A Synopsis of the Prionine Genus Spinimegopis stat. nov. (Coleoptera, Cerambycidae, Prioninae). (Revisional Studies of the Genus Megopis sensu LAMEERE, 1909-8)

2007·43.7 MB·English

by  Komiya, Z

Elytra,Tlokyo,35(1):345-384,May30,2007
ASynopsisof thePrionineGenusSpzmmegopfsstat nov.
(Coleoptera,Cerambycidae,Prioninae)
(RevisionalStudiesof theGenusMegop1issensu LAMEERE,1909-8)
Ziro KOMIYA
3-2-12,Shimouma,Setagaya-ku,Tokyo, 154-0002Japan
and
Alain DRUMONT
EntomologyDepartment,InstitutroyaldesSciencesNatureIlesdeBelgique,
29RueVautier,B-1000Bruxelles,Belgium
Abstract Spinimegopts K. OHBAYAsHI, 1963 is proved to be a full genus.
Megoplsfomosanafomosana,M f nlppo川ca,M f awazoe1,M f lshlga1ana,M f
1anhsuensis, M f yakushimana, M f okinawana,M f hachijoana,M piliventris,M
malastaca,M lividipennis,Mbuckleyi,M tibialis,Mnepalensls,M cingalensls,M
mediocostata, M moretto1 and M perroti are transferred to this genus. Megopis
fiavlpennisDEMELT,1989isregardedasajuniorsynonymofMmalaslacaHAYAsHl,
1976. MegoplsantennalisFUCHS, 1965isregardedasasubspeciesof M piliventris
GRESSITT,1950. ThreesubspeciesofM formosana,nipponlca,kawazoeiandlshigaki
anaarerevivedasindependentspecies. Fivenew speciesandanewsubspeciesare
describedunderthenamesSp加megopls ach1na,S fuJita1,Sdefaayei,S tla1andS.
curtlcornlsspp nov.,ands foirmosanatamdaoar1asubsp nov.Lectotypesandparalec-
totypesaredesignatedtors buckley1' (GAHAN,1894)andS.1ividipennls (LAMEERE,
1920). Akeytothespeciesof thisgenusisgiven.
Resume Danscet article, Spinimegopls K. 0HBAYAsHI, 1963est reconnu
commeungenrevalide. Megoplsfomosana,M f nip-Ilea,M f awazoe1, M f
ishigakiana,M f lanhsuensis,M f yakushimana,M f okinawana,M f hac11i、joana,
Mpiliventris,Mmalasiaca,M livldipennls,Mbuckley1,Mtibialis,Mnepalensts,M
cl'tgalensis,M 'nediocostata,M morettotetM perrotisent transferosclanscegenre.
MegopisflallipennisDEMELT,1989estreconnucommeunsynonymedeM malasiaca
HAYAsHI,1976. MegopisantennalisFUCHS,1965estconsiderecommeunesous-espece
deM ptliverttrisGREssITT,1950. Troissous-especesdeM fo(rmosanasent,quanta
e11es,eleveesaurangd'especesindependantesetvalides. Cinqnouve11esespecesetune
nouve11essous-especesent decritescommeSpinimegoplskachina,S fuJ'1tat,S dela-
hayei,S huaietS.curticornlsspp nov.,ainsiques formosanatamdaoanasubsp nov.
346 Ziro KoMIYAand Alain DRUMONT
DeslectotypesetparalectotypessentdesignespourS buckley1(GAHAN,1894)etS
1ivldipennts(LAMEERE,1920). Uneolepourtouteslesespecesdugenreestdonnee.
Spinimegoplswasdescribedby Kazuo OHBAYAsHI in l963asasubgenusof the
genus Megopfs to receiveM nplpo川caMATSUSHITAandM awazoe1 HAYASHI. He
characterizedthissubgenusnotonlyhavingthreelateralspinesofthepronotumbutalso
havinghair-fringesunder themaleantennae. HAYAsHI (1976, 1980) proposed to
enlargethissubgenusbyincludingseveralspecieswhichhavedistinct threespineson
eachsideofpronotumbutnothavinghair-fringesofantennae. HetransferredMegopis
(Aegosoma) tibialisWHITE to thesubgenusSpinimegopisanddescribed threemore
species,M.lanhsuensls,M malasiacaandM riepalensis,underthissubgenus. However,
hedidnotexaminetherestoftheknownspeciesinthegenusMegopiswhethersomeof
themshouldbe transferred to thissubgenusor not. In thispaper, wearegoing to
succeed these works.
Beforethissubgenuswasproposedby K. 0HBAYAsHI (1963), thephylogenetic
importanceofpronota1spineshadgenerallybeenoverlookedbypreviousworkers. Only
exceptionwasmadebyGREssITT(l940)whenheincludedthischaracter inaseriesof
differences between thesubgeneraAego11pton and Baralipton (see KoMIYA, 2005).
Concerning the Aegosoma group, in which the most species to be transferred to
Spinimegopishadbeenplaced,thischaracterwasindicatedonseveralspeciesbyGAHAN
(1906) but he did not suggest any moremeaning of this character state. After
examination of a longseriesof specimenswhich havespines on the pronotum, we
concludedthatthischaracterclearlydividesAegosomasensuLAMEERE(1909) intotwo
parts,viz.,AegosomaandSpinimeg〔)pls. Eachof thesetwogroupsbearssimilarity in
manyothercharactersandtheyareverydistinct fromeachother. Therefore,weregard
SpinimegopisasanindependentgenusasKoMIYA(2005) noted.
Thespeciestobetransferredtothisgenushavepreviouslybeenplacedinthegenus
Megopisbut theywereseparately locatedinmanysubgenerasofar. Inthefollowing
lines, wearegoingto transferMegopfs加c eyz, M fvidlpenms, M. perrotf andM
med1ocostata fromthesubgenusAegosoma, M clngalerislsandM antennalis(whichis
now regarded as a subspecies of S. piliventris) from thesubgenusBaralipton, M.
piliventris from thesubgenusAegolipton, M moretto1(whichwas not placed in any
subgenusat thetimeofdescription) from thesubgenusMegopis,M f(ormosana, M
formosananipponica, M f kawazoe1, M f yakushimaria, M f lanhsuerlsis, M f
hachijoana, M f okinawana, M f ishigakiana, M nepalensis, M malasiaca and M
tibialis from thesubgenusSpinimegopis (nipponica, kawazoei and ishigakiana are re-
gardedinthispaperasindependentspecies). Fivenewspeciesandanewsubspeciesare
describedundernamesS kachina,Sfujitai,S delahayei,S huaiands.curticornisspp
nov andS.fo(rmosanatamdaoanasubsp nov.Megopis avlpennisisregardedasajunior
synonymofM malasiaca.
The abbreviations andspecial terminologies used in this paper areas follows:
NSMT - NationalScienceMuseum(Nat.Hist),Tokyo;BMNH- TheNatural History
SynopsisofthePrionineGenusSpini1negopis 347
Museum, London; IRSNB-InstitutRoyaldesSciencesNature11esdoBelgique;CAS-
CaliforniaAcademyofScience, San Francisco; 0MNH- Osaka Museum of Natural
History; MNHN- Museum national d'Histoire nature11e, Paris; ZSMC -Zoo1ogische
Staatssammlung, Miinchen; ADC-collection of Alain DRUMoNT, Belgium; AWC-
co11ection of AndreasWEIGEL, Germany; CHC-co11ectionof Carolus HoLzscHUH,
Austria;EVC- co11ectionofEduardVIvEs,Spain;HFC- co11ectionofHiroshiFuJITA,
Japan; YKC-co11ection of Yoshiyasu KUsAKABE, Japan; ZKC- co11ection of Ziro
KoMIYA,Japan. Measurementsofbodyparts:BL-bodylength fromclypeustoapices
ofelytraorabdomen; HL-lengthof head fromclypeustobase; HW -widthof head
acrosseyes;PL- lengthofpronotum;PW-maximumwidthofpronotum;PA-apical
widthofpronotum;PB-basalwidthof pronotum; EL-lengthof elytra; EW-maxi-
mumwidthof closedelytra; AL-length of antennae;Am- lengthof (n)thantennal
segment. Terminologies: CI -first internal costaofelytron; C2-secondcosta(outer
internalcosta); C3- thirdcosta(innerexternalcosta);C4- fourthcosta(outerexter-
nal costa); Sa-spineof anteriorangleof pronotum; Sm-1atera1spineof pronotum,
whichisusuallyplacedinbasalthirdorclosertobase;Sh-spineofposteriorangle; hair-
fringe- longbristlyhair linesunderantennaewhichareobviouslylonger thanhairson
the other sideof antennae.
GenusSpinimegopis K. 0HBAYAsHI,1963
AegosomaWHITE,1853(notSERVILLE,pr'oparte).
Aegosoma:GAHAN,1894,1906(notSERvILLE,proparte).
Megopls:LAMEERE,1909(notSERVILLE,proparte).
Megopls(Aegosoma):MATSUSHITA,1933,1934(notSERvlLLE)
Megopis (Aegosoma):GRESSITT, 1950(notSERVILLE).
Megopls (Aegolipton):GREssITT,1950(not GREsslTT.1940).
Megopis(SpI川'negop1s): K.0HBAYAsHI,1963.
Megopls(Aegosoma):FUCHS,1965(notSERvILLE).
Megopls (Baralipton):FUcHs.1965(notTHOMSON).
Megopls (Sp加加1egop1s):HAYAsHI,1974,1975,1976.
Megopis (SpI川megopls):FUJITA,1980.
Megop1s (Spi川megopis): KUsAMAetaf.,1984.
Megop1s (Sp加1tlegopfs):N. 0HBAYAsHIetaf.,1992.
Typespecies:Megop1s(degosoma)nlppomcaMATSUSHITA,1934,Trans nat.Hist.
Soc. Formosa, 24: 538.
Body elongateandcylindrical. Bi t7-55mm, usuallybetween 30and 45mm.
Bodycolorbrown,oftenyelloworalmostblack. Headandpronotumcoveredwithlong
hairs,elytrahairedor glabrous, undersideusuallycoveredwithlongandthick hairs
exceptabdomenwhichisthinlyhaired.
Head short. Mandibles usually as longas one-fifth of head, each mandible
sickle-shaped,externalsidesmoothlyarchedorobtuselyangledandsometimesfurnished
withanexternaldent (tubercle), internalsidesharplybladedand furnishedwithadent
348 Ziro KoMIYAand Alain DRUMoNT
closetothebase. Eyesbulging,coarselyfaceted,interspacebetweeneyesofdorsalside
usuallyahalfaswideaseachlobe,lowereye-lobesongulamorewidelyseparatedthan
ondorsalside. Antennaltubercleslargebuttheir innerendsoftennotclearlydefined.
Antennae1l-segmented, AL/BL 0.8-1.2,早, 0.6-0.9;basal segments (1-4) sub-
,
cylindrical,apicalsegments(5-11)depressedandangledecto-apica11y,furnishedwitha
longitudinal carinaalongexternal line;segment 3about as longas or longer than
segments4十5十6,segment1furnishedwithdistinctlongitudinalgrooveoninnerside,
severalbasalsegmentshair-fringedinsomespecies.
Pronotum sub-rectangular or trapezoidal, 1.5-2.0 times as wide as long; lateral
marginedgedfrombasetoanteriormargin,usuallyfurnishedwithdistinctthreespines
(Sa,Sm,Sh) oneachside,sometimesoneor twospinesobsolete;disc, unevenand
convexatcentralpart. Scutellumlinguiformorsemicircular.
Elytrausually longer than2.5 times of united lengths of headandpronotum,
pubescentorglabrous,marginedwithdarkcolorinmostspecies;innercostae(CIand
C2) usuallydistinct,C3andC4mostlyabsent;suturalteethsmallerabsent.
Legsslender,thicklyhairedingeneralbutthelengthandthicknessofhairsarevery
differentbyspecies;metatibiaedepressedlaterallybutnotstronglyso. Pro-,meso-,and
metasterna,meso-andmetepisternaandpro-,meso-andmetacoxaethicklycoveredwith
longhairs.
Penislongandslender, lateral lobesincludingbasal ring0.5-0.7timesas longas
penis.
Notes. The genus Spinlmegopis is similar toAegosoma but is distinguished by
havingsegment3ofantennasubcylindrica1andwithoutinternalgroove,andpronotum
furnishedwith threedistinctspines on each side. InAegosoma, thesegment 3 is
furnishedwithalongitudinalgrooveinternallyandpronotumalwayslackinganyspines.
ThreespinesoneachsideofpronotumarealsoobservedinthegenusBaraliptonbutthe
genus Baralipton hassegment 1 of antenna furnished with distinct apical spineand
without anygrooveinside.
Sp加'megopistibiah'sSpecies-group
Spinimegop1istibialis,S buckleyi,S nepalensisandS kachinasp nov formthe
tibialisspecies-group,whichischaracterizedbyhavingglabrousandsemi-transparent
elytra furnishedwithstrongcostae.
Sp加'megopistibiah's(WHITE,1853),comb
nov
(Figs.1,2)
Aegosoma tiblaleWHITE, 1853,Cat.Coleopt. Ins.Coll.Brit.Mus., Longic.,7:32; 1853, Proc.zool.Soc.
London,1853:28. - GAHAN,1906,FaunaofBritish India,CeylonandBurma,Coleopt.,1:47.
Megopls(4egosoma)ttbialis:LAMEERE,1909,Annis.Soc.ent.Belg.,53:141;1919,Gen.Ins.,(172):74.
-
HuA,1982,CheckListof theLongicornBeetlesofChina,4.
Megopls(Sp加megopls) tibfafls: HAYAsHI,1971,Ent.Rev.Japan,23:84.
SynopsisofthePrionineGenusSpinirtlegopis 349
M ale. Body generallydark brownoralmostblack;elytraoften reddishespe-
ciallyinfemale; thicklyhairedonhead,pronotum,scutellumandundersideexcept for
middlepartsofabdomen,antennaeglabrous,elytraglabrous.
Headnot large;mandibleshort, theexternal linesabruptlybent inwardsatabout
middle,eachfurnishedwithasmall innerdentatbasalthird;apexofpalpustruncate;
interspacebetweeneyes0.4-0.6timesaslongaseachupper lobe. Antennacylindrical
and thickened in segments 1-4, weakly depressed dorso-ventra11y in several apical
segmentswhicharefurnishedwithlongitudinalcarinaerunningalongexternalmargins
andsometimeswithanothercarinarunningoninternalsideparallel to theexternalone;
segment l furnished with deep internal groove, closely granulate in segments1-4,
irregularlyfurnishedwithsparsepunctures,scatteredwithgranulesandshortwrinkles
insegments5-11;AL/BL0.86-1.06inmale(richinvariationbutusuallyover1.00),Al
3/All 2.9-3.4, Al3/Al4十5十61.00-1.07, Al7=Alli.
Pronotum wide, almost rectangular andslightlynarrowedanteriorly,widest at
middlespine(Sm), furnishedwiththreedistinctspinesoneachside,discmat,covered
withsmallandirregulargranulesandsparselonghairs. LP/WP0.49-0.68. Scutellum
linguiform, coveredwith longhairs.
Elytra long, semi-transparent, covered with shallow punctures at basal third,
puncturesgraduallymingledandtransitingtoindistinctgranulesatapicalhalves;lateral
sides widest at about basal fourth, then gradually narrowed and rather suddenly
roundedjustbeforeapex;eachelytronfurnishedwithtwodistinctcostae(CI,C2),CI
startingfromthesideofscutellum, runningparallel tosuturalmargininbasalfourth,
thenslightlybentinwardsanddisappearingjustbeyondmiddle,C2startingjust inside
ofhumeri,runningobliquelyinwardsinbasalthree-fifthsandthenbecomingsubpara11e1
tosutural lineanddisappearingjustbeforeapex;sutural toothindistinct orabsent.
Legs long and slender, thinlycoveredwithhairs; protibiae thick, tarsinarrow,
united lengthofthreetarsalsegmentsabout aslongastheclawsegment.
Fem ale. Similartothemalebutheadnarrower,pronotumwidestatbaseand
distinctlynarrowedapicad,antennaeshorter,AL/BL0.60-0.72.
BL:(f, 21.5-35.7mm,早,35.0-45.0mm.
Holotype ,preservedinBMNH.
Distribution. Northeastern India (area close to Nepal), Nepal, Bhutan (new
record,examinedspecimen:RordHongtsho-Lobesa,3kmeastofDochuLapass,2,800
m,6-VIII-2005,Kautt NAUMANNleg.inZKC),China(Tibet) (examinedspecimens:
3 ,Nyalam,3,200malt.,9~10-VII-1998,inZKC).
Specimensexamined. 19 l3早早, fromeasternNepalandnorthernIndia,other
,
thantheabovementionedexamples (IRSNB,ZKC,ADC).
Spinimegopz'sbuckleyi (GAHAN,1894),comb nov
(Figs 3,4)
Aegosomabuckley1GAHAN, 1894, Ann.Mag nat Hist., (6),14:227
350 ZiroKOMIYAandAlainDRUMoNT
Me9oPSI (‘4egosoma)buckleyt:LAMEERE,1909,Annis.Soc.ent.Belg.,53:142
ThisspeciesresemblesStibialisbutsi dsitinctinthefollowingcharacters:
Bodyslenderer;antennaemorerugoselygranulate,Al3relativelylong,Al3/All3.9
-4.3,Al3/Al4十5十61.03-1.12;elytrathicklygranulateandmat,slendererthaninS
tibialis,HL/HW2.5-2.7, (CI)extendingtoapicalfifthofelytron.
BL: ,40.0-42.6mm,早,39.8-45.0mm.
Distribution. NorthernIndia(Himacha1PradeshandUttarPradesh).
Typedesignation. Twosyntypes, ld'',1早,arepreservedinBMNH. Wedesignate
asthelectotypethemalespecimen, length40mmwithattached labels:“Syntype”,
“60.15E.L.C”,“N. India”,“AegosomaBuckleyi TypeGAHAN”. Thesyntype早is
designatedasaparalectotypewhichhaslength46mm, labels“Syntype”,“60.15E. L.
C”, “N. India”. According toDr. SharonSHUTEof BMNH, “60.15 E. L. C” is
registrationlabeland60.15standsforthe15thlotregisteredin1860andthesespecimens
werepresentedbythesecretaryoftheIndiaBoard.E.L.C.probablystandsforthename
ofacollectorof thislotwhoisregisteredonlyasDr.CANTOR.
Specimensex:lamined. 5 3-9--9, UttarPradeshbor.,JosimathAuli,2,800m.
,
alt.1-VIII-1994(ZKC); l f,samelocality,13~17-VII-l994;1早,Himacha1Pradesh,
Solang,2,700m,20-VII-1998(ADC).
SpinimegopisnepalensisHAYAsHI, l971, comb
nov
(Figs 5,6)
Megopis(Spinimegopis)tlepalenslsHAYAsHI,1971,Ent.Rev.Japan,23:83
ThisspeciesisclosetoS tibialisbut isconspicuouslydifferentinlargerbodyand
paleyellowishelytra.
Bodyrobust, integumentdarkbrownandusuallyreddishonpronotumandlegs,
elytrapaleyellowishbrown,semi-transparent,marginedwithdarkcolor.
Headlarger,antennaltubercledeveloped,antennaeshorter,AL/BL0.88-0.98in
male,0.76-0.89infemale,Al3/Alto38-0.41inmale,0.35-0.37in female,Al3/Al4十
5十61.07-1.12,segment11furnishedwithdistinctknotof fusedsegment12.
Elytra furnished with distinct CI and C2, CI extending to apical fifth and
disappearing,ormeetingC2orsuturalmargin;suturaltoothsmall.
Legsstout,thicklyhairedonundersideandthinlysoondorsalside,tarsalsegment
1aslongasunitedlengthofsegments2and3,clawsegmentslightlylongerthanunited
lengthof1-3.
BL: 37.5-54.5mm,早,43.7-54.4mm.
,
Distribution. Nepal (nearWalunchung,alt. l,620-3,050,west ofMt. Kanchen-
junga,nearTaplejung), India(Sikkim) (newrecord).
Types. Holotype Unnamedplace2,450mtoChowki,E. Nepal, 1,620m, 29-
,
VII-1963,)inNSMT.Twoparatypesweredesignatedbutwewereunabletofindthem.
Specimensexamined. 3 ,1早,Taplejung,easternNepal,VII-2000,8 8早早,
,
SynopsisofthePrionineGenusSpinimegopls 351
near Mangan, Kanchenjungaregion,SikkimHima1,2,225malt. 10~15-VIII-1995,
Viktor SINIAEF& Evgeniy AFoNIN leg. (ZKC, ADC); 1早, Sikkim, III-IV, H.
FRUHsToRFER (IRSNB) (this femalewasidentifiedwithAegosomabuckleyi GAH by
LAMEERE).
Spinimegopiskachinasp nov
(Figs 7,8)
M ale. Integument almost uniformly brown, head, pronotum, scutellum and
most part of undersidecovered with long hairs. Head rather small; each mandible
obtusely angled at themiddleof external line indorsal view, furnished withshallow
longitudinal foveaeonexternalsideofbasalhalf;antennal tuberclenotstrongly raised.
Antennaeshorterthanbody,AL/BL0.78-0.93;segment3relativelyshort,notstrongly
thickenedandslightlydepressedlaterally,undersidethinlyscatteredwithshorthairs;Al
3/All 25-2.7, Al3/Al4十5十6098-1.02.
. .
Pronotum mat, narrower as compared with that of other species of thesame
species-group and more strongly narrowed apicad; three lateral spines obtuse and
seldomformingacutespines;anteriorangleobsoleteandoftenabsent.
Elytraslender, glabrous,smoothandshiny, without granulesexcept onbasal part
ofcostae,EL/EW2.6-2.9,furnishedwithfeeblyraisedcostaewhichusually fadeout in
apical third.
Legs slender, thickly covered with short hairs underside; tarsi narrow, three
segmentsofthesamewidth,clawsegmentaboutaslongasunitedlengthofpreceding
threesegments.
Female. Similartomalebutantennaeshorterandslenderer, AL/BL0.58-0.70,
pronotummorestronglynarrowed forwards, elytrabroader.
BL: 27.8-38.7mm,早,34.5-39.6mm.
,
Distribution. Myanmar, (knownonly fromthetypelocality).
Typeseries. Holotype: ChuduRazimountain range, East Kachin, Myanmar,
,
VII-2004, deposited in NSMT. Paratypes: ti 149 ,samedataastheholotype,
,
(NSMT,ZKC),1早,samelocality,VI-2004, (YKC).
Notes. ThisnewspeciesisclosetoS tibialisWHITEbut quitedifferent inshorter
antennaewhicharedistinctlyshorterthanbodyinmaleandshorter than two-thirdsof
body in female, relativelyshortersegment3ofantennae (Al3/All 2.5-2.7) whichis
Figs.1-8 (onp352). Habitusof Spinitnegopisspp. - 1-2.S. tibialis: 1, male from Nepal, 2,
female from northern India. - 3-4.S b1lckleyi:3,male,4, female. - 5-6.S nepalensls: 5,
malefromNepal,6, female fromnorthernIndia. - 7-8.S kachtttasp nov:7,male holotype,
8, female,paratype.
Figs 9-17(onp353). Spinlmegoptsspp. - 9-10.S.cingalensismale:9,habitus,10,obliqueview.
- 11-13.S. mediocostata: 11,malehabitus, 12 femalehabitus, 13,malehead. - 14. S.
moretto1:malehabitusof paratype. - 15-17.S. perrotl: 15, male fromSapa, habitus, 16-17,
antennae.16.male17. female.
352 Ziro KOMIYAand Alain DRUMoNT
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SynopsisofthePrionineGenusSpinimegopis 353
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354 Ziro KOMIYA and Alain Di;uJMoNT
providedwithsparseshorthairsunderside,obtusespinesofpronotumandlesspromi-
nentcostaeofelytra.
2. SpinimegopislividipennisSpecies-group
Sp加megopfsffvldlpemfs,Smafasiaca,S fuJftaisp nov andS defa ayeisp nov.
conform the lividipennlsspecies-groupwhich is characterized by having a distinct
subvertica1processonanteriorendofprosternum(Figs 24-28).
Spinimegopismalasiaca(HAYAsHI,1976), comb nov
(Figs.18,25)
Megopis(Spinimegopis)malasiacaHAYAsHl,1976,Bull.Japanent.Acad.,9:25.
Megopis (Aegoso'na)?a、1ipennisDEMELT,1989,Entomofauna,10/14:241.Syn nov.
Thisspecies ismarkedlydifferent fromanyotherspeciesof thegenus inhaving
robustbodyanddevelopedsegment 3ofmaleantennae.
Integumentochreforthemostpart,scutellumandelytrayellowandmarginedwith
darkbrown,apicesofmandiblesandeyesblack;head,pronotumandundersideexcept
abdomencovered with longhairsandother portionsincludingelytramostly covered
withthinpubescence.
Antennaethick,AL/BL 1.03-1.10早,0.62-0.70,segments1-4roughlygranu-
,
late,segments6-11 furnishedwithtwocarinaerunningundersideandsparsegranules
along internal margin, segments 11almost devoid of knot, Al3/All 3.6-3.8, Al3>
Al4-7, Alli =Al5.
Pronotumstronglyconvex,providedwithapairof vaguebut largeprotuberances
ateachsideofmedianline; largegranulesscatteredontheseprotuberancesandnear
basalangle.
Elytrawidest just behindhumeriandslightlynarrowed tobasal fifth,thenbecom-
ing parallel again and suddenly rounded apicad. CI and C2strongly raised and
sometimesvery feebleC3orC4beingobserved.
Legs rather short ascomparedwithbody; tarsishort,segment 3distinctlywider
than tand2,clawshorter thanunitedlengthofthreeprecedingsegments.
Prosternumfurnishedwithadistinctprocessatthemiddleofanteriormargin(Fig.
25),whichisprojectedobliquelydownwards,obtuselypointedapicadandcoveredwith
longhairs.
BL: ,40.8-65.8mm.早.40.0-53.0mm.
Figs.18-28(onp355). Spinimegopisspp;18-23,habitus. - 18. S. ,rlalasiaca male. - 19.S.
1ividipennis male from Yunnan. - 20-21. S ftijital sp nov: 20,maleholotype, 21, female
paratype, both fromVietnam. - 22-23.S delahayei sp nov: 22, male holotype,23, female
paratype, both from Myanmar. - 24-28. Lateral view of prosternum. 24, S. lividipe,ulis,
female, 25,S malaslaca,male,26-27,S. /iujltaisp nov: 26,male,27, female, 28,S dela11ayel
male.