An annotated Check list of the Noctuidae (s. l.) (Lepidoptera, Noctuoidea: Nolidae, Erebidae, Micronoctuidae, Noctuidae) of the Asian part of Russia and the Ural region.

243 Pages·2005·3.435 MB·English

by  Vladimir S. Kononenko

N O C T U I D A E S I B I R I C A E
VOLUM E I
AN ANNOTATED CHECK LIST

NOCTUIDAE SIBIRICAE

VOLUME I

AN ANNOTATED CHECK LIST OF THE NOCTUIDAE
(S. L.) (LEPIDOPTERA, NOCTUOIDEA: NOLIDAE, EREBIDAE,
MICRONOCTUIDAE, NOCTUIDAE)
OF THE ASIAN PART OF RUSSIA AND
THE URAL REGION

VLADIMIR S. KONONENKO

ENTOMOLOGICAL PRESS
SORØ 2005
 Copyright 2005 by Entomological Press, Sorø, Denmark
All rights reserved. No part of this book may be reproduced or translated in any form, by print, photo-
print, microphilm, microphiche or any other means without written permission.

Editor in a Chief: M. Fibiger

Text composition and book design: Vladimir S. Kononenko

Frontpage drawing: Xestia penthima (Ersch., 1870) by Vladimir S. Kononenko

Text printed by: Litotryk A/S, DK – 5700 Svendborg, Denmark

Sold and distributed by: Entomological Press, Molbechs Alle 49, DK – 4180 Sorø, Denmark

This publication should be cited as:
V. S. Kononenko. An annotated Check list of the Noctuidae (s. l.) (Lepidoptera, Noctuoidea: Nolidae,
Erebidae, Micronoctuidae, Noctuidae) of the Asian part of Russia and the Ural region. – Noctuidae
Sibiricae. Vol. 1. Entomological Press, Sorø, 2005. 243 pp.

Author’s address:
Institute of Biology and Soil Science, Far Eastern Branch of Russian Academy of Sciences, Vladivostok
22, Russia 690022, e-mail: [email protected]

ISBN 87-89430-12-3
CONTENTS

EDITORIAL FOREWORD . . . . . . . . . . . . . .. . .. . ... . . . . .. . .. .. . . . .... . ... . . .. . .. . . 7
ACKNOWLEGEMENTS .. . . . .. . . . . . . . .. . . . . . . . . . . .. . . . . . . . . .. . . . . . . . . . .. . ... . 8
PREFACE . . . . .. . .. . .. .. . . .. . . . . . . . . . . . . .. .. . . .. . .. . . .. . . . . .... ... . .... . .... 9
INTRODUCTION . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11
MATERIAL AND METHODS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11
CLASSIFICATION . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . . . . . . . . . . . .. . . . . . . 12
BRIEF CHARACTERISTIC OF REGIONS . . . . . . . . . . . . . . . .. . . . . . . .. . .. . . . .. . .. . . . 17
THE URALS . . . . . . . . . . . . . . . . .. . .. . . . . . .. . ... . . .. . .. . ... . . .. . . . . . .. . . .. . .. 18
SIBERIA . . . . . . . . . . . . . . . . . . .. . . . . . ..................... . . . . . . . . . ......... 19
THE RUSSIAN FAR EAST . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. .. . . .. . .. . ... . . .. . .. 23
LITERATURE . . . . . . . ......... . . . .. . . . . . . ...... . .. . .. . . . . . . . . . .... . . . . .. . .. 29
ARRANGEMENT OF THE CHECKLIST . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33
ABBREVIATIONS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30
TAXONOMIC, NOMENCLATURAL AND FAUNISTIC SUMMARY . . . . . . . . . . . . . . .. . .. 35
GENERIC SYNOPSIS . . . . . . . .. . . . . . . . . . .. . .................................. . 36
DISTRIBUTIONAL CHECKLIST . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . .. . . . . . .. . .. . . 61
Family NOLIDAE . . . . . .. . . . . . . . . . . . . .. . .. . ... . . .. . .. . ... . . . .. . .. .. . . . . . . . . 61
Subfamily NOLINAE . . . . . . . . . . . . . . .. . . .. . .. . ... . . .. . .. . ... . . . .. . .. . .. . . .. . . 61
Subfamily CHLOEPHORINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . .. . . .. .. . . 61
Subfamily EARIADINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 63
Subfamily ELIGMINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 63
Family EREBIDAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . ........... . 63
Subfamily RIVULINAE .. . . . . . . . . . . . . .. . . . . . . . . . . . . . .. . .. .. . . .. . .. . ... . . . .. . 63
Subfamily BOLETOBINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . . . .. . .. 63
Subfamily HYPENODINAE . . . . .. .. .... . . .. . . . . . .. . . .. . .. . ... . . .. . .. . ... . ..... 63
Subfamily ARAEOPTERONINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . .. . .. . . 63
Subfamily EUBLEMMINAE . . . .. . .. .. . . .. . .. . . . . . . .. . .. . ... . . .. . .. .... . . . . ... 63
Subfamily HERMINIINAE . . . . . . . . . .. . . .............. . . . . ................ . ... 65
Subfamily HYPENINAE . .. . . . . . . . . ........................ . . . . . . . . . . . ....... 67
Subfamily PHYTOMETRINAE . . . . . . . . . . . . .. . .. . . .. . . .. . .. . ... . . .. . .. . . . . . ... . . 67
Subfamily AVENTIINAE . . . . . . . . . . . . . . . . .. . . . ... . . .. . .. . ... . . . . . . . . . . . . . . . . 68
Subfamily EREBINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . .. . . .. . .. . ... . . . . . . . 68
Subfamily CALPINAE . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . .. . .. . .. . . . . . .. . 68
Subfamily CATOCALINAE . . . . . . . . . . . . . . . . . . . .. . . .. . .. . ... . . .. . .. . ... . . .. . .. 69
Subfamily EUTELIINAE . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . .. . . .. . .. . ... . . .. . .. 73
Family MICRONOCTUIDAE . . . . . . . . .. . . . . . .. . . .. . . .. . .. . ... . . .. . .. . ... . . .... 73
Family NOCTUIDAE . . . . . . . . . . . . . . . . . . . . . . . ......... . . ................... . 73
Subfamily PLUSIINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . ... . . .. . .. . 73
Subfamily EUSTROTIINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . .. .... 75
Subfamily BAGISARINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77
5
Subfamily ACONTIINAE . . . . . . . . . . .. . . .. . .. . ... . . .. . .. . ... . . .. . . .. . .. . . . . . .. 77
Subfamily PANTHEINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . ............ 77
Subfamily DILOBINAE . . . . . . . . . . . . . . . . . . . .. . . . . . . . .. . .. . .. . . .. . .. . ... . . .... 78
Subfamily RAPHIINAE . . . . . . . . . . . . . . . . . . . . .. .. . . .. . .. . ... . . .. . .. . ... . . .. ... 78
Subfamily ACRONICTINAE . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . . . . . . . . . . . ......... 78
Subfamily METOPONIINAE. . . . . . . . . . . . . . . . . . . . . . . . .. . .. .. . . .. . .. . ..... .... . 80
Subfamily SINOCHARINAE. . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . . .. . .. . ..... . .. ... 80
Subfamily AGARISTINAE . . . . . . . . . . .. . . .. . .. . ... . . .. . .. . ... . . .. . .. .. . . . . . .. . 80
Subfamily CUCULLIINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . .. . .. . ... . . . . .. . . 80
Subfamily ONCOCNEMIDINAE. . . . . . . . . . . . . . . . . . . . . .. . .. .. . . .. . .. . ... . . .. ....82
Subfamily AMPHIPYRINAE . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . .. . . .. . .. . ... . . .. . . 82
Subfamily PSAPHIDINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . . .. . .. ...83
Subfamily HELIOTHINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . .. .. . . .. . .. . ... . . .. . .. . 83
Subfamily CONDICINAE . . . . . .. . .. . . . . . . .. . .. . .. . . .. . .. . ... . .... . .. . .. . . . . . . 84
Subfamily ERIOPINAE . ... . . .. . .. . ... . . .. . .. . ... . . .. . .. . ... . . .. .... . ...... .. 85
Subfamily BRYOPHILINAE. . . . . . . . . . . . . . . . . . . . . . . .. . .. . .. . . .. . .. . ... . . .. . .. .85
Subfamily XYLENINAE. . . . . . . . . . . . . . . . . . . . . .. . . .. . .. . ... . . .. . .. . ... . . . ... ..86
Subfamily HADENINAE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ... . . . . . . . . . . . . . . . . 86
Subfamily NOCTUINAE . . . . . . . . . . . . . . . . . . . . . . . . . . .. . .. . .. . . .. . .. . ... . . .. . .. 103
COMMENTS . . . . . . . . . . . . . . .. . .. . .. . . .. . .. . ... . . .. . .. . ... . . .. . .. . ... . . .. . .. 111
BIBLIOGRAPHY . . . . . . . . . . . . . . . . . . . . . . . . . .. . . .. . .. . ... . . .. . .. . ... . . .. . .. . . 195
INDEX . . . . . . . . . . . . . .. . . .. . .. . ... . . .. . .. . ... . . .. . .. . ... . . .. . .. . ... . . .. . .. . 219

6
EDITORIAL FOREWORD

Hopefully this book will open the new series, language, often published in only a few stencil-
“Noctuidae Sibiricae” dealing with the fauna of copies, as a thesis, or they are published as small
the Noctuidae (sensu lato) of the Asian part of and little known local faunas that have all been
Russia (often called “Siberia”). This series con- out of print for a long time. Therefore one of the
tinues and extends the issue Noctuidae Europaea first goals was to gather all available information.
from the Ural Mountains (the geographical limits
A decision was made to split the work, Noc-
for Noctuidae Europaeae) to the Pacific coast of
tuidae Sibiricae, into three volumes: the first
Russia.
goal being:
Five years ago, during the first of many pleas-
– to produce a complete list of all Noctuidae (s.
ant visits to me in Sorø by Vladimir Kononenko, I
l.) taxa that have been reported from Siberia;
complained about the lack of knowledge of the
– to write taxonomic and faunistic comments to
Noctuidae (s. l.) fauna of Siberia – i.e. the large
all questionable taxa or data;
gap between the chain of the Ural Mountains and
the Pacific coast / North America. I mentioned the – to integrate the East Palaearctic and Oriental
need for a genuine book on the subject. We dis- original taxa to the modern classification of
cussed the possibility of producing and overcom- the Noctuidae (s. l.);
ing such an immense task, working our way – to compile a complete list of relevant literature
through the various problems and the possible from Siberia;
solutions. After a few hours of fruitful discussion
The above constitutes the contents of volume
Vladimir volunteered to undertake the work.
1, the text of which has been edited by David
Vladimir has been working more than 30 years
Agassiz. However, just as the first version of the
in the taxonomy and zoogeography of the Noctui-
completed book was finished, Fibiger and Lafon-
dae, mainly the East Palaearctic fauna. He is the
taine (2005) finished their work on the Noctuidae
author of more than 100 publications, among them
(s. l.) systematics, and Vladimir decided to update
three monographs. Vladimir conducted his field
all the text according to this new systematic order,
work in many remote corners of East Siberia and
which is also reflected in the new ‘European List
the Far East. He took part in three expeditions in a
of Noctuoidea (Fibiger & Hacker, 2005).
joint project between Finnish and Russian Acad-
emies for comparative study of the insect fauna of The subsequent volumes of Noctuidae Sibiricae
Siberia and Northern Europe initiated by Kauri will contain a text for each taxa, mainly covering
Mikkola in the early nineties. With his knowledge short diagnosis, the distribution data, larval food
of the Noctuidae and relevant Russian literature, plants, the period activity for larvae and imagines,
Vladimir is currently the only person qualified to colour photos of all species and subspecies from Si-
perform this challenge. Furthermore, he is familiar beria, and illustrations of male and female genitalia.
with all museums and collections that possess ma-
Many lepidopterists and other entomologists
terial collected from Siberia and, through col-
have supported this study and have been ex-
leagues in the Russian Academy of Sciences and
tremely helpful during this exhaustive work and
other former Soviet museums and institutions, it
they will be properly acknowledged in the au-
has been possible for him to study almost all criti-
thor’s foreword. It is the hope that all friends of
cal records or reports from Siberia. For the process-
lepidopterology will appreciate this new series
ing of plates Vladimir has studied and designed
that will give necessary information for biologists
application of the computer graphics for scientific
both inside and outside Russia.
illustrations with excellent, professional results.
The knowledge about the Noctuidae (s. l.) of
the Asian part of Russia is in a diversity of publi-
cations, which makes it difficult to survey. It ap-
Sorø, 4th June 2005
pears in small or less well known journals, en-
tirely in Russian – with no abstract in a foreign Michael Fibiger, Editor in Chief
7
ACKNOWLEDGEMENTS

The preparation and publication of this book Europe and Siberia and the exchange programme
would be impossible without the encouragement between entomologists of Finnish and Russian
and friendly help of many Lepidopterist col- Academies of Sciences for his support and en-
leagues, both professional and amateur. They couragement for this work, for discussions and
have assisted me through consultation, in discus- valuable critical comments.
sions, by making comments or criticisms; by per-
My special thanks expressed to late Prof. C.
mission to work on their collections, with loans of
M. Naumann, for his kind support of museum
material, genitalia slides and type–specimens
work in preparation of this book, as well as other
from the collections under their curation. I express
projects on Palaearctic Noctuidae.
my cordial thanks to the following:
My great thanks to Dr. L. Ronkay and Mr. G.
Prof. Dr. C. M. Naumann, Dr. D. Stüning, Dr.
Ronkay for fruitful discussion of many critical
W. Speidel (ZMFK, Bonn, Germany), Dr. W.
taxa and their kind help given me in course of
Mey (MNHU, Berlin, Gernany), Mr. G. Behounek
study of the collection of HNHM.
(Grafing b. Münich, Germany), Mr. H. Hacker
I am indebted to Dr. M. Fibiger (Sorø, Den-
(Staffelstein, Germany), Mr. B. Shmitz (Ger-
mark) who inspired me to start the “Noctuidae Si-
many), Mr. L. Kühne (Potsdam, Germany); Mr.
biricae” project, critically revised the manuscript
M. Honey, (BMNH, London, Great Britain); Prof.
and made generous efforts for its publication.
Dr. N. P. Kristensen, Mr. D. Nilssen, Mr. Alex
Madsen, Mr. Gert Jeppesen, (Copenhagen, Den- My special thanks to Mrs. Mariann Fibiger
mark), Dr. B. Gustafsson, (Stockholm, Sweden), (Sorø, Denmark) and Dr. D. Agassiz for linguistic
Prof. Dr. K. Mikkola, Mr. J. Kullberg (ZMHU, correction of the English.
Helsinki, Finland), Mr. M. Ahola, Mr. K. Nuppo- The museum work for Noctuidae Sibiricae pro-
nen, Mr. J. Junnilainen (Finland); Mr. H. Thöny
ject has been supported in part by grants from the
(Potü, Brasil); Dr. J. Rawlins (Carnegie Natural
Finnish Academy (in the year 1998, and in the year
History Museum, Pittsburgh, USA), Dr. J. D. La-
2003) and the Finnish Natural History Museum in
fontaine (Agriculture Canada, Research Branch,
the year 2001, the Deutsche Forschungsgemein-
Ottava, Canada); Dr. M. Hreblay (Erd, Hungary),
schaft (DFG) in the years 1999, 2001 and 2004 and
Dr. L. Ronkay (HNHM, Budapest), Mr. G. Ron-
by Ernst Mayer Traveling grant in the year 2004.
kay (Budapest, Hungary); Mr. S. Sugi, Dr. M.
I hope that this book will be helpful to noctuid
Owada, Dr. S.-I. Yoshimatsu (NSM, Tokyo,
researchers, both professional and amateur lepi-
NIAST, Tsukuba, Japan), Dr. J. Viidalepp (IZB,
dopterists, as an introduction to the noctuid fauna
Tartu, Estonia), Prof. Dr. H. J. Remm (Tartu Uni-
of the Urals, Siberia and the Russian Far East, as
versity, Estonia); Dr. P. Ivinskis (Vilnus, Latvia);
well as a stimulus for further research into insect
Mr. N. Savenkov (Riga, Latvia); Dr. A. Zilli (It-
diversity in these regions.
aly, Roma); Prof. Dr. Z. F. Kljutshko (Kiev State
University, Ukraine); Dr. I. L. Sukhareva, Dr. A. At present a large amount of material of Noc-
L. Lvovsky, Dr. A. Matov, (ZI, St. Petersburg, tuidae from Asiatic Russia is accumulated in
Russia); Dr. A. V. Sviridov (ZM Moscow State Western museums and in many private collec-
University, Russia); Dr. A. V. Nekrasov (Mos- tions. I would be greatly appreciate from my
cow, Russia); Dr. V. V. Dubatolov, the late Dr. colleagues any further comments, critical notes,
G. S. Zolotarenko (ZM IASE, Novosibirsk), Dr. and additional information on the specific diver-
E. A. Beljaev (IBSS, Vladivostok) and many sity and distribution of the Noctuidae (s. l.) in
other colleagues. Asian Russia.
I am deeply grateful to Prof. Dr. K. Mikkola,
initiator and organiser of numerous expeditions for Vladimir Kononenko
comparative study of the insect fauna of Northern Vladivostok, July 3rd, 2005

8
PREFACE

It gives me great pleasure to present to my col- this huge country were initiated in the mid 18th
leagues and other interested readers an annotated century and continued during the next century by
Checklist of the Noctuidae s. l. (i.e. Nolidae, Ere- expeditions of the Russian Academy of Sciences
bidae, Micronoctuidae and Noctuidae) of the (I. Gmelin, P. Pallas, A. Middendorff, G. Radde,
Asian part of Russia. R. Maak, L. Shrenk, N. M. Przhevalsky, I. Cher-
The Noctuidae (s. l.) is the largest, most di- sky, A. Chekanovsky, O. Herz, E. Toll and other
verse complex of related families of the order explorers) and continued for more than 200 years,
Lepidoptera, superfamily Noctuoidea with a the inventory of such huge family as Noctuidae is
worldwide distribution with over 25000 species in not yet complete.
the World, i.e. about 20–25% of the total number The most comprehensive guides to the World
of Lepidoptera species. Well over 10000 species
or Palaearctic fauna of the Noctuidae, including
are known from the Palaearctic region and at least
those of the Asian part of Russia, since the begin-
about 1400 species are known from the Asian part
ning of the 20th century were for a long time the
of Russia. The representatives of this complex of
classical catalogues by Staudinger & Rebel
families have great ecological as well as economic
significance. They are an important component of (1901), Hampson (1903–1913, 1914, 1920) and
natural or agricultural systems due to their high the colour plate monographs by Seitz (1909–1914,
specific diversity, the abundance of many species, 1931–1938). Since the publication of the last fa-
high migratory ability of imagines and phyto- mous issue numerous descriptions of new taxa,
phagous feeding habits of larvae. taxonomic revisions, and faunal lists have been
The classification of the higher taxa of the published for the Palaearctic by R. Püngeler, O.
Noctuidae (s. l.) is not completely resolved. The Bang–Haas, C. Corti, M. Draudt, Ch. Boursin, E.
rank, taxonomic position and composition of Dufay, A. Kostrovicky, I. Kozhantschikov, S.
some subfamilies and tribes are still disputed, Sugi and others.
uncertain, or not fixed yet. For this reason and
During last two decades the Noctuidae have
because of the former family Noctuidae recently
become the subject of intensive study by many
has been splitted for three families (Fibiger &
lepidopterists in Europe, Asian countries and
Lafontaine, 2005) I have entitled my present work
North America. Our knowledge of noctuids of the
“Checklist of the Noctuidae (sensu lato)”. I have
Palaearctic region as a whole, as well as of the
included in the list the family Nolidae with sub-
Asian part of Russia has been improved consid-
families Sarrothripinae, Chloephorinae, Eariadi-
erably. The taxonomy and classification of the
nae and Eligminae which were considered by
Noctuidae is undergoing dramatic changes. Nu-
former authors either as subfamilies of Noctuidae,
merous species have been described or discovered
or (some of them) as distinct families, or as sub-
from the study area and neighbouring regions.
families of Nolidae and most recently fixed by
Some subfamilies, many genera or other taxo-
Holloway (1998) in Nolidae. On the other hand
nomic groups have been revised; numerous
the subfamily Pantheinae is treated here as a noc-
changes have been introduced in the nomencla-
tuid subfamily, while Rawlins & Kitching (1999)
ture. Guide books, checklists and catalogues of
raised its rank to family level.
the Noctuidae have been published for Asian
The Asian part of Russia is a vast territory ex- countries: Japan (“Moths of Japan” by Sugi,
tending from the eastern slopes of the Ural Range 1982a, Owada, 1982); China (“Iconographia In-
in the West to the Pacific coast of North Asia in sectorum Sinicorum” by Chen, 1982; “Noctuidae
the East, and from the Arctic Ocean in the North of Xizang” by Chen et al., 1989; “Noctuidae
to the Kazakhstan and the Mongolian Steppes in Sinica” by Chen, 1999); Taiwan (“Checklist of
the South. Even though the faunistic surveys and the Lepidoptera of Taiwan” by Inoue, Sugi,
collecting of insects, particularly Lepidoptera, in Owada, 1992); Korea (“Illustrated Catalogue of
9
the Noctuidae in Korea” by Kononenko, Ahn, author in the years 1972–2005 in the Russian Far
Ronkay, 1998); the Himalayan region (series East and East Siberia.
“Moths of Nepal” by Haruta, Yoshimoto, 1992– Of special help for this project has been the
1998; Hreblay & Ronkay, 1998); the Near East, vast material of Noctuidae collected during the
including South East part of Russia (“Die Noctui-
joint Finnish – Russian expeditions to Siberia and
dae Vorderasiens” by Hacker, 1990) and Europe:
the Russian Far East (project No 20 between the
(“Systematic list of the Noctuidae of Europe” by
Russian Academy of Sciences and the Academy
Fibiger & Hacker, 1991, 2005; “Noctuidae Eu-
of Finland, coordinated by Prof. Dr. Kauri Mik-
ropaeae” by Fibiger, 1990, 1993, 1997; G. Ronkay
& L. Ronkay, 1994, 1995; Ronkay, Yela, Hreblay, kola in the years 1981–1993 and by Mr. Seppo
2001; Hacker, Ronkay, Hreblay, 2002; Goater, Koponen, Lic. Ph., in the years 1994–1998) and
Ronkay, Fibiger, 2003; “The Lepidoptera of during subsequent expeditions of private Finnish
Europe. Noctuidae” by Nowacki & Fibiger, 1997; collectors to the Southern Ural and to the Altai
“Noctuidae of Central Europe” by Nowacki, 1998). Mts. (K. Nupponen and others). Details are given
Worldwide nomenclature catalogues of generic and
in connection with each area.
specific names of the Noctuidae have been pub-
The Checklist is given in the form of a table, it
lished (Nye, 1975; Poole, 1989).
includes 1413 species known to occur in the Urals
At the same time no modern general guide,
and neighbouring regions, Siberia and the Russian
catalogue or checklist exists for Noctuidae from
Far East. The distribution of each species in 17
the vast territory of Asian Russia between Europe
regions from the Urals to the Pacific coast is tabu-
and the Far East. Various local faunal, descriptive
lated. Although the Urals belong to Europe I in-
and taxonomic articles dealing with the Noctuidae
clude in the list the faunal data for this zo-
from different regions of the Asian part of Russia
ogeographically very important region as a transi-
have been published by both Russian and foreign
tional area between Europe and Asia. The distri-
authors. However they are dispersed in separate
bution of each taxon is sketched using appropriate
articles published in small local journals or as
codes. Only specific names are listed in the table.
collections of papers and are often inaccessible to
Subspecific taxa, synonyms (mainly regional and
researchers. Although considerable museum mate-
some recently introduced for widely distributed
rials and literature on taxonomy, distribution, and
species) uncertain taxa, and taxonomically of
ecology of noctuids in Asian part of Russia have
faunistically problematic cases are discussed in
been accumulated, these data are difficult to use
the chapter “Comments”. The taxonomic changes
due their scattered nature.
made in this book are listed in the taxonomic
The present Checklist aims partially to fill this summary. The generic synopsis includes names of
gap and summarise our modern knowledge of the genera recorded in the region with complete syn-
fauna, taxonomy and nomenclature of the Noctui- onymy. The literature sources for Noctuidae of
dae (s. l.) of Asian part of Russia within its ad- the Asian part of Russia and neighbouring coun-
ministrative borders. It is based on collections tries as well as general publications on taxonomy
of museums and private collectors listed below, of the Noctuidae are listed in the “Bibliography”
literature sources and materials collected by the with a guide in the chapter “Literature”

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