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ANNOTATED CHECKLIST OF THE VASCULAR FLORA OF THE LOBLOLLY UNIT OF THE BIG THICKET NATIONAL PRESERVE, LIBERTY COUNTY, TEXAS PDF

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ANNOTATED CHECKLIST OF THE VASCULAR FLORA OF THE LOBLOLLY UNIT OF THE THICKET NATIONAL BIG PRESERVE, LIBERTY COUNTY, TEXAS MacRoberts Barbara R. Larry Brown and Michael MacRoberts H. E. Herbarium, Spring Branch Science Center Bog Research, Museum of Sciences Life i, [email protected] Louisiana State University in Shreveport Shreveport, Louisiana, 71 U.S.A. 15, 1 Warren W. Pruess and Sandra Suzanne Birmingham Walker Elsik I. Department of Ecology and Azimuth Forestry Services, Inc Evolutionary Biology 14671 Highway 87 South State Rice University Texas 75973 Shelbyville, U.S.A. Houston, Texas 77005, U.S.A. INTRODUCTION This paper is the 6th part of an inventory, based on available voucher specimens, of the vascular flora of the Big Thicket National Preserve in southeastern Texas. In 5 previous papers, we provided annotated checklists of the Hickory Creek, Turkey Creek, Big Sandy Creek, Lance Rosier, and Beech Creek units (MacRoberts et al. Brown 2002; et 2005, 2006a, 2006b, 2008). At this time, the Menard Creek Unit has been inventoried, and al. a paper is anticipated in the near future. Here, we provide an annotated checklist of the Loblolly Unit. AND METHODS SITE The Big Thicket located in the West Gulf Coastal Plain in southeastern Texas Diggs 2006 is (see et al. for and Peacock literature description). (1994) reported 12 units for the Big Thicket National Preserve. Since then the Big Sandy Creek Corridor (1,798 hectares), the Canyonlands (350 hectares), and Village Creek Corridor (2,059 hectares) have been added. The addition of these 3 units has added 4,207 hectares to the 34,000 reported by Peacock in 1994. However, Deanna Boensch, Big Thicket ecologist comm.), fire (pers. now indicated the total hectares are 40,088. The units range from 222 to 10,000 hectares and are scattered over 7 Texas counties. The Loblolly Unit consists of 222 hectares in Liberty County (Fig. 1). It is located on the Beaumont formation on Aldine-Aris/Gyron-Aldine, Vamont, and Waller and topographically soils, is flat, and poorly drained with very slow permeability (Deshotels 1978; Aronow 1981; Shelby et 1992; al. Griffith 1996). Elevation about 30 meters. After heavy rain standing water present in the flatwoods is falls, is for extended periods of time. The Unit consists of basically plant community complex: Floodplain and/ 1 & Hardwood common or Flatland Marks Harcombe Forest, a association in the area (Ajilvsgi 1979; 1981; Bezanson 2000). 2006). However, others have questioned this interpretation maintaining that has always been Flatland al. it & Hardwood although may have been (Harcombe Marks Our Forest cleared in the past 1979). study of it the Loblolly Unit found no prairie species or any remnant prairie habitat. Therefore, the flora is clearly a & & Hardwood (Harcombe Marks Marks Harcombe Flatland Forest 1979; 1981). Like units of the Big Thicket National Preserve, the Loblolly Unit has not previously been the subject all of a detailed floristic inventory. known Because our purpose is to produce a list of taxa to occur on the Loblolly Unit, a vouchered A specimen was considered to be the only evidence acceptable for inclusion in the total of 471 herbarium list. specimens form the data for this report. All specimens have been annotated by Larry Brown. Brown Larry collected 277 specimens during 11 days from 5 August 1978 14 June 2008. The field to 2 8 1 Brown et al., Vascularflora of the Loblolly Unit, Big Thicket 1483 MacRoberts' collected 127 specimens during 5 field days from 30 July 2007 to 18 May 2008. Sandra Elsil* and Warren Pruess collected 35 specimens during 2007. Susan Walker 23 specimens on 22 collected Octobei Watson 2007. Geraldine collected 9 specimens over several years in the late 1970s and early 1980s. most In nomenclature cases, follows Jones Kartesz and Diggs et al. (1997); (1999); et (2006). In cases al. of multiple collections, no more than 4 are listed for each taxon. RESULTS The following is an annotated list of the Loblolly Unit taxa. Abbreviations used are given below. GW = Geraldine Watson. Her specimens TAES. are at LB = Larry Brown. His specimens with numbers TAES collection above 29,000 are and those few at with numbers below 29,000 SBSC. are at MM = Barbara and Michael MacRoberts. Their specimens TAES. are at SEWP = Sandra and Warren Elsik Pruess. Their specimens TAES. are at SH = Shawn Harper, grandson of LB. SW = Susan Walker. Her specimens TAES. are at MM & Hygrophila lacustris (Schlecht. Cham.) Nees, 32266 Arisaema triphyllum Schott, 8004 (L.) M Justicia ovata (Walter) Lindau var. lanceolata (Chapm.) R.W. A ™is*«» MM Long, LB 32200, 29281, 29298; 7930 MM Ruellia caroliniensis Gmel.) Steud, LB 29318; 82: (J.F. Arecaceae LBSH 33675 5abal minor (Jacq PerS LB 3241 5 ') Aceraceae " MM SEWP Aristolochiaceae 3044; 7906, 8094 .., MM Acmella oppositifolia (Lam.) Jansen, LB 32206; 7< MM Ambrosia trifida L, LB 32431; 7975 Centellaerecta(U) Fernal- MM 7998 tlimifolia L, Chaerophyllum Hook, tainturieri v Conoclinium coelestinum DC, LB 32269 (L.) SEWP 3044 \ MM Coreopsis tinctoha Nutt, LB 32261; 7926 MM Ecliptaprostrata L, 7934, 801 LB 5849 (L.) 0; *Cyclospermum leptophyllum Sprague (Pers.) MM Elephantopus carolinianus Raeusch., LB 32425; 7 MM Eiephantopus tomentosus LB 32414; 801 L, Cynosciadium digitatum DC, LB 32202 MM LB 32274, 3241 -Jifolia (L.) Raf., 3; £ Eryngium prostratum Nutt. ex DC, LB 32187 ^3m^ MM HHyyddrrooccoottyyllee vveerrttiicciilllaltaataTJhhuunnbb..,, LLBB 332211 9933,, 2299:323; 791 MM Small 7999 Limnosciadiumpumilum & & (Engelm. A. Gray) Mathias Cons- Eupatohum serotinum Michx., LB 32284 Euthamia leptocephala (Torr. & A. Gray) Greene ex LB 32290 Britton, Gamochaeta SEWP pensylvanicum 3030 Cabrera, (Willd.) Trachelospermum difforme (Walter) A. Gray LB 3241 Gamochaeta purpurea Cabrera, LBSH 33305, 33488 (L.) amarum •lenium Rock amarum, LB 32265; Mlv (Raf.) var. 7902 Chambers gia cespitosa LBSh (Raf.) K.L. var. cespitosa, 33292 33666 1 Pluchea camphorata DC, LB 32426; 8002 (L.) SW SEWP Cornus florida L, 3005; 45; LBSH 33672 1 SW Cornus foemina LB 29320, 29304; 38 Mill., 1 Pyrrhopappuspauciflorus (D. Don) DC, LB 29309a Nyssa biflora Walter, LB 32304; LBSH 33301 MM MM Solidago altissima L, 7976 Nyssa sylvatica Marsh., LB 29306; 8085; LBSH 3: Solidago rugosa Mill. var. asper (Aiton) Female 32293; Penthorum sedoides LB 32433, L, MM & Cayaponia quinqueloba Shinners, Symphyotrichum Love 8099 (Raf.) lateriflorum Love, 5 (L.) Symphyotrichum racemosum Nesom, 32270 LB (Elliott) MM 7908 Vernonia missurica Raf., 33504 H *Youngiajapomca DC, LBSH 33308 MM (L.) a Dewey, 8233 & CarexcomplanataTorn Hook., LB 32248 Carex carolinianaScbwe\r\., LBSH 33293 Podophyllum peltatum L. Carexcherokeensis Schwein., LBSH 33289 SEWP Carexcrus-corvi Shutlew. ex Kunze, LB 32189; 302: Carexdebilis Michx., LB 32246 Carexflaccosperma Dewey, LBSH 33487 MM Carexfrankii Kunth, LB 321 2931 7927, 8230 91 9; , Carex intumescens Rudge, LBSH 33485 MM Carexjoori L H. Bailey, LB 32277, 3724; 7996 MM a LB 29296, 29300, 3241 79< Bailey, 9; LBSH 33287 1 Bailey) Small, Carex triangularis Boeck., LBSH 33285, 33481 LB 29324 ; 32186,3 MM *Heliotropium indicum L, 81 00 Heliotropium procumbens LB 29312, 5852 Mill., Bockeler, LB s 3 a Engelm., LBSH 33297 I59 MM Cyperus erythrorhizos Muhl., 8005 b MM Cyperus ochraceusVahl, 7977 Cyperus pseudovegetus LB 32283, 3231 Steud., Cyperus virens Michx., LB 32247 SW usneoides 37 Tillandsia L, (L.) 1 LBSH 33668 //'Steud., 8022 1 Polypremum procumbens L, LB 32437 a (Kunth) Roem. & Schult, LB 32285 SEWP 32267; 3024 Campanulaceae i MM MM *Fimbristylis miliacea Vahl, 8089 (L.) Lobe/id cardinalis L, 8009; LB 14856 Isolepis carinata Hook. & Am. ex Torn, LBSH 33299 MM +Kyilingapumiia Michx., LB 32258, 7501a; 7987, 8080 1 Rhynchospora caduca LB 32286, 32307, 321 321 92 Elliott, 96, MM Rhynchospora corniculata (Lam.) A. Gray; LB 32245; 7922 Rhynchospora glomerata Vahl, LB 32301, 32317; LBSH (L.) 33678 Rhynchospora inexpansa (Michx.) Vahl, LB 32201; LBSH 33674 LB 29295; LBSH 33480 fa Britton, .,LB32212;SW132 32423 Scleria oligantha Michx., LB Scleria triglomerata Michx., LB 29286 pseudocaudatum (Clute) r. 7 5 5 1 Brown et al., Vascular flora of the Loblolly Unit, Big Thicket MM (Michx.) Schott, 8095; I 29284 LB LB 29294 arshall, Acalypha gracilens A. Gray, LB 32308 Walter, LB 3241 1,33132 is/s Juncus validus Coville, L MM 'Phyllanthus urinaria L, 791 MM *Triadica sebiferum Small, LB 32268; 7905 (L.) MM 09 ssin Durazz., 81 / 8000 /1 GW Centrosema virginianum (L) Benth., 21 56 GW MM Lycopus virginicus L, 2086; 8072 GW Desmodium glabellum (Michx.) DC., 215 Micromeria brownei Benth. (Sw.) LBSH 33494 SEWP LB32207; 3037 MM *Medicago 8228 lupulina L, 29308 Prunella vulgaris L, LB *Trifolium campestre Schreb., LBSH 33291 SEWP 3040 Salvia lyratal., LBSH 33283 'pens L, Scutellaria integrifolia L, LB 32287 iana Nutt. subsp. ludovidana, LBS! Stachys tenuifolia Willd., LB 32190, 32260; 33669 Quercusa/fc>aL.,MM8101 MM Quercus falcata Michx., 81 03, 8093 :t.)Nees,SEWP3014;MM7 Quercus laurifolia Michx., LB 32275a MM Quercus michauxii Nutt, LB 29283; 791 SEWP MM 3010 (L)Britton, Quercus nigra L, 8084, 8092; LB 32275 Quercus phelios LB 32250, 29290 L, SW Quercus shumardii Buckley, 35 32439 LB 1 MM SW +Quercus velutina Lam., 8083, 8090; 1 33677 Gelsemium SEWP on 301 f, Mitreola petiolata Gmel.) Torr. Gray, (J. F. \. L SW 7931; 155 Lygodiumjaponicum SEWP (Thunb. ex Murray) LB 2931 Sw., 6; Magnolia grandiflon Malvaceae SEWP Modiola caroliniana Don, G. 3 (L.) MM MM Sida rhombifolia L, LB 32294; 7970 Hypericum hypericoides Crantz, LB 293 791 (L.) 5; 1 MM Hypericum mutilum LB 32299; 8087 L, Triadenum walteri G. Gmel.) Gleason, LB 2929I 7978 (J. : MM MM Moms 8237 Rose, rubra 7903 L, // 33287 //(Baker) Brackett, Myricaceae Morella cerifera Small, (L.) 5 SEWP isulatum Bicknell, LB 29311; 3026; E.P. 2 8 Journal of the Botanical Research Institute of Texas 2(2) +*Echinochloa colonum (L.) Li *Eleusineindica Gaertn., IV (L.) Elymus virginicus L, LB 32203; GW & Eragrostis hypnoides (Lam.) B Sterns Poggenb., i, 2168 GW SW Leersia lenticularis Michx., 21 62; LB 32204; 58 1 Ludwigia d 32208 Willd, LB Ludwigia glandulosa LB 3221 Walter, Oplismenus hirtellus Beauv. subsp. setarius (Lam.) K (L.) P. Ludwigia LB 32438 hirtella Raf., GW MM Ekman, LBSH 33670 231 8067; 3; Panicum anceps Michx., LB 32300, 3231 LB 32194 Elliott, MM .) Panicum dichotomiflorum Michx., 8076 t,GW2399 MM Panicum gymnocarpon LB 5595, 32428; 792 Elliott, SEWP Oenothera speciosa Nutt, 3035 Panicum LBSH 33503 rigidulum Nees, MM *Paspalum dilatatum LB 32409; 7974 Poir., +Paspalum conjugatum LB 32303 Bergius, P.J. Paspalum laeve Michx. var. laeve, LB 32440 Paspalum notatum Flugge, LB 32273 Paspalum praecox Walter, LB 32302 Paspalum repens Bergius, LB 7502 P.J. 1 Paspalum setaceum Michx., LB 3231 Passifloraceae *Paspalum LBSH 33497 Steud., urvillei MM Passiflora incarnata L, LB 32298; 8098 MM Phalarisangusta Nees LBSH 33498; 8235 exTrin., LB 32253 Passiflora luteal., *Poa annua LBSH 33294 L, SW Saccharum giganteum 147 (Walter) Pers., GW 2324 *Sacciolepis indica Chase, (L.) Setaria parviflora Kerguelen, LB 32209 (Poir.) Sphenopholis obtusata (Michx.) Scribn., LBSH 33500 Platanus occidentalism 32420; 33 Polypodiaceae & Agrostis hyemalis (Walter) Britton, Sterns Poggenb., LBSH Andrews & Windham Pleopeltis polypodioides subsp. E.B. (L.) michauxiana (Weatherby) Andrews & Windham, E.B. Axonopus Kuhlmann, (Raddi) LB 32314, 32303a; fissifolius GW2177 Axonopus furcatus (Flugge) A. Hitchc, LB 32427 S. SEWP *Anagallis arvensis L, LB 29322; 3036 L *Bromusjaponicus Thunb. ex Murray, LB 32181 Samolus valerandi subsp. parviflorus (Raf.) H^ 293 1 3, GW Chasmanthium laxum 2314 32182;SEWP3018;MM7920 (L) H.O. Yates, Dichanthelium acuminatum Gould & (Sw.) C.A. Clark \ acuminatum, LB 29301a, 32320 Dichanthelium acuminatum Gould & (Sw.) C.A. Clark var. i & dheimeri (Nash) Gould C.A. Clark, LBSH 33489 Berchemia scandens 29321, 32322; commutatum comn Dichanthelium Gould subsp. (Schult.) ( tatum, LB 29287, 32422; LBSH 33679 Dicnanmenum aichotomum Gould (L) : MM MM LB 29301; 7988 Crataegus marshallii EggL, 8097 Dichanthelium dichotomum Gould (L) Crataegus opaca Hook. & Am. ex Hook., LB 32282, 29 & (Muhl. ex Freckmann Lelong, LB 32197, 32257, Elliott) 29307 32323; LBSH 33478 SW SEWP *Duchesnea indica (Andr.) Focke, 3033; 54 1 Dichanthelium laxiflorum (Lam.) Gould, LB 32435 SEWP Prunus serotina Ehrh. var serotina, 301 LBSH 33' 2; MM Dichanthelium ovale Gould &C.A. Clark subsp. SEWP ( Elliott) Rubus argutus Link, LB 32280, 3023, 3007; 79 simum Freckmann & (Nash) Lelong, LB 32321 SEWP Dichanthelium sphaerocarpon Gould 3020; (Elliott) , LBSH 33491 2 ; a of the Loblolly Unit, Big Thicket MM Diodia virginiana L, LB 32259; 791 SEWP *Galium aparine 3046 SW L, LBSH 29282; 33300; 3 Galium tinctorium L, LBSH 33304, 33306 Houstonia pusilia Schoepf., LBSH 33283a Mitchella repens L, LB 2931 (Desv.) Morton, LB 29281a, 32185; 7 I Ulmus americana LB 32249; SEWP 300 L, Bacopo monnieri Penn., LB 32262; (L.) IV SEWP Gratiola virginiana L, LB 31 938; 3' Leucospora multifida (Michx.) Nutt, LB MM Boehmeria cylindrica Sw., LB 5595; 7 MM (L.) 7969 Valerianaceae Valerianella radiata (L) Dufr., LBSH 33286 MM Mecardonia acuminata 7972 (Walter) Small, Verbenaceae MM Mecardonia procumbens SEWP Small, 3029; 791 MM (Mill.) 9, Callicarpa americana L LB 32281 7898 7971 MM Phyla nodiflora Greene, 7929 (L.) " Micranthemum umbrosum " 1 Gmel.) Blake, (J.F. S.F. LBSH 33667 I, MM SW Mimulus alatus Aiton, 7932; 143 MM Verbena scabra Vahl, 7981 Verbena urtia L, LB 32183 'folia a peregrin< SEWP (Green) McKinney, Smilaxglai EWP Smilax rotundifolia L, S 3008; 29297; SEWP Smilaxsmallii Morong, 3006 Parthenocissus quinquefolia LB 3231 Planch., (L.) 1; Physalis cinerascens (Dunal) Hitchcock A.S. \ LB 293 10a cinerea (Engelm.) Engelm. ex Millardet Vitis var. MM 32256; 7910 3 MM r olium Michx., LB 32263; 7893 Xyridaceae ambigua ex GV 32436 Xyris Bey. 2224; species is usually found in low moist areas which are frequent in this Brown Unit. See et al.(2007) for a discussion of this species in Texas. Kyllinga pumila. Kyllinga odorata is the most common Kyllinga in the Houston area (Harris and surrounding counties) but not present in this Unit. K. pumila is rare in the Houston area but common along C.R. 2071 that extends through the center of the Unit. Quercus woody velutina. This plant, as of now, has not been observed as a tree in the Houston area but grows as a small understory shade species. It is separated from the similar Q. jalcata by the longer erect branched on On hairs the abaxial surface of the shade leaves. Q. jalcata these trichomes are smaller and appressed [see the key in Smith (1994)]. Echinochloa colonum. Most books use Ward specific epithet. See (2005) for this change in ; orthography. Paspalum conjugatum. Turner at (2003) mapped this species in Cameron County in extreme south Texas al. with record from Hardin County Pineywoods 1 in the region of east Texas. Diggs et al. (2006) consider the Hardin County specimen to be from a plant cultivated as a lawn grass. For now, the only east Texas collec- tions of native plants are from Liberty County in the Trinity River National Refuge and the Loblolly Unit. The plants were frequent along the southern edge west from the junction with C.R. 2071. Additional specimens SBSC at are from the various units of the National Wildlife Refuge complex in Brazoria County. DISCUSSION There are 88 families and 201 genera for the 305 taxa (271 native) on the Loblolly Unit Eight families, list. Poaceae Cyperaceae Asteraceae Scrophulariaceae Apiaceae (42 taxa), (36 taxa), (31 taxa), (11 taxa), (10 taxa), % Lamiaceae (9 taxa), Fagaceae (8 taxa), and Fabaceae (7 taxa) account for 50 of all taxa collected. We compared the present list of species for Loblolly Unit with the species reported by Turner et al. % (2003) in their Atlas the Vascular Plants Texas and found that 115 taxa, 37.7 of the species on the of of County Loblolly Unit are not reported in Liberty in that source. This result not surprising considering list, is that, until recently, focused collecting has been undertaken in the Big Thicket region. little we we While the case that our incomplete, as are believe that have collected prob- it is list is all floras, % We ably 90+ of the taxa. estimate that the Loblolly Unit has about 281 to 290 native species/taxa. Of the 7 Units inventoried so far, the following taxa are reported only for the Loblolly Unit: Hymenocallis liriosme, Cynosciadium Rorippa Commelina Ipomoea Hy- digitatum, lacunosa, Eleocharis engelmannii, sessiliflora, diffusa, drolea uniflora, Hypoxis wrightii, Micromeria brownei, Ludwigia sphaerocarpa, Paspalum conjugatum, Ranunculus new and Only Thus muricatus, Physalis cinerascens var. cinerascens, Verbena 14 taxa are to this Unit. urticifolia. 95% of the taxa in the Loblolly Unit had already been collected in other units of the Big Thicket National Preserve, which is to be expected since most of the other units possess Floodplain and/or Flatland Hardwood communities. forest ACKNOWLEDGMENTS This study was supported in part by an All Taxa Biological Inventory (ATBI) grant from the Big Thicket Asso- We ciation of Saratoga, Texas. thank Paul Roling, liverwort and moss student of the Big Thicket; and Deanna new Boensch, Big Thicket fire ecologist, for providing information regarding the Big Thicket Units. Also thanks to Ray C. Telfair and Eric Keith for their careful reading of the paper and for the improvements. II REFERENCES A&M G. 979. Wild flowers of the Big Thicket, east Texas, and western Louisiana. Texas Univ. Press, Col- Ajilvsgi, 1 lege Statior on Aronow, 1981. Notes the geologic units: Big Thicket National Preserve. Big Thicket National Preserve, S. Beaumont, Texas. Bezanson, D. 2000. Natural vegetation types of Texas and their representation conservation areas. M.A. Thesis, in Univ. ofTexas, Austin. Brown, L.E., B.R. MacRoberts, M.H. MacRoberts, PA. Harcombe, W.W. Pruess, I.S. Elsik, and D. Johnson. 2005. Annotated checklist of the vascular flora of the Turkey Creek Unit of the Big Thicket National Preserve, Tyler and Hardin 21:1807-1827. counties, Texas. Sida Brown, MacRoberts, M.H. MacRoberts, PA. Harcombe, W.W. and 2006a. Annotated L.E., B.R. Pruess, I.S. Elsik, S.D. Jones. checklist of the vascular flora of the Big Sandy Creek Unit of the Big Thicket National Preserve, Polk County, 22:705-723. Texas. Sida Brown, MacRoberts, M.H. MacRoberts, PA. Harcombe, W.W. and 2006b. Annotated L.E., B.R. Pruess, I.S. Elsik, S.D. Jones. checklist of the vascular flora of the Lance Rosier Unit of the Big Thicket National Preserve, Hardin County, 175-1 Texas. Sida 22:1 189. Brown, L.E., E.L Keith, DJ. Rosen, and Liggio. 2007. Notes on the flora ofTexas with additions and other significant J. records Texas 1:1255-1264. Bot. Res. Inst. J. III. Brown, MacRoberts, M.H. MacRoberts, PA. Harcombe, W.W. and Walker 2008. Annotated L.E., B.R. Pruess, I.S. Elsik, S.B. checklist of the vascularflora of the Beech Creek of the Big Thicket National Preserve, Tyler County, Texas. Texas 2:651-660. Bot. Res. Inst. J. Brown et al., Vascular flora of the Loblolly Unit, Big Thicket 1489 Deshotels, J.D. 978. Soil survey for the Big Thicket National Preserve, exas. USDA Soil Conservation Service and 1 Texas Experiment Agricultural Station, College Station, Texas. Diggs, G.M., B.L. Lipscomb, M.D. Reed, and RJ. O'Kennon. 2006. Illustrated flora of east Texas. Sida, Bot. Misc. 26:1- 1594. USDA Griffith, K.L 996. Soil survey of Liberty County, Texas. and Texas Agriculture Experiment College 1 Station, Station, Texas. PA PL Harcombe, and Marks. 1979. Forest vegetation of the Big Thicket National Preserve. Unpublished. Rept. New Southwestern Region: National Park Service, Santa Mexico. Fe, Jones, S.D, J.K. Wipff, and P.M. Montgomery. 997. Vascular plants of Texas. Univ. of Texas Press, Austin, Texas. 1 A synonymized and Kartesz, J.T. 999. checklist atlas with biological attributes for the vascularflora of the United 1 Canada, and Greenland. and Meacham. States, 1st ed. JT. Kartesz C.A. Synthesis of North American In: flora. Version North Carolina Botanical Garden, Chapel 1.0. Hill. MacRoberts, B.R., M.H. MacRoberts, and L.E. Brown. 2002. Annotated checklist of the vascular flora of the Hickory Creek Unit of the Big Thicket National Preserve, Tyler County, Texas. Sida 20:781-795. PA Marks, RL. and Harcombe. 981 Forest vegetation of the Big Thicket, southeast Texas. Ecol. Monogr. 51:287- 1 . 305. A&M Peacock, H.H. 1994. Nature lover's guide to the Big Thicket. Texas Univ. Press, College Station, Texas. M. McGowen, Aronow, McGowen, Shelby, C.A., K. S. W.L. Fisher, LF. Brown, J.H. C.G. Groat, and V.E. Barnes. 992. Geologic 1 Beaumont atlas of Texas: Sheet Map. Bureau of Economic Geology, Austin, Texas. Smith, E.B. 1994. Keys to the flora of Arkansas. The Univ. of Arkansas Press. Fayetteville. Turner, B.L, H. Nichols, G. Denny, and O. Doron. 2003. Atlas of the vascular plants of Texas. Sida, Bot. Misc. 24: A Ward, D.B. 2005. case of disputed orthography: Echinochloa colona; or Echinochloa colonum (Gramineae). is it is it Sida 21:21 71 -21 83. Watson, G.E. 1979. Big Thicket plant ecology, an introduction. 2nd ed. Big Thicket Museum, Saratoga, Texas.

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