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CaribbeanJournalofScience,Vol.41,No.3,374-391,2005 Copyright2005CollegeofArtsandSciences UniversityofPuertoRico,Mayagu¨ez An Introduction to the Geology of the Bocas del ToroArchipelago,Panama A.G.COATES1, D.F.MCNEILL2, M-P.AUBRY3, W.A.BERGGREN4, L.S.COLLINS5 1SmithsonianTropicalResearchInstitute,Apartado2072,RepublicofPanama 2DivisionofMarineGeologyandGeophysics,RosensteilSchoolofMarineandAtmosphericScience,Universityof Miami,Florida,33149,USA 3DepartmentofGeologicalSciences,RutgersUniversity,WrightLabs,610TaylorRoad, Piscataway,NewJersey,08854-8066 4WoodsHoleOceanographicInstitute,DepartmentofGeologyandGeophysics,WoodsHole,Massachusetts,02543, USAandDepartmentofGeologicalSciences,RutgersUniversity,WrightLabs,610TaylorRoad,Piscataway,New Jersey,08854-8066,USA 5DepartmentofEarthSciences,FloridaInternationalUniversity,Miami,Florida,33199,USA Correspondingauthor:[email protected] ABSTRACT.—WereviewthestratigraphyoftheNeogenerocksoftheBocasdelToroarchipelago,western CaribbeancoastofPanama,andprovidenewgeologicalmapsandapreliminarydescriptionofnewNeogene formationsontheislandsofBastimentosandColon.ThePuntaAlegreandValienteformationsrangeinage from19to12Ma.Afterahiatusfrom12to8Ma,atransgressive/regressivesedimentarycycleisrecordedby theTobobe,NancyPoint,SharkHole,CayoAgua,andEscudodeVeraguasformations(=theBocasdelToro Group),thatrangeinagefrom7.2-5.3to1.8Ma.Incontrast,inthenorthernregion,theOldBank,LaGruta, andGroundCreekunitsandtheSwanCayFormationonlyrangeinagefromabout3.5to0.78Ma.Thehiatus represented by the unconformity is from 12 to 3.5 Ma. We integrate the geology of the Bocas del Toro archipelagointoabriefhistoryoftheNeogeneofthelowerCentralAmericanisthmus. KEYWORDS.—Neogene,BocasdelToro,CentralAmericanIsthmus INTRODUCTION framework for the evolutionary and eco- logical studies of the macrobiota. Background This overview paper provides an intro- Regional Geologic History duction to the geological foundation of the modern biological systems found today in TheCentralAmericanisthmusformsthe theBocasdelTororegion.Thestudyofthe westernmarginoftheCaribbeanPlateand geology of Bocas del Toro was undertaken lies at the center of a complex intersection aspartofthePanamaPaleontologyProject of the Pacific Cocos and Nazca plates with (PPP; Collins and Coates 1999), a collabo- the Caribbean Plate and the small Panama rativeresearchprogramtostudysediments Microplate(Fig.1).Thedominantlyoceanic deposited during the last 20 million years CaribbeanPlateliesbetweentheNorthand (involvingtheMiocene,Pliocene,andPleis- South American plates. Its relative east- tocene Epochs, which are together known wardmotion,withrespecttotheNorthand as the Neogene) along both sides of the South American plates is accommodated isthmus of Central America and of north- bystrike-slipfaultstothenorthandinpart ern South America (Fig. 1). tothesouth(nowconfoundedbycompres- Wesystematicallymappedthewholere- sionfromthewest-northwestward-moving gion, named the physical stratigraphic for- South American Plate). In the east, it is mations (Fig. 2) and located the rich and boundedbytheLesserAntillessubduction diverse macrofossil sites. After additional zone.ThewesternmarginoftheCaribbean paleomagneticstudiesandradiometricdat- Plateismorecomplex;inthenorthernpart, ing, these studies provide a detailed geo- the Cocos Plate is in contact with the Car- logic history of the region and a temporal ibbean Plate. 374 GEOLOGYOFBOCASDELTOROARCHIPELAGO 375 FIG.1. MapofsouthernCentralAmerica(darkshading)andthePanamamicroplate(paleshading).Dashed lineswithteethmarkzonesofconvergence;zipperedlineisPanama-Colombiasuture.Veryheavydashedline markslocationofNeogenevolcanicarc.FinearrowsarePaleogenefaults;thickarrowsarelateNeogenefaults. PrincipalNeogenesedimentarybasinslocatedbystripedovals.SpottedpatterndefinestheCocosRidge.Arrows ontheinsetindicatedirectionsofrelativemotionsoftheplates.PFZ=PanamaFractureZone In the southern portion of the western andisassociatedwithactiveseismicityand margin, a triple junction brings the Cocos volcanism. Northern Central America also and Nazca Plates in contact with the small hasabroadzoneofoldercontinentalcrust, Panama Microplate (Fig. 1). The Panama and has a geologic history extending back Microplate appears to have formed by totheearlyPaleozoic(Donnellyetal.1990). northwardescapefromcompressionofthe In contrast the Nazca Plate has a relative South American and Cocos/Nazca plates easterlymotionbutisnotactivelysubduct- (BurkeandSengor1986;Coatesetal.2004). ingunderPanama.ThePanamaMicroplate Much of Central America lies either on the is formed of oceanic crust that is typical of trailing western edge of the Caribbean the widespread basalt plateau that under- Plate or on the Panama Microplate but a liesmuchoftherestoftheCaribbeanPlate portion also lies on the southwestern cor- (Case et al. 1990). ner of the North American Plate (Fig. 1). Three major movements have affected SincetheirformationintheMiocene,the the Neogene tectonic evolution of the Nazca and Cocos plates have impinged on southern Central American isthmus (CAI; Central America with different motions. Kolarsky et al. 1995; Coates and Obando The Cocos Plate (Fig. 1), with relative 1996). The first is convergent tectonics of northeasterly motion, is subducting vigor- theeasternPacificsubductionzone(Fig.1), ously under northern Central America as the primary driving force that created the far south as the Costa Rica-Panama border Central American isthmus by forming a 376 A.COATESETAL. western South America (Coates et al. 2004; Silveretal.1990;MannandKolarsky1995). The relative northwestward movement of South America (Fig. 1) has increasingly compressed the southern Caribbean Plate margin in the late Neogene. This conver- gencehasupliftedeasternPanamaandthe northern Andes of Colombia and Venezu- ela. The resulting shoaling of the CAI (de- tected paleoceanographically by a marked divergence between planktonic foraminif- eraloxygenisotoperecords(aproxyforsa- linitychanges)wastolessthan100mfrom 4.7-4.2Ma(Keigwin1982;HaugandTiede- mann 1998; Gussone et al. 2004). This initi- atedthedevelopmentofthemodernAtlan- tic-Pacific contrast in sea surface salinities (SSS), with higher SSS in the Caribbean. RecentstudiesbyTiedemann,Steph,and Groenveld and others (Poster sessions, American Geophysical Union Meeting, 2004andpers.comm.2005)havesuggested 2.8 as the time of final closure of the CAI. FIG. 2. Physical stratigraphic nomenclature of the UsingCa/Mgmeasurementsofplanktonic NorthernandSouthernregionsoftheBocasdelToro foraminifera,theyshowthatCaribbeanand archipelagowithagesinmillionsofyears Pacific Sea Surface Temperature (SST) rec- ordsaresimilarfrom5-2.8Ma.Thenlarger volcanic arc extending southwards from scale sea level fluctuations with 41 kyr. cy- North America. It forms part of an exten- clicity became dominant in response toam- sive zone of subduction that runs the plificationoftheNorthernHemispheregla- length of the western margin of North, ciation. After 2.8 Ma, glacial stages, when Central, and South America (Astorga et al. sea level is lowest, show minimum Pacific 1991). SSTs, but maximum SSTs occur in the Car- The second tectonic effect was subduc- ibbean. This anomaly is explained by low tion of the Cocos Ridge (Kolarsky et al. seastandspreventingcoolerlesssalinewa- 1995;Collinsetal.1995),alighterandrela- ter from entering the Caribbean from the tively thick welt of Pacific oceanic crust, Pacific (the CAI is emergent), and the re- undertheCentralAmericanvolcanicarcin verseappliesininterglacialswhensealevel Costa Rica (Fig. 1). This hard-to-subduct is high, breaching the CAI. ridge rapidly elevated the Central Ameri- The closure of the Isthmus of Panama can Isthmus from the Arenal volcano in triggeredprofoundenvironmentalchanges CostaRicatoElValleinPanama(deBoeret on land and in the sea. The formation of a al. 1988) culminating in the center where bridge between the North and South theTalamancaRangerisestoabout4000m. American continents gave rise to the Great TheelevationoftheTalamancarangeprob- American Biotic Interchange on land ably substantially reduced the number of (Webb and Rancy 1996; Webb 1999). Less marineconnectionsbetweenthePacificand well known are the timing and nature of Caribbean. thechangesinthesea,consequentuponthe The third tectonic influence on southern growth of the Central American volcanic Central America was the collision of the arc and its eventual collision with South CentralAmericanvolcanicarc(thewestern America.Thiscreatedamarinebarrierthat margin of the Caribbean plate) with north- increasinglyaffectedoceancirculationdur- GEOLOGYOFBOCASDELTOROARCHIPELAGO 377 ingNeogenetime.Intheprocess,astriking inthesetwoareas(Fig.2).Wethenattempt contrast evolved between the relatively to synthesize the geologic history of Bocas warm, more saline, nutrient-poor Carib- del Toro and compare it with adjacent re- beanandthemoreseasonal,lesssalineand gions in Costa Rica, Darien, Panama, and morepelagicallyproductiveeasternPacific the Atrato Valley, Colombia. (Jackson and D’Croz 1999). These environ- mental changes apparently altered the 1) The northern region comprises Swan course of evolution, first of the deep-water Cay,Colon,Pastora,SanCristobal,Cari- planktonic organisms like radiolaria and nero, and Bastimentos islands, and the diatoms from about 15 Ma, and then suc- Zapatillo Cays. cessively shallower taxa until complete 2) The Southern region comprises the is- emergence at about 2.8 Ma. By this time, landsofPopa,Deer,CayoAgua,andEs- theeasternPacifichadevolvedrichpelagic cudodeVeraguas,andtheValientePen- biotasbutpoorlydeveloped,lowdiversity, insula. coral reefs (Jackson and D’Croz 1999). In Ingeneral,thenorthernregionexhibitsa contrast, the Caribbean had become domi- latePliocene-Pleistocenesuccessionofshal- nated by coral reef-seagrass-mangrove low water sediments, especially coral reef coastal ecosystems (Collins et al. 1996) and deposits,thateitherunconformablyoverlie a profound taxonomic turnover had oc- middle Miocene volcanic arc basalt (Basti- curred in corals and mollusks (Budd and mentosIsland)orrestonathick(>2500m) Johnson 1997, 1999; Jackson et al. 1993). siliciclastic shale (Colon Island) that is late Much of this history is documented in Pliocene at its top and of unknown age at the long geological section preserved its base. This suggests that there is a major around the Bocas del Toro Archipelago structural break in the volcanic arc base- whosedepositsrangeinagefromtheearly ment rocks between Bastimentos and Co- Miocene (about 20 Ma). The sequence can lon islands. The southern region reveals a beassignedtofourphasesintheriseofthe more extensive volcanic arc suite of lower isthmus as follows: 1) Deposition of lower and middle Miocene rocks, including a se- bathyal,pre-isthmian,oceanicsedimentsin quenceofdeep-seaooze,basalt,coarsevol- early Miocene time (21.5-18.5 Ma); 2) canicsedimentsandsmallscalepatchreefs, Growth of a volcanic arc during middle Miocenetime(∼18to∼12Ma)characterized that tracks the onset and rise of an active volcanic arc in the Bocas del Toro region. by terrestrial and marine deposits includ- Subequently, an unconformably overlying, ing columnar basalt and flow breccia, non-reefal, upper Miocene and Pliocene, coarse volcanic sediments, and scattered marine, transgressive/regressive shelf se- small coral reefs; 3) Extinction of the arc quence was deposited. about 12 Ma and subsequent extensive emergence and erosion; and 4) Subsidence of the volcanic arc during the latest part of Stratigraphic definitions Miocene time (∼7.2-5.3 Ma), resulting in a marinetransgression,followedbyamarine Stratigraphy of sedimentary rocks in- regression, that culminated in the wide- volves the integration of four main kinds spread development of Plio—Pleistocene of units, each of which provides the basis reefs (McNeill et al. 2000; Coates et al. of a subdiscipline of stratigraphy. The 2003). first,lithostratigraphy,describesthephysi- cal stratigraphic units and their three- dimensional geometry across the region STRATIGRAPHY studied.EachmappableunitorFormationis For the purposes of this paper, we first distinguished by its rock type or lithology describe the stratigraphic units that occur andisnamed,usuallyfromtheplacewhere inthesouthandnorthoftheBocasdelToro it most typically can be observed. A series Archipelago. These are discussed sepa- of formations deposited in vertical succes- rately because the sequences are different sion in a genetically related depositional 378 A.COATESETAL. episode may be lumped together as a (i.e., Milankovich periodicity), usually in Group. conjunction with paleomagnetic stratigra- If the contact of two lithostratigraphic phy, to determine the ages of chronostrati- units shows evidence of a break in sedi- graphic and biostratigraphic boundaries. mentation this line is called an unconfor- i.e. the stratigraphic level common to two mity. Unconformities usually reflect major successive stages or biozones. Numerical geologicalevents(liketectonicupliftorsea ages vary depending on the criteria se- levelchange)intheareawheretheyoccur. lected in the building of a gechronologic The significance of unconformities will frameworkortimescale.Theagesgivenbe- vary depending on how different the units lowrefertothetimescaleofBerggrenetal. are above and below, and how much time (1995). The boundaries between lithostrati- thebreakrepresents.Fromthenatureofthe graphic units may be of the same age re- different sediments and their associated gionally, but over larger regions they may sedimentary structures and fossils, the become younger or older in age as the physical environment and water depth in physical environments they represent mi- which the formation was deposited can be grated across the sedimentary basin reconstructed.Fig.2outlinesthenomencla- through time. They are then said to be di- tureofthelithostratigraphyoftheNeogene achronous. formations of the Bocas del Toro archi- pelago. The southern region The second subdiscipline is biostratigra- phy, which takes advantage of the fossil content of rocks, and is based on the range Eachlithostratigraphicunit,startingwith ofspecies,i.e.,thestratigraphicintervalbe- the lowest, will first be named and de- tween two specific stratigraphic horizons scribed, and then evidence for its age and corresponding to the lowest (LO) and the environment of deposition will be added. highest(HO)occurrenceofataxon.Theba- For a more detailed description of the sic unit of biostratigraphy is the biozone, a stratigraphy of these units see Coates et al. stratigraphic interval defined between LO (1992, 2003) and Coates (1999). and HO of taxa selected for their distinct Punta Alegre Formation.—This formation morphology and vast geographic distribu- is named for Punta Alegre, the nearest tion.Biozonesallowstratigraphiccorrelation, smallvillagetoitssolitaryexposure.Punta the fundamental procedure of establishing Alegre is located on the western tip of the that two horizons have the same relative northcoastofBahiaAzul(BluefieldBay)in age. Chronostratigraphy, the third element the northern part of the Valiente Peninsula ofstratigraphy,isasystemofreferencefor (Fig.3).Theformationcanbeobservedina the relative dating of sediments. Its basic prominent small cliff about 1.3 km north- unit is the stage, defined by its base (= a westofthevillage,onthewest-facingcoast stratigraphic horizon) and a stratotype (a 1 km south of Valiente Point. It consists of stratigraphic interval). Correlation of rocks clay and silty ooze containing abundant inagivenbasintoastageisestablishedvia calcareous nannofossils and benthic and biostratigraphy. Stages are grouped in a planktonic foraminifera (see Coates et al. nested hierarchy of Series, Systems, and Er- 2003,Tables1,2),manyofthelatterareeas- athems. Their temporal equivalents are ily seen by a hand lens and by the naked called Ages, Epochs, Periods, and Eras, respec- eye. The formation can be seen to lie un- tively. derneath weathered coarse volcanic flow The fourth subdivision, geochronology, breccia(thebrokenupboulderyrockthatis is the science of numerical time. Whereas crated when lava flows and cools at the chronostratigraphydealswithrelativetime same time). From its abundant planktonic (younger/older), geochronology trans- foraminifera and calcareous nannofossils, forms specific stratigraphic horizons into the age of the Punta Alegre Formation is numerical markers. Geochronology uses 19-18.3 Ma. (Coates et al. 2003), and by its radioisotopic ages and/or astrocyclicity benthic foraminifera it is interpreted to GEOLOGYOFBOCASDELTOROARCHIPELAGO 379 have been deposited in water depths of flows and coral reef lenses. The latter con- 100—2000 m (Coates et al. 2003). sist of several species of Montastraea (in- The Valiente Formation.—This unit over- cludingM.imperatorisandM.canalis),mas- lies the Punta Alegre Formation and crops sive Porites waylandi and thin branching out extensively on the Valiente Peninsula Stylophora(possiblyS.granulate;AnnBudd (Fig. 3) and nearby Deer and Popa islands pers. comm. 2005). These facies intercalate (Fig. 4). The Valiente Formation is of com- andreplaceeachotheroververyshortdis- plex and highly variable lithology because tances,bothlaterallyandvertically(forde- itrepresentsthesuiteoffacies,marineand tailed descriptions of the facies, see Coates terrestrial,igneousandsedimentary,thatis etal.2003).Fig.3showsthatthebasaltlava associated with an active emergent volca- and flow breccia facies form a core around nic island arc. It includes columnar basalt, which the other facies are distributed pe- basalt flow breccia, pyroclastic tuff (air- ripherally as terrestrial, coastal or marine borne volcanic ash deposits), fluviatile/ slope deposits. Planktonic microbiotas and estuarine conglomerate, marine debris radiometrically dated basalt in the se- FIG.3. Geologicalmapandcrosssection(a–á)oftheValientePeninsula,BocasdelToro,westernPanama, showingthedistributionofthePuntaAlegreandValienteformationsandtheBocasdelToroGroup.Thefive lithofaciesoftheValienteFormationareindicatedbyseparatecolorsandnumbersonthekey(upperright)as follows;v1)basaltlavaandflowbrecciafacies;v2)coarsevolcaniclasticfacies;v3)pyroclasticfacies;v4)coral reeffacies;v5)marinedebrisflowandturbiditefacies. Fi 3 li 4/C 380 A.COATESETAL. pelago above the Valiente Formation, sig- nifyingextensiveemergenceanderosionof thevolcanicarcfrom∼11.5-∼7.2Ma(Fig.2, 3). This unconformity can be clearly ob- served in the inner islands of the Plantain Cays and along the coast immediately to the west. The underlying columnar basalt of the Valiente Formation can be seen on the north side of the Cays and the fossilif- erous conglomerate and sandstone of the Tobobe Sandstone on the south side. TheBocasdelToroGroupsignifiesanew depositional cycle as a marine transgres- sion submerged the older eroded volcanic arcrocks.Theoldestsedimentsthusrepre- sent the first shallow-water, beach, and near-shoresanddeposits.Asthetransgres- sion developed, subsequent overlying ma- rine units reflect deeper water deposition until the reverse occurred and the sea re- gressed to shallow water again. Thus, the units of the Bocas del Toro Group are ge- FIG.4. GeologicalmapofPopaIsland.Section(b-b’) neticallyrelatedanddocumentasinglema- showstheValienteFormationonPopaIsland,where it is unconformably overlain by the Pliocene Cayo rine transgressive/regressive event. The Agua Formation (ca). Numbers and symbols as for most continuous section lies along the east Figure3.OnPopaIsland,onlythev1basaltflowfacies and west coasts of the Valiente Peninsula andthev2coarsevolcaniclasticfacies(notassociated (Fig. 3) where it ranges in age from late withreeflenses)arepresent,withthinlayersoflow Miocene (Messinian, 7.2-5.3 Ma) to late rankcoal,anexampleofwhichisexposedalongthe Pliocene (∼3.5 Ma). coastimmediatelynorthofPuntaLaurel.Aprominent basalt dike is exposed at the tip of the Punta Laurel Tobobe Sandstone.—This lowest unit is a (hereshowninredasv1facies)whereitcutstheVa- pebble conglomerate that passes up into lienteFormation. clean, hard quartz sandstone containing sand dollars, spatangoid echinoids, mol- lusks, including the large, thin-shelled bi- quenceindicatethattheValienteFormation valve Amusium, vermetids and serpulids, ranges in age from 16.5-11.5 million years as well as an array of infaunal burrow old and includes marine deposits from structures. It is best observed on the small near-shore to as deep as 1000 m, as well as Plantain Cay and an adjacent smaller un- severaltypesofterrestrialdeposits(Coates named island to the west (Fig. 3). The de- et al. 2003). posit represents a beach and near-shore Bocas del Toro Group.—Columnar basalt sandbodyanddocumentstheearlieststage and pyroclastic and fluviatile rocks of the of the marine transgression. Planktonic Valiente Formation required extensive foraminifera from laterally equivalent de- emergenceofpartsofthegrowingvolcanic positsonToroCay(Fig.3)indicatethatthe arc(columnarbasaltisformedbythecool- age of the unit is Messinian (7.2-5.3 Ma). ingoflavapondedlakes).About12Ma,the Here the deposit contains a variety of mol- archadbecomeinactiveandrecentstudies lusks including turritellids and pectens, inDarien,Panama(Coatesetal.2004),sug- erect bryozoa, and echinoids. Classic ex- gestthatthecollisionofthesoutherntipof amples of complex thalassinoid burrow the Central American volcanic arc with systems constructed by callianasid crusta- South America was initiated at the same ceans are also present. time. These events may explain the major Nancy Point Formation.—It was named unconformity in the Bocas del Toro Archi- for the promontory called Nancy Point Fi 4 li 4/C GEOLOGYOFBOCASDELTOROARCHIPELAGO 381 which lies 2.5 km southwest of the village only along the southern coast of the Va- of Tobobe (Fig. 3). The formation is almost liente Peninsula, east of Secretario. The continuouslyexposedsouthwardalongthe Shark Hole Formation consists of mica- coast as far as Chong Point (= Nispero ceous, clayey siltstone that is pervasively Point). The Nancy Point Formation is con- bioturbated and rich in large scaphopods. formablewiththeTobobeFormationbelow The uppermost part of the formation also andtheSharkHolePointFormationabove containsabundant,thin,shellybedsandin- (Fig. 2 and 3). It consists of shelly, muddy traformational slumps with pillow folds and silty sandstone, and muddy siltstone and rip-up clasts. Benthic foraminifera in- with occasional coarse volcaniclastic and dicatethatthepaleobathymetryofthisunit bioclastic sandstone beds. It contains scat- ranges from 100-200 m (Collins 1993). This tered mollusks and occasional leaves and represents the first stage of the regressive plant fragments with diverse moderately phase of the Bocas del Toro Group trans- abundant molluscan assemblages through- gressive/regressive cycle. Calcareous nan- out the section and several, rich, low- nofossils and planktonic foraminifera are diversity,thick-shelledmolluskbedsatthe abundant in several horizons and suggest base. the age of the formation is early Pliocene ThetransitionfromTobobeSandstoneto (5.3-3.6 Ma). NancyPointFormationisbestseenonToro Escudo de Veraguas Formation.—The Cay (Fig. 3) where dark blue-gray, silty stratigraphic order of the formations de- sandstone, typical of the Nancy Point For- scribed above has been determined by mation,containsoccasional,clearlydefined physical superposition. The three remain- infaunal burrow systems and extremely ingformationsoftheBocasdelTorogroup abundant and diverse mollusks. It overlies are known only on islands and their posi- coarse, channeled Tobobe Sandstone with tionrelativetotheotherunitshasbeende- only a 10-m exposure gap. Nancy Point terminedthroughstratigraphiccorrelation. Formation benthic foraminifera indicate The Escudo de Veraguas Formation is that deposition of the Nancy Point Forma- known only from the island of the same tionwasin200-500m.waterdepth(Collins name that lies 27 km east of Nancy Point 1993) demonstrating that relatively rapid (Fig.5).Itsstratotype,forthelowerpartof deepening took place from near-shore (To- theformation,isalongthecoastontheeast bobe Sandstone) to upper slope (Nancy side of the V-shaped embayment situated PointFormation).Thisrepresentsthemaxi- inthecentralpartofthenorthcoast,about mumdepthofthetransgressive/regressive 1 km east of Long Bay Point (Fig. 5). The cycle. Planktonic foraminifera and calcare- stratotype for the upper part of the forma- ousnannofossilsdatetheNancyPointFor- tionliesalongthewestcoastforaboutone mationasMessinian(7.2-5.3Ma),thesame km south of Long Bay Point. time range as for the Tobobe Sandstone. Lithologically, the Escudo Formation Within that time frame the Nancy Point consists of, in the upper part (1.8 Ma), per- Formation is younger because it overlies vasively bioturbated clayey siltstone and the Tobobe Sandstone. silty claystone, with frequent concretions, Shark Hole Point Formation.—It is named and scattered shelly hash, often with scat- for the promontory of the same name that tered whole and diverse mollusks, small, lies 3 km east of Chong Point (Fig. 3) and cornute,ahermatypiccorals.Thelowerpart the stratotype lies along the coast between oftheformation(3.5Ma)ismoreindurated ChongPointandBrunoBluff,southofOld withverycommonanddenselypackedce- Bess Point. The Shark Hole Point Forma- mented burrow concretions and thalassi- tion conformably overlies the Nancy Point noid galleries at the base, scattered aher- Formation(Fig.2and3)andisoverlainby matypic corals through the middle part, an unnamed conglomeratic, cross-bedded, aswellasvariablyabundantmollusks,and coarser grained sequence of volcaniclastics at the top a coral biostrome dominated by containing large pieces of wood and plant Stylophora and sand dollars. The top and fragments. This unnamed unit is exposed botton units suggest deposition in inner 382 A.COATESETAL. mon in the middle of the formation. Com- pared to the Shark Hole and Escudo de Veraguas formations, the Cayo Agua For- mation is consistently coarser-grained, with common basalt grains and granules, phosphatic pebbles, and wood fragments. A distinctive marker bed of corals occurs near the top of the formation and is well exposedatTiburonPointandtheunnamed point to the south. The corals appear to be mostly free living, hermatypic, grass flat corals,suchasPlacocyathusandManicina(= Teleiophyllia), as well as ahermatypic spe- cies, and they suggest deposition in water depthsof30-50m(AnnBudd,pers.comm. FIG. 5. Geological map of Escudo de Veraguas is- 2005). Evidence from benthic foraminifera land. Ages in millions of years in brackets for lower (Collins1993)indicatespaleobathymetryof and upper parts of the Escudo de Veraguas Forma- tion. 20-80 m, overlapping the inference from li- thologyandcoralfaunathattheCayoAgua Formation represents a shallow-water fa- neriticwaterdepths,butthemiddlepartof cies. thislowersuccessionhasabundantbenthic The age of the Cayo Agua Formation is foraminifera indicating deposition in outer datedatthebase∼5.0-3.5Maandatthetop neritictoupperbathyalwaterdepths(Col- 3.7-3.4 Ma, which suggests it is a contem- lins 1993). porary, shallow water equivalent of the Cayo Agua Formation.—This formation SharkHolePointFormationandthelower- was named for the island that lies about 6 most part of the Escudo de Veraguas For- kmtothewestofToroPoint,ValientePen- mation. insula.Theformationiswellexposedalong theeastcoast(Fig.6)andconsistslithologi- The northern region cally of pervasively bioturbated, muddy, The islands in the northern region of the silty sandstone with common horizons of BocasdelToroarchipelagohaveadifferent abundant thick-shelled mollusks and aher- geologic history than the southern region. matypic corals. Occasional horizons of Volcanic arc columnar basalts of the Va- pebble conglomerate and very coarse- liente Formation form the basement that grained volcaniclastic sandstone are com- underlies the marine late Neogene succes- sion, as in the southern region. While no radiometric dates for these basalts are available, we can presume they are part of thesamevolcanicarcthatwentextinctand cooled after the middle Miocene (∼12 Ma). In all the Northern Region, the sediments unconformably overlying the Valiente For- mation basalt are Plio-Pleistocene in age. The upper Miocene succession of Tobobe Sandstone, Nancy Point, and Shark Hole formations are absent. Furthermore, the northernunitsarecharacterizedbyamajor component of coral reef limestone. The geology of Colon and Bastimentos islandsarecurrentlybeingstudiedandthe FIG.6. GeologicalmapofCayoAgua.Agesinmil- formaldesignationofthelithostratigraphic lionsofyears. units and their ages is not yet finalized. In Fi 5 li 4/C Fi 6 li 4/C GEOLOGYOFBOCASDELTOROARCHIPELAGO 383 general, the sequence is younger than the Valiente Formation.—The spectacular ex- section to the south with the exception of posures along the northwestern coast of the Escudo de Veraguas Formation. The BastimentosIsland(Fig.7)ofcolumnarba- oldest units on Colon and Bastimentos is- salt and flow breccia are assigned to the lands may overlap in time with the top of Valiente Formation solely on their great the Cayo Agua and Shark Hole Point se- similarityinlithology.Theexposuresrange quences at ∼3.5 Ma. from Toro Point to the eastern side of FIG.7. Geologicmapandsection(a-a’)ofBastimentos Fi 7 li 4/C

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logic history of the region and a temporal framework for the evolutionary and eco- logical studies of the macrobiota. Regional Geologic History.
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