[Ants of the genus Tetramorium (Hymenoptera, Formicidae) in the fauna of the USSR. Part I.]

11 Pages·1992·0.41 MB·English

by  Radchenko, A. G.

. . .
300JI0rHMECKHft ^YPHAJI
1992, TOM 71, Bbin. 8
^AVCHbiNKO
yjlK 595.796 (47+57)
©
1992r. A.r.PAflHEHKO
MYPABLH POflA TETRAMORIUM (HYMENOPTERA, FORMICIDAE)
OAYHbl CCCP. COOBIUEHME
1
BcTSTbeflanoSsoppofl3 TetramoriumMayr (JjayHu CCCP. OnHcaHu flBa hobwx ana HaygHbh-
Ha: T. vernicosus (othochtch krpyime T. inerme) h T. bursakovi (othochtch k rpynne T.ferox).
T. kabulistanicumPisar.Bnepsbie npHBOAHTca juih (Jjaynbi CCCP. flaHa onpeneJiHTentHaa laSnHua
BHHOBpoAa Tetramorium{JjayHuCCCP (nopa6o4HMh caMKaM)
Tetramorium Mayr - ouym H3 KpynHeftumx no micny bhhob KOCMonojiHTHMecKHx
poflOB MypaBbCB. CoBpcMeHHbie peBH3HH poaa npOBeneHbi anH BcexperaoHOB, KpoMe
najieapKTHKH (Bolton, 1977, 1979, 1980)'. Bojitoh (Bolton, 1980) npHBOflHT ana
MHpoBOH (j^ayHbi Tetramorium iw^jpy 328 bhaob, b tom mhcjic b llaneapKTHKe - 25.
rio HauiCMy MHeHHK), uh4jph 3™ secbMa 6jih3Kh k peanBHbiM (b llaneapKTHKe hhcjto
BHrtoB,BepoHTHO, HeMHOiUM Qoiiee 30)
IlepBOH peBHSHeii,BKHMMaiomeH najieapKiHiecKHx Tetramorium, 6biJia pa60Ta 3Me-
pH (Emery, 1909), rae aarop npHBeJi nHTbBHnoB (HCKjuoHan HHTpoHytuipoBanHbie) m
OKOjio 20 noflBHnoB H BapHexcTOB, "HcTb H3 KOTOpbix — HacTOHiUHe BH^bi, a npyrae
nonacHbi paccMETpHBaTbcJ! KaK cHHOHHMbi. no3AHee B Karanore MypaBbeB MMpoBoii
4)ayHw (Emery; 1921) ajih IlajieapKTHKH yKasanw MeTwpe BHaa c 38 noflBHaaMH h
BapHeieTaMH. B AajibHefimeM pasAHHHwe aBTOpw (Santschi, 1921, 1927; Emery,
1925; Karawajew, 1926, 1926a, 1926c, 1931; Wheeler, 1923, 1927; Stitz, 1934;
Roszler, 1935, 1936; Kratochvil, 1944 et al.) onHcajiH eme uenbift pan BHHOBbix h
HH(J)paBHAOBbix (JjopM H3 pasJiH^Hbix paHOHOB IlajieapKTHKH. HaKoneu, aecKOjibKO
BHflOB oHHcano cpaBHHxejibHO HenasHo (Pisarski, 1967; XUiyccKHH, 3a6eJiHH, 1985;
^yccKHii H Ap., 1990; PaA'KHKO, ApaKenHH, 1990). TaKHMo6pa30M,K nacTOHmeMy
BpeMeHHB riajieapKTHKe onncaHO 6onee 60 bhaobwxh HH(})paBHAOBbix4)opM Tetramo-
rium. CnoacHocTb h 3anyTaHHOCTb CHCTeiviaTHKH «aHHoro poAa cBHsana c tcm, hto 3Th
MypaBbH BecbMa HSMeHiHBbi b CKynbnrype, OKpacKe h 4)opMe lena, h flOcTOBepHbe
onpeaeneHHe 6e3 caMOK b pHne cnyiaeB BecbMa caojkho hhh Aa)Ke npaKTHiecKH ne-
BO3M03KHO.
IlpH npoaeaeHHH HacTOHmefi peBH3HH 6bin H3yqeH MHoro<iHcJieHHbiH MarepHaji, b
TOM HHcne THnoBOH, npaKTHqecKH h3 BcexpafioHOB IlajieapKTHKH, xpaHHiuHHCHb koji-
neKUHHX SooAoraqecKoro Mysea MrV (3M) MocKsa, HncxHTyTa 30onorHH AH YKpa-
,
HHbi (H3), KneB (BKAiOHaH KOAAeKUHKi B.A. KapaBacBa), 3HH PAH, CaHKT-IleTep-
6ypr, HHCTHTyra 3oonorHH HAH (H3 HAH), BapmaBa, a TaK^Kc onncaHHH h pHcyHKH
THHOB H3paanH^Hbix My3eeB EBponbi, cAenaHHwe npo(}). B. IlHcapcKHM^
B HacTOHiuee Bpeivia na reppHTOpHH CCCP AOCTOBepao H3BecTHo 22 bhab MypaBbes
H3 poAa Tetramorium, pHA cxapbix yKasanMH Bbi3biBaeT coMHCHHe (cm. HH5Ke). Mx
Bcex MOJKHo oBbCAHHHTb B HecKOJibKO Tpyno.
'CnHCOKJiHTepaTypfainpHseneH boBTopoMcooSmeHHH.
'BMpaJKaio HCKpcHHioio BnaronapHocTb f.M. UnyccKOMy k B. llHcapcKOMy sa jiio6e3Ho npe-
HocraBJieHHyK) B03Mo>KHocTb o5pa6oTKH KonneKUHOHHoro MaTepnajia, a xaKJKe ueHHue coBCTbi
HyKasaHHHnpHnonroTOBKenaHHoftpaSoTti.
39
i\
. .
—
1. Fpynna T. striativentre y caMOK h pa6o'iHx I reprHT GprouiKa b rycTofl cTpyfi-
iHTOH CKyjibnrype: striativentre, schneideri, kabulistanicum.
2. Fpynna T. karakalense - y caivioK h pa6oMHX SpwiuKo r/iaAKoe, ho yero ocnoBa-
HHH ecTb leiKHe seepooSpasHO pacxoflnnmecH MopmHHKH: b rpynne ofljiH bha.
3. Fpynna T. caespitum — oinnMaioTCH uiHpoKHM h sbinyKjibiM cKyryMOM caMOK,
B npo4)HJib HaBHcaiomHM Haa npoHoiyMOM; yrnbi npoHoiyivia cBCpxy He bhahh:
caespitum u, CKOpee Bcerojacoti.
4. Fpynna T. semilaeve — caMKH c nnocKHM cKyiyivioM, He HaBHcaroniHM cBepxyHan
npoHotyMOM; yrjibi npoHoryMa bhahw cBepxy. FonoBa h rpynb paSoTOx MopmnHHc-
Tbie semilaeve,forte, turcomanicum, reticuligerum.
:
5. Fpynna T. inerme — caMKH no 4)opMe rpyflH cxoflHbi c snaaMH npeAbinymefl
rpynnbi, OAHaKo ronosa h rpynb paSoMHx, no KpanHen Mepe CBepxy, rjiaxiKHe h 6;iec-
TamHe,jih5o c oncHb cjia6oH noBepxHOciHOH CKyjibnTypofl: inerme,armatum, nitidis-
simum, concaviceps, kisilkumense, vernicosits.
6. Fpynna T. ferox — caviKH c oieHb xapaKTepHbiMH cnnbHo pacujHpeHHbiMH iJieHH-
KaMH cieSenbKaH rnanKHMH MaHAH6yjiaMH:/ewx, aegea, perspicax, densopilosus,
conftnis: k 3toh >Ke rpynne, no-BHflHMOMy, othochtch hbursakovi
FIpoMepbi H HHfleKCbi: HF — nnHna ronoBw crporo cnepeAH ot saTbinouHoro KpaH
flo KpaH HajiHUHHKa; ecjTH aaTbuioMHbiH KpaH Bornyr — ot jihhhh, coeOTHHromeH san-
HHC yiTibi; lUr — MaKCHMajibHaH uiHpHHa ronoBbi cnepean; HFn — HanSojibrnHH ayi-
aivierp rnaaa; JJC — AJiHHa cKanyca; ^M— Hnnna rpyjiji c6oKy no flHaronajiH ot Mec-
la coijieHeHHH c ronOBOH (y pa60MHx) HnH ot nepeHne-BepxHero Kpaa MesoHOTyMa
(caMKH, caMnbi) AO KOHua DHHcTepHaiibHMX jionacTeii; BM ^ BbicoTa rpyziH c60Kyot
HHKHeH ToqKH McsonneBp nepneHAHKynapHO jxo Bepxa MeaoHOTyivia; Mil — luHpHHa
yaenKa neTHOjiH CBepxy; JliUn— to >Ke nocTncTHOjia.
B nepBoii nacTH paSoTw npHBeaeHw onHcaHHe hobwxbhaob h onpeflenHTenbHaa Ta6-
HHua Tetramorium t^aynbi CCCP, bo BTOpofl — cHCTCMaTHMecKHH ogsop.
Tetramorium vemicosus Radtschenko, sp.n.
MaTepHaji: caMKa (rojioTHn), FHccapcKHH xpeBer KoHjiapa, 21.VI 1973
(B. HnyuieB) ; napaTHnw: 9, PencTeK, loro-BocTOHHbie KapaKyMbi, 19.V 1982 (KpH-
BoxaTCKHK) 5 99, KHprH3HH, AKcy-Jl)Ka6arJifai, 1300 m nan yp. m., 20.VI 1961
;
(F. flnyccKHH); 1099, YaeeKHcTaH, ceBepo-aanaanee HuiKeHTa, 1500—1700 m na/i
yp. M., 3.V 1942 (K: ApaojfibflH) ; 1099, IOjkhwh YaeeKHCTaH, Bapranaa, N" 78-137,
30.V 1978 (F. ^InyccKHH) 7 pa5oHHx, yaeeKHcTan, KauiKa-jJapbH, 26.V 1942 (K. Ap-
;
HonbHH) Innbi xpanaTCH a 3M MFY h H3 AH yKpaHHbi.
OnHc.aHHe. C a m k a (pHC. 1,a, 5).
FonoBa nonepeqnaH (/fr-0,95-0,96) c napajinejibHbiMH BoKOBbiMH cTopoHaiviH h
,
cJierKa BornyTbiM 3aTbUiovHbiM KpaeM. Jlo6 c MHoroHHCJieHHWMH tohkhmh Mopmnn-
KaMH, mCKH H BHCKH C UITpHXOBaTOH MOpUlHHHCTOCTblO. CKanyc He flOCTHraeT jaTbl-
noMHoro KpaH (//C-1,22—1,24) »
CKyryivi rnaxncHH « gjiecTHUiHH; cKyTennroM rnanKHH (ronoTHn) h;ih c tohkoh no-
BepxHocTHoii MopmnHHCTOcTbio (napaTHn) ; nponoaeyM csepxy c He>KHbiMH Mopmnn-
KaMH. BoKa rpy^H co cnaSbivm, kopotkhmh MopmHHKaMH. nponoaeyivi c kopotkhmh,
OCTpblMHJpeyrOJIbHblMH 3y6wHKaMH.
yjieHHKM CTeSenbKa He pacujHpeHHbie (/fy7-2,2—2,8, /ffl/Z-l,8-2,1), qeuiyHKa ne-
THona ynnomeHa,b n-po^vuihKjiHHOBHflHaH.
OTCTOHiuHe BonocKH HMeraTCH na Ji6y, HaimMroiKe, TeivieHH, no 6oKaM h cHH3y
ronoBbi, na CKyTyivie, cKyTCJijiMMe, HJieHHKax cTe6enbKa h na sceii noBepxHOCTH Tep-
FHTOB epjounca.
Teno HpK0-pbi>Kee, Bneciamee.
FasMcpbi: ^r-1,09, Z^/F-l,15-1,13 (1,15), ^C-0,88-0,9 (0,9), jm-\,99-2,Q6
(2,06) MM (b cK06Kax — roftoTHn).
Pa 5o qH e (pHC. 1.e, ^)
40
Phc. 1. Tetramorium vemicoms, sp. n.: a, 6 - caMKa, rojioTwi; e, ^ - paGo^Hft.
napaxHn (a, e - rojiosa cnepenw, 6 - rpyflt h cTeSeneK csepxy, a- rpynbH
CTe6eneK bnpo(J)Hnb)
FonoBa yflnHHeHHan (i/T-l,1—1,2), c napajiJiejibHWMH Bokobhmh CTOpoHaMH, luh-
poKo 3aKpyrjieHHbiMH saflKHMH yraaMH h npHMWM saTbuipqHbiM KpaeM. MopuiHHHC-
TocTb rojioBbi HOKHaH, HOBepxHocTHaH, HO MopinHHKH HBCTBeHHbie; HHOFflacKynbnTy-
pa crjiaaceHHaa h rojiOBa noMXH rjiaAKaH. Tnaia cpeoHHx pasMepoB (///yz-5,2-5,9),
CKanyc HCMHoro He hoxohhtno saTbuioqnoro Kpan (/fC-1,16—1,22).
Fpynb nnHHHaa (//A/-2,4—2,8), c HBCTBeHHWM MeaonponofleajibHbiM BflaBJieHHeM
(y HeKOTopbix 3K3eMnjiHpoB H3 KauiKa-ilapbH oho sbipajKeHo cjia5ee), cBepxy h c
BOKOB C He^HblMH nOBepXHOCTHblMH IHTpHXOBaTbIMM MOpmHHKBMH, 6jleCTflIUaH.
OrcToaiuHe bojiockh na lene lOHKHe, HeMHoro^mcneHHwe.
YsejioK neraonH saxpyrneHna seprnHHe, c HepesKoii njiomaflKoii; mjichmkh creSenb-
Ka rjiaflKHC h 6jiecTHmHe.
Tejio oxpHCTO-jKCJiToe.
Pa3Mepbi:/7-r-0J3-0,78,j!tfr-0,66-0,71,^C-0,59-0,69,,iZM-0,88-0,98 mm.
C a M U bl HeK3BeCTHbI.
XI,H(J)4)epeHu,Ha7ibHbiH aHarHos. BuusaHHMaeTnpoMOKyroqHoe nojio»e-
HHe MCJKAy T. semilaeve h T. nitidissimum - T. kisilkumense. Pa6oTOe oinHiaioTCH
cKynbniypoH rejia, cbmrh - cKy.nbnTypOH Me30H0TyMa h 5ojibmHMH pasMepaMH.
Tetramoriumbwsakovi Radschenko, sp. n.
MarepHaji: caMKa (rojioran) K)>KHbiH KaaaxcraH, BOCToqHbieoiporH Kaparay,
,
crenb, N° 48. 13.VI 1979 (C. BypcaKos) 8 99, 9 99 h 9 do.(napaTHnbi), H3 Toro kb
:
rHesna. THnw xpaHHTCHb 3M MTVh 143 AH YKpaHKbi.
OnHcaHHC. CaMKa
(pHC. 2,a-e) TonoBa cnerKa nonepeinaH (Mr-0,94—
.
0,97), HeMHOro cy>KeHa k nepeflaeMy KOHiiy, saxbinoHKbiH Kpaft npsMOH. Bch ronosa
noKpblTa MHOroqHCJTCHHblMH HBCTBeHHblMH npOnOJlbHblMH MOpillHHKaMH, 3axOnHmH-
MH Ha saibuioK; MaHflH6yiibi c nepesKHMH MopinHHKaMH. CKanyc He flocTHraer sa-
TbiJiOHHoro Kpan (//C-1,35-16,38).
Fpy^b CBepxy m c 6okob c MHoroqHcneHHbiMH npoHOjibHbiMH MopinHHKaMH, HHoraa
cKyTe.njiioM qacTMiHO rjiaaKHHh SnecTHiuHH, he nponofleyMC npeoSnaflaioT uferpeHCBaH
cKyjibHTypa h nepesKHe nonepe^Hbie Hyroo5pa3Hbie MOpmHHKH; uiHnHKH flOBonbHo
WTHHHbie, OCTpbie.
41
Phc. 2. Tetramoriumbursakovi, sp. n.:a-e - camKa, ronomn; ^,^ - caMeu; e -
pa6oHHft, napaTHnu (a - ronoBa cnepepiH; 6 — rpyjib H cTe5eneK eeepxy; e -
rpyflb H cTe5eneK b npo(J)Hnb; 8, ^ - CTe6eneK b npo(J)KJib; d - to )Ke CBepxy)
IjieHHKHcTe6ejibKapacampeHbi cJia5ee, hcm ybhaob rpynnw 7! ferox, ho cHJibHee,
HCM y npeACTaBHTejiCH npyrax rpynn (/f/7-2,0—2,75, /fl777-l,49—1,58). MeiuyftKa
nexHOJiH c KopoTKOHHaKiroHHOH BeiuHHHOH ruiomaffKOH, ee nepeflHHH Kpafi cBoKy He-
MHoronpHocTpcH, c Heray5oKOHBbieMKOH nocepeflHHe.
Ha n6y, saxbuiOMHOM Kpae rojioBbi, HajiHMHHKe h BHHsy rojioBw HMerarcH mhofo-
iHCJleHHbie OTCTOHIKHe BOnOCKH. CKytyM H CKyTCJIJllOM C MHOrOHHCJieHHblMH OTCTOH-
iUHMH BonocKaMH; qjieHHKH CTe6eJibKa c flnHHHbiivm, HanpaBneHHbiMH HaaaA otctoh-
iUHMH BonocKaMH. npHJie5Kainee onymcHHe Qprnuaca penKoe, paccTOHHHe Me>KAy
BojiocKaMH npHMepHo paBHo Hx OTHHe; oiCTOHume bojiockh HMeioTca wa bcch no-
BepXHOCTH leprHTOB H CTepHHTOB.
Ubct TeMH0-6ypbiH, HorH»H ycHKH CBemee.
PasMepw: ^/'-0,82-0,87 (0,84), Z(tt"-0,88~0,91 (0,88), MC-0,6-0M (0,62),
iW-1,47-1,6 (1,53) MM.
C a Mu bi (pHc. 2,a, d).
FonoBa cy5KBaHpaTHaH (fff-l,19—2,2), rnaaa KpynHbie (///>z-2,15-2,34). Jlo6
c HBcTBeHHbiMH Herpy6biMH MopiuHHKaMH, cxoflHiuHMHCH K rna3KaM, B ueHTpe cnpo-
aojibHOH, 6ojiee hhh MCHee rnaflKOH nonocofl; MOKay rjiasKaMHh rjiaaaMH MopiuHHKH
qacTHMHo saMeHCHbi npoflojibHWMH panaMH tomck. HeMHorouHcneHHbie ji/iHHHbie ot-
CTOHiuHe BOJiocKH MOryT 6biTb Ha HaiiHiiHHKe, Ji5y h 3anHHXyrnax; cHH3y rojioBbi ne-
CKOirbKO KOpOTKHXBOJIOCKOB.
Fpyflb nonHocTbio noKpbira MHoroqHcneHHwMH tohkhmh MopiuHHKaMH; crjia>KeH-
Han cKyjibnrypa mokct 6biTb Ha MesonjieBpax h b bhac ysKoA nonocbi Me>K;]y Horay-
nHMH; nponofleyM Sea 3y5iHKOB, caMoe Sojibuiee - c rynbiMH 6yrpaMH. OrcTOHiuee
onymeHHe BapbHpyer, ho Bcerna ecTb no KpaHHeft Mepe necKOjibKo nnHHHbixbojiockob
Ha cKyxyMC h cKyrenjiioMe.
4jieHHKH CTe6ejibKa pacuiHpeHb! OTHOCHTenbHO cJia6o (ii77-l,58—1,91, /ffl/7-1,12^
1, 27). rieTHOjib He ynjiouien, b npo(|>Hjib c KopoiKOH BepuiHHHofl nnoiuaflKOH. yjicHH-
42
. .
Phc. 3.flexajiH cxpoeHHHT.striativentre (a, e~d),T. kabulistanicum (6). T. schne-
ideri (e-3), T. reticuligerum (u) (a, 6 epiouiKO pa6oMnx caepxy; e, e, u -.ro-
noBa pa6o<rax cnepeflH; d, 3 - to >Ke camoK; e, jk - rpym> h creSeneKpaSoMHX
Bnpo4)HJib)
KH CTe6ejtbKa C P/IHHHblMH OTCTOHIUHMM BOJlOCKaMH; Ha OplOIllKe OTCTOHmHe BOJIOC-
KHecTb no KpEHM TeprHTOB H Ha aceftnosepxHOCTM cxepHHTOB.
TenoleMHo-Sypoe ao wepHoro,hofhh ycHKH oxpHcibie.
-Pa3Mepbi:^r-0,62-0,67,/i/r-0,64-0,69,;7C-0,28~0,29,iZM-l,85-1,9 mm.
PaSoHHe
(pHc. 2, e).
TojioBa npHMoyrojibHan (///"-l,02-1,06), c napajiJiejibHbiMH 6oKOBbiMH cTOp^oHa-
MH H npHMbiM saTbiJioHHbiM KpacM. CKanyc HeMHoro He flocTHraeraaibiJioqHoro Kpasj
(/fC-1,33—1,46) BcH ronoBa c MHoroqHCJieHHbiMH HerpyBwMH npoHo;ibHbiMH MOp-
.
mHHKaMH. Tnaaa cpennHx pasMepoB (MF/i-5,5-6,0)
Tpynb c HBCTBCHKbiM MeaoHpoiipaeanbHbiM BAasneHHeM, umnHKH nponofleyMa p
BHne pasHoBenpeHHoro TpeyrojibHHKa, ocipbie. Ilpo- h MesoHoryM c apononbHbiMH
MopuiHHKaMH, nponofleyM cBepxy Meacny uiHnHKaMH c marpeHeBoii cKyjibniypOH
IneHHKH cTcgeJibKa cBepxyrnanKHe, 6JiecTHmHe.
Bee TejIO C MHOrOHHCneHHWMH otctohikhmh BOnOCKaMH.
Ubbt KpacHOBaro-SypbiH.
Pa3Mepbi:^-0,71-0,78,/tfr-0,7-0,76,^C-0,52-0,57,.3M-0,9-0,99 mm.
43
.
Phc. 4. ileTajiHcTpoeHHfl T. kamkaleme (napaTHn) (a), T. caespitum (6, e), T.ja-
coti (a) H T.forte 'd) (a- SpwuiKo pa6o>Mx csepxy, 6-d ~ rpynt cumok
B npo<j)HjibHCBepxy)
ijH(j)(J3epeHuHajibHbiH aHarH0 3. OnHcwBaeMWH bhhsaHHMaernpoivieacy-
TCJHoe nojiOKeHHe Me5KAyrpynnaMH T. ferox h T. semilaeveh xopoiuo oiramaeTcH ot
BcexBHAOB cTpoeHHeM cieSeJibKa caMOK. PaBoTOe cxoahm c T. densopilosus, omHiaioT-
CH cKyubnTypoH rpyflHh MCHbumMH pasMepaMH.
OnPEAEHHTEnbHAH TABnHUA MVPABbEB POflA TETRAMORWM
MAYR (DAVHW CCCP (PABOHHE H CAMKH)
1(6) 1 leprHT epiomKa xoth Sbi nacraqHO a npoflOJibHoH cTpyiiMaTOH cKynbnrype,
uienKOBHCTbiH (pHc. 3,a, 6). TojiOBa h rpynb rpy6o cKynbnxHpoBaHbi.
2(3) CTpyfiqaiaH CKyjibniypa pasBHia b nepeaHeft nonoBHHe I reprHra 5pK)ujKa,
3aflHHH qacTb c cna5oH marpeHCBOH cKyjibniypoH h^h rJianKaH (pHC. 3,6)
;
rojiosaH rpyAb paSoiMx KpacHOBaTo-KopHimeBbie, 6pK)uiKO TeMHo-6y-
poe T. kabulistanicum Pisarski
3(2) I TeprHT SpwiuKa nojiHocTbio b CTpyHHaroH CKyjTbnrype; reao pagoHHx oa-
HOUBCTHoe, TeMHo-5ypoe.
4(5) Pa6oqHe: noQ c pesKHMH, rpygbiivra npoflOjibHbiMH MOpiuHHKaMH. hx kojih-
MecTBo Me>KAy jioShwmh BajiHKaMH Ha ypOBHe raaa ne 6oabiiie 20 (pHC. 3,e)
ripononeyM c rpeyronbHbiMH 3y6iHKaMH (pHC. 3,z). YsenoK ncTHOJiH cBepxy
cnerKa nonepeiHbiM. CaMKH: jio6 c rpy6wMH npoaojibHbiMH MopmMHKaMH,
44
.
MopmHHHCTaH cKynbHTypa npeoSnanaeT MOKfly rnaaaiviH h jioBhmmh sanH-
KaMH (pHC. 3,d). Me3onjieBpbi cocrnaaceHHOHMopmHHMCTocTBK) H pasBHToft
uiarpeHeaoH cKyjibnrypoH. lUnnbinpoiioHeyiviaKopoqe 7! striativentre Mayr.
5(4) PaScme: no6 c MHoroqHcneHHbiMH rycTbiMH nerpyBbiMH MopmHHKaMH
(pHc. 3,e). riponofleyM canHHHbiMHiuHnHKaMH (pHc. 3,j«f). VaenoK nerao-
nH cBepxy cnerKa ynnHHeHHbiH. CaMKH: no6 c HerpySwMH nponojibHbiMH
MopmHHKaMH, Me>Kfly rnaaaMH h noSHbiMH sanHKaiviH npeoSJiaflaei uiarpeHe-
Baa CKynbHTypa (pHC. 3,3). Mesonnespbi cnponoJihHOH MopiuHHHCTOCTbio
IlbinbinponoAeyManjiHHHee T. schneideri Emery
6(1) I TeprHT BpiouiKa rjiaHKHH h 6necTHinHH, jihBo c noBepxHociHOH HaoMCxpH-
TOCKOH cexqacTOH CKyjibHTypoH, jim6o c kopotkhmh rpyBbiMH MopmHHKaMH
yocHo:^aHHH (pHC. A,a).
7(8) riepeAHHH nacib rojioBbi paBoHHx h caMOK MopmHHHCTaa, saflKHH — c hbct-
BCHHOH luarpeHeBOH cKynbnTypoii (pHC. 3,m). CKyiyM caMOK xoth 6bi b ne-
peflHCH TpeiH rjianKHH h BjiecramKH. TeJio <iepHoe hhh Bypoaaro-qep-
aoe T. reticuligerum Bursakov
8(7) FojioBa paBoMHx jihBo nojiHocTbio MopmnHHciaH, jihBo rjiaflKaa; tarn saa-
HHH nacTb rojioBbi caMOK c marpeHCBOH cKyjibniypoH, to Teno BypoBaxo-
KCJiToe.
9(10) BpwuiKO caMOK h paBoqnx c KOpoxKHMH rpyBbiMH npoflonbHbiMH MopmHH-
KaMH y ocHOBaHHH (y lacxn paBoHHX MojjmnHKH Moryx-Bbixb necKOJibKO
crjiajKeHbi) (pHC. 4,a). Y paBoiHX sepx rpyflH h yaejiKH cxeBejibKa MOp-
mHHHcTbie T. karakalense DJus. et Zabelin
10(9) BpwiiiKO pa6o»mxh cbmok Bes MopiUHHOK y ocHqaaHHa.
11(32) laeHHKH cieBeJibKa caMOK He paciuHpeHHwe, nexHonb HBciBeHHo yace nocr-
neiHOJiH (pHc. 4,e, d\ 5, a) (HUn He Menee 1,8; ///777 paBoMHx 2,35-2,9,
y T. caespitum - 2,28-2,7)
12(13) Fpyflb caMOK BbicoKaa, b npo(J)HJTb cJierKa BbinyKnaH, cKyxyM HaaHcaex
Han npOHoiyMOM h nojiHocxbio npHRpwaaex ero caepxy (pHc. 4,5, e) Fono-
.
aa paBoTOx c HBcxBeHHOH npoflonBHOH MopmMHHcxocxbK), no Bokbm rpynH
npeoBnanaex rpaHynHpoaaHHaa cKy/ibnxypa, ysejiKH qjieHHKoa cxeBenbKa
caepxyrnayKHe h BnecxHUOie T. caespitum (L)^.
13(12) Fpyflb caMOK HeBbicoKaH, cKyiyM He npHKpwaaex caepxy nepeflHHx yrJioa
npoHoiyMa (pHc. 4,d, e)
14(15) Fpyflb caMOK a npo(J)HJib cnexKaawnyKnan,/W-1,6-1,8 (pnc. 4,^). CKyxyM,
no KpaHHefi Mepe b nepeflHefl nojioBHHe, rnaflKHH h BnecTHmnH. KpynHee:
JIM Bojibme 2 mm. Fonoaa paBoMHX MopmHHHcxafl, ineHHKH cxeBenbKa rjiafl-
KHe H BjiecTHiflHe, HHorfla c HerpyBoH noaepxHocxHOH cKy.nbnxypovi. Ilaex
paBoiHxH caMOK ox xcMMHoM-Byporo flo qepnoro . . . T.jacotiWheeMleMr
15(14) Fpyflb caMOK njiocKaH, Bojibuie 2,1 (pnc. S,6, a); jwenbqe: ne Bojiee
1,9 MM, ecnH Bonbme, xo rpyflb caepxy nonHocrbw rnaflKaa, 6.necTHiuaH h
uaex opanaceabiH. Fonoaa h rpynb paBoiHX nnBo cKynbnxHpoaaHHbie, nnBo
raaflKHe.
16(17) Fpyflb caMOK csepxy rnaflKaH h BnecTHman, ^M-2,0-2,1 mm. PaSoMne Meji-
KHe, rojioaa c cHjibHo crjia>KeHHbiMH noaepxHocxHbiMH MopuiHHKaMH, Bnec-
XHiflan; rnasa cpeflHHX pasMepoa, MFn-S,3—5,1\ rpyflb a npo4)Hjib y Bojib-
umHcxaapaBoHHXc HBcxaeHHbiM MesonponofleaiibHbiMBflaaneHMeM T. vemi-
cosus sp. n. \
17(16) Fpyflb caMOK ceepxy BojibuieH qacxbio hjih nonHocxbM c aacxaeHHbiMH npo-
flonbHbiMH MopinHHKaMH.,iZM He Bonee 1,9 mm. Ecjih paBoqne Kenxwe h co
crnajKCHHOH CKyjibnxypoH ronoBbi, xo rnasa Mejibie (IWjI-6,0—8,6) h mbso-
nponofleajibHoe BflaanenHe ne Bbipa)KeHo.
^UnaKptiMa H VpajibCKofto5n. (HacoHOB, 1889; Py3CKHft,1905) 6bijiyKaaan T. meridionale
Emery. Pa6o<JHe 3Toro BHfla oxjiHijaiOTcaox T. caespitumHanmneM nonepeiHUX MopiUHHOK Ha 3a-
xbinKe.
45
i\iaiU
(.^
Phc. 5. flpiajiHcTpoeHHH T. armatum (a-e), T. inenne (e, d), T. kisilkumense (napaTHn)
(e).r. nitidissimum {cvBmmhi)(X'-u), T. concavkeps (napa-TKa) (v) fa - rpyatcaMKH
cBcpxy; 6, ^, jc - rpyflb hcTe6ejiekcawoKb npo^unb; e, l e, u ~ rpynt\\ cTeBeJicKpa-
60MMX B npoiJjHiTb; 3, /<• - roJioeapaSoMHxcnepeaw)
18(19) rpyflb caMOK cBepxy nojiHocTbio noKpwTa rpy5wMH npoaojibHUMH Mop-
uiHHKaMH, lUHnbi nponoaeyMa ocrphie, wacTo najjOHKOBHflHbie (pHC.4,<));
Ha 5oKax rpyoH pesKHe MOpiuHHKH, uiarpeneBaH cKyjibniypa iie pasBHia.
KpynHee:,ZJ')^-],7-),9 mm. ro;ioBa h rpy^b pa5o>jHX, BKJUowaH Meaorwespbi,
c rpy6biMH npoflonbHbiMH MopiuHHKaMH; ysenKH iJieHHKoB cieSenbKa caep-
xy MopuiHHHCTbie. CaMKH H paSowe wepHbie, HHorna ronoBah rpyub paBowx
KpacHOBaTO-Bypbie 7! /«r/e For.
19(18) Tpyflb caMOK caepxy c HerpyQbfMM MOpiuMHKaMH, cKyTe.n;noM h riepeaHaa
nacTb CKyryMa xoth 6b( iiacTuiHo rnajiKMe h SnecramHe (pHc. 5,a); Ha fioKax
rpynH, noMHMO mopihhhhctoh, pasBHTa ujarpeHeaaH CKy.(ibnTypa. FonoBa
pa6oMHx MOpmHHHCTafl HJiH iTiaflKan, yaenKH qjieHHKoa creBeJibKa caepxy
no KpaijHeH Mcpe nacTbio rnajiKHe h enecTHuiHe.
20(21) CaMKH Kpynnee, co cnaSo ynjiomeHHoii rpyflbw (iZM-J ,45 - 1,55 mm,
/2M-2,15-2,2), TeMHO-Bypbie. Fojioaa h aepx rpysH pago^jHx cpesKHMH npo-
AOJibHbiMH MopuiHHKaMH, Ha SoKax rpyaH, HapHfly c npoflOJibHoii MopmH-
HHcTocTbK),HMeeTCH 6yropHaTafl cKyjibnrypa; yaejiKH nerHojiH caepxy
46
-
. .
qacTbK) cKyjibnTHpoBaHbi; reno TeMHO-6ypoe no yepHoro T. turcomanicum
Emery
21(20) CaMKH Mejil.qe (aW-1,25-1,4 mm), ecJiH Kpynnee, to lejio KopHMHeBaTO->Keji-
Toe. FonoBapagoMHx rjiaflKan, HHoraa c noBepxHocTHoft lUTpHxoBaTOJi Mop-
mHHHCTOcTbM; eoiH MOpiuHHHCTaH, TO MopiuHHKH T.oHKHe, rycTbie, yjcnKH
qjieHMKOB cTeSejTbKa cBepxy nonHocTbio rjianKHe h ubct Tena 5ypoBaT0->Ken-
Tbiii.
22(23) CaMKH MexiKHe (.iJW-1,25-1,3 mm), c yMepeHHO ynjiomcHHOH rpyflbio
(MM-2,l9'-2.n) mcKH, bhckh h npocTpaHCTBo Me^uy BHCKaMHH jio6HbiMH
;
BanHKaMH C nepeSKHMH, ho HBCTBeHHWMH npOflOJIbHblMH MOpmHHKaMH. Fo-
noBa paSowx c rycTbiMH npoflojibHWMH MOpmHHKaMH, rpynb cnepxy Mop-
lUHHHCTaa T. semilaeve E. Andre
23(22) CaMKH: Ha5op npH3HaK0B hhoh. Fojiobb paSoHHx rjiaaKaa h 5;iecTHmaH,
HHorna no6 c oieHb nexcHbiMH nosepxHocTHbiMH MOpmHHKaMH, ho rpyflb
CBepxy FJiaflKaH hhh cnaSo nyHKrapoBaHHaH.
24(27) Fpynb caMOK c;ia5o yruiomena (MM-2,0-2,2) (pHc. 5,6, e); pa6oTOe Bypwe
HJiH >KenTOBaTO-KopHqHeBbie, c KpynHbiMH rjiaaaMH (///yj-4,7—5,5).
25(26) nponoaeyMcaMOK6e3flBcTBeHHbix3y6uoB, b npotJjHJibyrjiOBaTWH (pHC. 5,e)
;
nponojieyM pa6o>oix c oqeHb kopotkhmh 3y5iHKaMH anw. 6e3 hhx
(pHc. 5,d) T. inerme Mayr
26(26) riponoaeyM caMOK h paSowx c HBCTBeHHbiMH (bcrpbiMH syfiuaMH (pnc. 5,6,
8) T. armatum Emery
27(24) FpyAh caMOK cHjibHo ynnoiuena (Mf-2,3-2,6) (pHc. 5,:w) (caMKa T. kisil-
kumense HenaBecTHa) pa6oqHe xenibie hjih KpacHOBaTO-jKenTwe, rna3a
;
MCJifaie (/f/yz-6,0-8,6).
28(29) F;ia3a paOoTOx Kpynnee (//ry2-6,0-6,3) ysenoK neTHOjiHb npocjjHnh saKpyr-
;
jieHHbiH, 5e3 HBcTBeHHOH BepmHHHOH njlomarfKH (pnc. 5, e) . . . E. kisilkumense
Dlussky
29(28) Fjraaa paeomx Meiibqe (//r«-6,7-8,6) ysejiOK neTHOJiH b n^o^vinb c hbct-
;
BCHHOH BepmHHHOH lUIOmanKOH (pHC. 5,M).
^M
30(31) Kpynnee: i7M caMOK 1,47-1,55 mm, pa5o™x 0,9-1,0 mm. FojioBa pa6o-
TOx c HBCTBCHHo BOFHyTbiM saTbmoMHbiM KpacM, masa Kpynnee {MF/i-SJ
31 (30) M7e,j0i)b^(ep:HcM.M5,«c:)bmok 1,3-1,4 mm,TM. cIonpcaav6iocqeHpXs0B,u7r-s0a,k8ovmm. FojiOBa paSoMHX c
npHMbiM Hjm MyTb BorayTbiM aaTbinoTObiM KpaeM, rna3a Menb^ie {MF/i-l,Q~
8,6) (pHc. 5,3) T. nitidisimum Emery
32(11) MjieHHKH cTeSenbKa caMOK cH^bHo pacmnpeHbi, ncTHonb HCMHoro ywe hoct-
neTHOjiH; M7/7-1,1-1,4 (pnc. 6,a,d) (y T.bursakoviHUn-l,5-1,6),HHH p-i-
fcifflx 2,0—2,32, ecJiH Sojibme, to ronoBa rnaaKaa h 6jiecTHmaa.
33(44) CaMKH.
34(35) I TeprHT SpiomKa c rycTbiM npHJie)KainHM onymeHHeM,fljinna bojiockob b hcc-
KOjibKo pa3 Bojibme paccTOHHHH Me5Knyhmmh (pnc. 6,a) T. densopilosus
Rad. et Arakel.
35(34) PaccTOHHHe Me>Koy npH.ne5KamHMH sonocKaMii Ha I TeprHTe SpiomKa npHMep-
Ho paBHo HxnJiHHe hjih Sojibme (pnc. 6,6)
36(39) CKyiyM h ronoBa rnaaKHe.H BnecTHmHe, HHorAa no6 c cna6biMH noBepx-
HOCTHblMH MOpmHHKaMH (pHC. 6,s). TCJIO KpaCHO-KOpHHHeBOe.
37(38) nocTneTHOJibmHpe; ;f7777-l,04-l,17 T. ferox Rnisky
38(37) nocTneTHojiby>Ke:/fy777-l,27-l,36 T. ae^ea Bar. Urb.
39(36) CKyTyM h ronoBa c MHoroqHcneHHbiivtH pesKHMH npoflo/ibHbiMHMopmHHKaMii
(pHc. 6,a,Tx)
40(41) HneHHKH cTe6ejibKa cJiaSee pacmnpeHbi: M7/7-1,5-1,6 (pHC.6^)... T. bursako-
vi sp. n.
41(40) IneHHKHCTeeejibKa mHpe:M777-l,l- 1,2 (pnc 6,a).
47
.
Pwc.6.HeTanacrpoeimsi T. densopilosus (ronoTHn) (a), T. aegea (chhthd T. levigata Karawj {6-8),
T. confinis (ronoTHn) (z-e). T. bunakovi (napaTWi) (jr). T. perspicax (napaTWi T. feroxoides Dlus.
et Zabelin) (3) (a, 6 - lacTb Teprnxa 6pK)UjKa cawoK; e, d - rojioBa caMOK cnepenH; ^ - rpynb
caMKHcBepxy; e, 3 - rpyni.hCTe6eneKcaMOKb npo())Mnb; m - cTe6eneK caMKH CBepxy)
42(43) CKyryM c MHoro>oicjieHHbiMH otciohiuhmh BOjiocKaMH. Kpynnee: JHM-1,9-
2,0 MM T. confinis Rad. et Arakelian
43(42) CKyiyM ronbift hhh c enHHHMHbiMH otctohiuhmh BOnocKaMM (pHC. 6,3).
Menbqe:/P/-1,5—1,6 mm T. perspicax Sant.
44(33) Pa5oTOe.
45(46) FonoBa h rpynb csepxy rjiaflKaa h 6JiecTHmaH, caMoe Soxibiuee b HH>KHeH qac-
TH ronoBbi Moryi 6biTb cjiaBwe 'noBepxHociHbie uiTpHXOBaTbie MOpmHHKH,
npoHoiyMrnaflKHH T. aegea Baroni Urbani"
*CxoAeH c T. inerme, oinwiaetcsi MenKHMH rnaaaMH (/f/Ji-5,8-6,5) h 6onee CBexnoft oKpacKott.
48