179individuals repre- Hartman inBrittonia., andthe senting eight populationsfrom elsewhere throughout the rangeof soon to be published this species in NorthAmerica. Starch gel electrophoresis ofsolu- Oonopsispuebloensis. ble enzymatic proteins coupled with substrate specific staining wasusedtodeterminetheamountanddistributionofgeneticvari- This study is only part of a ation in these populations. These data were compared with data larger Flora of the Rocky previously reportedforEuropean populations ofthis species. Mountains Project being conducted by Dr. Hartman, Usingthesetechniques,nogeneticvariation,eitherwithinoramong Curator of the Rocky populations from Colorado or elsewhere in NorthAmerican, was Mountain Herbarium at the detected. However, NorthAmerican populations were genetically University of Wyoming differentiated from European populations, with significantly more (http://www.rmh.uwyo.edu). diversity maintained by the latter. The surprising lack of genetic Many thanks are due to the diversity in NorthAmerican populations is probably thecombined Colorado Native Plant resultofhigh levelsofselling and inbreeding, restrictedecological Society (andespecially tothe amplitude, and genetic drift. Genetic bottlenecks are presumed to supporters of the John Marr have occurred as a result of climate changes associated with Fund), as well as to Nancy Pleistocene glaciationorfoundingevents associated with coloniza- Ryke of the San Isabel tionofNorthAmericabyproposedancestral Europeanpopulations. tionalForestforgenerous- I would like to thank Dr. Leo R Bruederle for his guidance ^Vsupportingthis research. throughout this project. I am also grateful for support from the Eustomagrandiflorum Colorado Native Plant Society, Sigma Xi, and the Department of Artist:JanA^ngate Biology atthe University ofColoradoatDenver. Page 6 Aquilegia Vol. 24 No. 5 CHAPTER NEWS Boulder Chapter Storage Laboratory (NSSL) onthecampus is primarily montane, with both lowerand ofCStJ. The NSSL is locatedjust west of upper montane forest well-represented. Monthly meetings are held October the railroad tracks, about one block north Leo P. Brnederle (Associate Professor, throughAprilattheFoothillsNatureCenter, of Pitkin Street (enter on the west side of CU-Denver) will discuss ongoingresearch locatedinBoulderat4201 NorthBroadway, the building). For more information, con- conducted along with CoNPS members opposite its intersection with Sumac Ave. tactChapterPresidentDonHazleltat(970) SaraE. HillandKevinFehlberg,theobjec- The Chapter now meets on the second 834-1493. tiveofwhichwastocircumscribetheflora Wednesday of each month, at 7:00 pm for ofthe RampartRange. business,aspeaker,booksales,andsocializ- November 9, 7:00pm ingWithrefreshments. Pleasejoinusat5:00 Weed Warrioring and other Botanical January 30, 7:30PM PM for supper with our speaker at Efrain’s ExploitsatDinosaurNationalMonument New Species in the Myrtle Family from Mexican Restaurant, 1630 N. 63rd St, just Tamara Naumann (Botanist, Dinosaur the Rich Flora ofQueensland,Australia north of Arapahoe in Boulder. For more National Monument) will repeat her Queensland is mega-rich in its biodiversi- information, contact Chapter President November8 BoulderChapterProgram. ty,asevidencedbyits9,000speciesofvas- Caiyl Shieldsat(303)665-6312orProgram cular plants. Many new species are ChairCarolynCrawford(303)665-6903. December5, 7:00pm described annually, particularly from the Ve^tableMotor-oilforColoradoVehicles more tropical areas of the state. Recent November8j7:00PM Duane Johnson (Professor ofAgriculture, research has focused on describing new Weed Warrioring and other Botanical CSU) will discuss his extensive work with species and taxonomicallyrevising several ExploitsatDinosaurNationalMonument leSserknownColoradocropsandplantprod- genera of fleshy-fruited members of the Tamara Naumann (Botanist, Dinosaur ucts. He will also review informationon the Myrtlefamily(Myrtaceae). Dr, NeilSnow National Monument) will present some of toxicityofBTcompollentonativeinsects. (Assistant Professor, UNC) will provide a htheerMexopneurmieenncte,sionvcelrudtihneglatshtosseevdeeraalliynegarwsitaht FTeobbreuaarnyno6u,n7c:e00dpm bwritohada poavretrivcuileawrfoofcutshoenAQuuseteranlsilaannd.flora, weeds,rareplants, tourists,and ranchers. December 13,7:00pm Metro-Denver Chapter Plateau Chapter The World of Wildland Fires, Indoor monthly meetings are held at7:30 Workshops, field trips, and other activities PrescribedBorns,andFireEcology PM from September through April in the are scheduled throughout the year. For Coloradojustexperienced one 6fits worst Morrison Center at the Denver Botanic more information, please contact Chapter years for wildland (forest, brush, and Garden. For more information, contact PresidentJeanneWengerat(970)256-9227 range) fires. Justin Dombrowski (Fire PresidentDeniseLarsonat(303)733-4338 or by e-mail at [email protected] or Management Officer, Boulder Fire or Leo P. Bruederle at (303) 861-4119 or ProgramChairLori Brummerat(970)641- Department) is involved in many facets of bye-mail at [email protected]. 3561 [email protected], wildland fire management throughout the fireseasonandacrosstheState. Hewilldis- October31, 7:30 pm November 18, 10:30am cuss his experiencesduringthe pastyear. StatusoftheGreenSedgein Colorado Plateau Chapter Annual Planning The green sedge, Carex viridula Meeting and Potluck January 10,7:00pm (Cyperaceae), occurs in wetland habitats Please note this change in date! TheRocky MountainArsenal scattered throughout northern and central Following our organizational meeting The RockyMountainArsenal wasformer- NorthAmericaandEurasia. Itsdistribution (workshops, field trips, and elections) and ly involved in weapons work during the alsoextends to several disjunct sites in the lunch, we’ll have a seed and catalog Cold War. Since being decommissioned, alpine wetlands of Colorado, where it is exchange. Members will then have an much of the area has been restored as a critically imperiled. Shannon Kuchel, opportunity to show off some ofthe great wildlife refuge. Suzanne O’Neill 1999 recipient of a John W. Maix Fund slides they took over the year. Dust your (President and Director, Rocky Mountain research grant, will discuss her research cameras offand buy some slide film! This Arsenal Wildlife Society) will discuss the addressing genetic diversity maintainedby should be fun! Meet at the Forest Service changes that have occurred over the years Colorado populations ofthe green sedge. building in Delta. andthefutureofthe site. December5,7:30 pm Southwest Chapter Fort Collins Chapter Floraofthe RampartRange. The Rampart Range is one ofa numberof For Chapter news and activities, pleasf Indoor monthly meetings are scheduled distinct geologic features comprising the contact Chapter President Sandy Friedle^L:::;?^ through April at 7:00 PM in the Main Front Range of the southern Rockies in at (970) 884-9245 or by e-mail at fried- Conference Room of the National Seed Colorado. The vegetation ofthe Ramparts [email protected]. . Vol. 24 No. 5 Aquilegia Page 7 Colorado Native BoardofDirectors Plant Society Jill Handwerk(01) . . FortCollins. . 970-491-5857 BillJennings(01). . .Louisville . ..303-666-8348 AndyKratz(01)... .Lakewood. . .303-914-8291 PatPloegsma(01). . . Strasburg. . . . 303-622-9439 Aquilegia is published four to six times per Ericyne(01) Lakewood. . . 303-239-4182 year by the Colorado Native Plant Society. DavidAnderson(02) FortCollins. . 970-484-0774 This newsletteris available to members ofthe GayAustin(02) .... Gunnison. . . . 970-641-6264 Society and toothers with aninterestin native KathyDarrpw(02). . CrestedButte. 970-349-0743 The Colorado Native Plant Society is a non- plants. Contact the Society for subscription GwenKittel(02).. . .Boulder 303-258-0908 profit organization dedicated to the apprecia- information. Articles for Aquilegia may be Laurel Potts(02). ...FortCollins. . 970-484-4076 tion and conservation of the Colorado native used by other native plant societies or non- Neil Snow(02) Greeley 970-330-4823 flora. Membershipis open toall with an inter- profitgroups iffullycitedtoauthorandattrib- ChapterPresidents est in our native plants, and is composed of utedtoAquilegia. Boulder.......Caryl Shields ....303-665-6312 plant enthusiasts both professional and non- professional. Pleasedirectallcontributionstothenewsletterto; FortCollins....DonHazlett ..... 970-834-1493 LeoP.Bruederie Metro-Denver . . DeniseLarson . . .303-733-4338 Pleasejoinus in helping to encourage interest Biology,Campus Box 171 Plateau JeanneWenger. . .970-256-9227 in enjoying and protecting Colorado’s native University ofColoradoatDenver Southwest SandyFriedley. .1970-884-9245 plants. The Society sponsors field trips, work- P.O.Box 173364 shops, ahd otheractivities throughlocal chap- Denver,CO 80217-3364 Committees Conservation...VACANT tersandstatewide. ContacttheSociety,achap- E-Mail: [email protected] terrepresentative,orcornmitteechairformore Education .....TinaJones 303-794-2647 information. Shortitems suchas unusual informationabout andCo-chairVACANT aplant,alittle known botanical term,etc.,are FieldStudies .. . LoraineYeatts . . .303-279-3427 ScheduleofMembWshipFees especially welcome. Previouslypublishedarti- FieldTrips.....VACANT ySufpeporting : . . . . ,. . . .. ..$.2$5500 cslioens.sCuabmmeitrtae-drefaodryrelpirneintaritngorreoqtuhierer iplelrumstirsa-- HoRretsitcourlattuiroenan.d...KMadthLyisDaaTmaasske.r.....390730--554443--31643932 Organization . . . . .$30 tions are also solicited. Pleaseinclude author's Legislative FamilyorDual .$15 nameandaddress,althoughanonymitymaybe Affairs ..VACANT . . ndividual .$12 requested. Articles submitted via e-mail or on Membership. . . .EricLane..... . 303-239-4182 IStudentorSenior .$8 disks(MACpreferably,orIBM)areappreciat- andSueMartin. . . 970-226-3371 ed. Please indicate word processing software Newsletter LeoP.Bruederie. .303-556-3419 MembershipRene^al/Information andversion;ifpossible,submitasanRTF(rich Publications....JeffDawson 303-722-6758 Please direct all membership applications, textformat) file. andVelmaRichards303-794-5432 renewals, and address changes to the Publicity VACANT Membership Chairperson, Colorado Native Officers RarePlant ... Eleanor Plant Society, P.O. Box 200, Fort Collins, CO President JillHandwerk . .970-491-5857 Monograp.h. . . . VonBargen ....303-756-1400 80522.Heasedirectallotherinquiriesreg^ding Vice-President. . .DavidAnderson. 970-484-0774 WebSite BobQarke 970-242-6067 theSocietytotheSecretaryatthesameaddress. Secretary AliceGuthrie. . .303-651-3127 Workshops BillJennings ....303-666-8348 Treasurer DeniseCulver . .970-225-1930 MEMBERSHIP APPLICATION FORM Name(s) MEMBERSHIPCLASS (Selectone) Address ' Individual, $12.00 ^ Family/dual, $15.00 (Address) Senior/retired, $8.00 Student, $8.00 City - State Zip Corporate, $30.00 Supporting, $50.00 Phone Li 1 E-mail . Chapter; Boulder ____ Fort Collins Metro Denver Plateau Southwest Inadditionto rny membership, I have included $ as a contribution to theJohn Man- Fund (endowment in supportofsmall grants-in-aid ofresearch), $ as acontribution to ^.le MymaP. Steinkamp Memprial Fund (endowmentin support ofsmaH grants-in-aid of research), or $ as a general contributionto the Society. CONPSISA NON-PROFITORGANIZATION -DUESAND CONTRIBUTIONSARE TAX-DEDUCTIBLE CHAPTER MEETINGS AND EVENTS Plateau Chapter (continued) Boulder Chapter January 27 Worikshop: How to know the Families ofthe Western Slope November8 WeedWarrioringandotherBotanical ExploitsatDinosaurNat’l. Monument March 17 Workshop: Grasses ofthe Western Slope December 13 TheWorld ofWildland Fires, Prescribed Bums, and Fire Ecology SOCIETY EVENTS January 10 The Rocky MountainArsenal Board Meetings Fort Collins Chapter November4 Longmont, Colorado November9 WeedWarrioringandotherBotanical Workshops ExploitsatDinosaurNat’l. Monument November4, 5 How to Know the Families December5 VegetableMotor-oilforColoradoVehicles First session: Saturday Second session: Sunday Metro-Denver Chapter December9, 10 Small Plant Families October31 Statusofthe Green Sedge in Colorado First session: Saturday December5 FloristicAnalysisofthe Rampart Range Second session: Sunday January 30 New Species in the MyrtleFamily from January 13, 14 Grasses: TheTribe Hordeae the Rich FloraofQueensland,Australia First session: Saturday Second session: Sunday Plateau Chapter November 18 Plateau ChapterAnnual Planning Febraary 10, 11 Cirsium: The Native Thistles Firstsession: Saturday Meeting and Potluck Second session: Sunday ColoradoNativePlantSociety P.O. Box 200 FortCollins,Colorado80522 http://www.conps.org TIME SENSITIVE MATERIAL