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Aquatic and Wetland Plants of Northeastern North America: Angiosperms: Monocotyledons (Volume 2) PDF

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Aquatic and Wetland Plants of Northeastern North America 2 Angiosperms: Monocotyledons Garrett E. Crow and C. Barre Hellquist Aquatic and Wetland Plants of Northeastern North AIllerica A revised and enlarged Volume 2 edition of Norman C. Fassett's Angiosperms: Monocotyledons A Manual of Aquatic Plants The University of Wisconsin Press Figures 60. 63.75.112. 118. 136. 147. 193.216.259.261. 297. The University of Wisconsin Press 316.317.318.319.324.330.419.421.585. and 586 are reprinted 2537 Daniels Street in part or in whole. with permission. from The New Britton and Brown Madison. Wisconsin 53718 Illustrated Flora oj the Northeastern United States and Adjacent Canada. by Henry A. Gleason. Copyright © 1952 by The New York Botanical 3 Henrietta Street Garden. London WC2E 8LU. England Figures 32. 34. 35. 151. 153.168. 269.414.415.416.418.421. Copyright © 2000 422.423.429.431.433.437.443.444.445.493.506.514.518. The Board of Regents 519.525.529.600. and 601 are reprinted in part or in whole from of the University of Wisconsin System Vascular Plants oj the Pacific Northwest. by C. L. Hitchcock. A. Cronquist. All rights reserved M. Ownbey. and J. W. Thompson. by permission of the University of Washington Press. Copyright © 1955-1969 by the University of 543 2 Washington Press. Printed in the Figures 139.149.151.193. and 196 are reprinted in part or in United States of America whole from The Woody Plants oJ Ohio: Trees. Shrubs. and Woody Climbers. Native. Naturalized. and Escaped (or. are reprinted from The Monocotyledoneae: Cat-tails to Orchids. The Vascular Flora oj Ohio. Vol. 1) Library of Congress Cataloging-in-Publication Data by E. Lucy Braun. by permission of the Ohio State University Press. Copyright © 1961 by the Ohio State University Press. Crow. Garrett E. Aquatic and wetland plants of northeastern North America: a Figures 354. 451. 519. 530. 586. and 591. are reprinted in part or in revised and enlarged edition of Norman C. Fassett's A manual of whole from The Monocotyledoneae: Cat-tails to Orchids. The Vascular Flora aquatic plants / Garrett E. Crow and C. Barre Hellquist. oJ Ohio. Vol. 1) by E. Lucy Braun. by permission of the Ohio State 456 pp. cm. University Press. Copyright © 1967 by the Ohio State University Press. Includes bibliographical references and indexes. Contents: v. 1. Pteridophytes. gymnosperms. and angiosperms: Figures 28.29.31. 74. 96. 144. 152. 223. 231. 234. 242. 252. 254. dicots-v. 2. Angiosperms: monocots. 262.264.266.273.322.328.343.368.381.389.391.393.395. ISBN 0-299-16330-X (v. 1: cloth: alk. paper). 435.452.461.463.466.467.489.490.507.508.509.511.518. ISBN 0-299-16280-X (v. 2: cloth: alk. paper) 522.523.524.528.576.577.578.580. and 603 are reprinted in 1. Aquatic plants-Northeastern States Identification. part or in whole from A Flora oj the Marshes oj CaliJornia by Herbert L. 2. Wetland plants-Northeastern States Identification. 3. Aquatic Mason. by permission of the University of California Press. Copyright plants-Atlantic Provinces Identification. 4. Wetland plants © 1957 Regents University of California; 1985 Herbert Mason. Atlantic Provinces Identification. I. Hellquist. C. B. II. Fassett. Norman C. (Norman Carter). 1900-1954. Manual of aquatic Figures 8.19.20.21.22.23.24.25.26.27.29.42.44.48.49. SO. plants. III. Title. 53.54.55.57.58.59.60.61.62.63.64.65.74.81.83.84.85. QK1l7.C84 2000 87.89.90.92.93.94.140.259.263.479.547.595. and 604 are 581.7'6'0974-dc21 99-19556 reprinted from bulletins published by the New York State Museum. Figures 56. 68. 75. 77. 80. 83.97.98. 103. 104. 108. 124. 128. 129.165.166.170.171.174.175.176.177.178.179.180.181. 183.184.198.201.204.213.224.225.238.245.246.247.248. 249.250.253.255.270.274.275.276.278.280.281.283.286. 287.288.289.338.342.360.362.382.388.396.403.404.407. 408.409.410.412.415.416.417.420.422.42~426.42~428. 430.433.441.443.446.449.453.454.455.456.458.464.470. 471.472.473.474.475.476.477.478.479.480.481.482.488. 491.492.494.495.496.497.498.499.500.501.502.503.505. 508.509.510.511.512.516.517.520.521.525.527.532.533. 534. 535. 536. 552. 553. 554. 561. 568. 585. 590. 592. 594. 595. 597. 598. and 60S are reprinted in part or in whole from Aquatic and Wetland Plants oj the Southeastern United States by Robert K. Godfrey and Jean W. Wooten. by permission of the University of Georgia Press. Copyright © 1979. 1981 by the University of Georgia Press. Contents Preface vii Subclass Commelinidae 85 Acknowledgments ix Xyridaceae 85 Introduction xi Eriocaulaceae 92 Map of geographical area xiii Juncaceae 97 Nuisance Aquatic Plants of the Northeast xv Cyperaceae I 2 I General Keys xix Poaceae (Gramineae) 252 Sparganiaceae 295 Monocotyledons 3 Typhaceae 306 Subclass Alismatidae 3 Subclass Zingiberidae 3 I 0 Butomaceae 3 Marantaceae 3 I 0 Alismataceae 3 Subclass Liliidae 3 I 2 Hydrocharitaceae I I Pontederiaceae 3 I 2 Scheuchzeriaceae 33 Haemodoraceae 3 I 7 Ju ncaginaceae 33 Liliaceae 320 Potamogetonaceae 36 lridaceae 323 Ruppiaceae 58 Burmanniaceae 328 Zannichelliaceae 58 Orchidaceae 328 Zosteraceae 68 Abbreviations 351 Najadaceae 68 Glossary of Plant Terms 353 Subclass Arecidae 76 Glossary of Habitat Terms 363 Acoraceae 76 References 365 Araceae 76 Index 373 Lemnaceae 78 v Preface The first edition of Fassett's Manual of Aquatic Plants ap Newfoundland. The coverage of Virginia in the first edi peared in 1940 and quickly became a classic in its field. tion was necessarily incomplete, and that gap has been Its success was, in large part, due to Dr. Fassett's aim to closed in this treatment. Fassett's treatment of 752 taxa provide a manual which would make possible the identi (plus 95 named forms) has been greatly expanded in this fication of an aquatic plant in sterile as well as reproduc edition to include 1186 taxa. tive conditions. He made an all-out effort to construct his Dr. Fassett's decision to exclude bog species in the first keys as simply as possible. coupled with flowering or fruit edition left many aquatic biologists and students in ing characters where essential for proper identification. northern regions without a comparable guide to the iden He copiously cross-referenced characters in his keys to la tification of species occupying those wet, boggy habitats. beled illustrations to facilitate the identification process Thus, we have included those species which typically further. grow in saturated peat. Likewise, we have attempted to Shortly before his death in 1954, Dr. Fassett invited Dr. meet the needs of aquatic biologists along the Atlantic Eugene Ogden to prepare a revised edition. This effort, coast by including species of salt marshes and tidal wa which appeared in 1957, resulted in a 22-page appendix ters, a study of which began Dr. Fassett's career as a be aimed at bringing the nomenclature of the original text ginning student in Professor Fernald's Botany 7 class at up to date. It also provided a much improved supplemen Harvard University and later became the focal point of tary key to the genus Potamogeton. Both Drs. Fassett and his doctoral dissertation. Ogden readily acknowledged that the addition of an ap In the original edition Dr. Fassett also included a few of pendix, though extremely useful, fell far short of the need the more common examples of various groups of algae for a complete revision. and some common mosses and liverworts. There are ex This new edition provides the badly needed updating, cellent references now available for the identification of yet attempts to retain the features which made Fassett's these groups of plants. We feel that to give adequate at work a classic. The region covered by the first edition, tention even to just the aquatic bryophytes would be be Minnesota to Missouri, eastward to the Gulf of st. Law yond the scope of our work. rence and Virginia, has been extended slightly to include vii Permission graciously provided by the New York State Museum is acknowledged for use of illustrations pub- lished in several of their bulletins (NYSMuseum); Rich- ard S. Mitchell and E.O.Beal (1979), Magnoliaceae through Ceratophyllaceae of New York State; Richard S. Mitchell and J. Kenneth Dean (1978), Polygonaceae (Buckwheat Family) of New York State;Richard S. Mitchell and J. Ken- neth Dean (1982), Ranunculaceae (Crowfoot Family) of New York State;Richard S. Mitchell and Charles J. Sheviak (1982), Rare Plants of New York State; Eugene C. Ogden (1981), Field Guide to Northeastern Ferns; and Charles J. Sheviak (1982), Biosystematic Study of the Spiranthes cernua Complex.

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This comprehensive and up-to-date reference work is an identification manual covering more than 1100 species of vascular plants that are native or naturalized in aquatic and wetland habitats in the northeastern region of North America, from Newfoundland west to Minnesota and south to Virgina and Mis
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