A study of family Polythoridae, with details on the genus Euthore Selys, 1869 (Zygoptera)

14 Pages·1992·1 MB·English

by  Bick, G H

Odonatologica21(3):275-288 September I.1992
A study of familyPolythoridae, with details on
the genusEuthoreSelys, 1869
(Zygoptera)
G.H. Bick¹ and J.C.Bick¹
1928 SW48Avenue, Gainesville, Florida 32608, United States
Received November 19, 1991 /Revised andAcceptedMarch 2, 1992
The 8 spp. ofEuthore arekeyed, discussed andtheir wingcolor patterns figured
and tabulated,E. fasciata (Hag.) is polymorphic with3 forms: sulfurata,fasciata.
plagiata,noneofwhicharevalidspeciesorsubspecies. The3commongenera;Cora,
Euthore, Polythoreare compared in detail and a key to males ofall 8 polythorid
genera is given.
INTRODUCTION
We continuehere our studies ofPolythoridae with detailed considerationof
thegenusEuthore.Afterthisatabularcomparison ofthe3 mostcommongenera,
Cora, Polythore, Euthore,is given followedby akey toall8 generaofPolythori-
dae.
THE GENUS EUTHORE
SELYS(1869) establishedthegenusEuthorewithfasciataas thetype species.
MONTGOMERY (1967)keyed 9species and detailedtheirtype depositions. DE
MARMEES (1981, 1982, 1988a)consideredtaxonomy, ecology, distributionof
Venezuelan species and BICK & BICK (1990) removed Euthore terminalis
(McL.) from Euthore, returning it to Cora.
The 8 species inEuthore with theoccurrence ofeachby country are:
fasciata (Hagen in Selys, 1853):Venezuela,Colombia, Ecuador.
fassli Ris, 1914: Col.
1 Research Associate, Florida Slate Collection ofArthropods,Gainesville, Florida, USA.
276 G.H. Bick & J.C.Bick
fastigiata (Selys, 1859):Venez. Col.
hyalina (Selys, 1853): Guyana, Venez., Col., Peru.
inlacteaCalvert, 1909:Peru.
leroiiRis, 1918: Col.
meridanaSelys, 1879:Venez.
mirabilisMcLachlan, 1878:Ec.
We attempted to study specimens in most ofthe large Odonata collections,
but, except for E.fasciata, disappointingly few were available, a contrast with
otherPolythorid generawhichwepreviously studied.However,B.E. Montgome-
ry’s unpublished notes andfigures, mostlyoftypes, wereavailableintheArchives
of the InternationalOdonata Research Institute, Gainesville, Florida, and we
examineddirectly types offasciata, plagiata (Selys) andmirabilis.
Abbreviationsused are:
ANSP Academy ofNatural Sciences, Philadelphia
BMNH British Museum (NaturalHistory), London
CC Carl Cook,Center, KY
DM J. de Marmels, Maracay,Venezuela
FO FelipeOvale
FSCA Florida State Collection ofArthropods,Gainesville
IRSN InstitutRoyal des Sciences Naturelles, Brussels
MCZ Museum ComparativeZoology, Harvard Univ., Cambridge
R.G. R.W.Garrison, Azusa, CA
SMF SenckenbergMuseum, Frankfurt
WCM W.C. MacIntyre
UMMZ University ofMichigan,Museum ofZoology,AnnArbor
USNM National Museum, Smithsonian Institution,Washington,D.C.
Becausefemalesof3speciesremainundescribedandweknowofnodescription
ofthelarval stage,thispaperdealsprimarilywithmales.Thesequenceofspecies
is alphabetical withoutindicationofrelationship. Basic collectiondata are given
in thefollowing sequence:COUNTRY (caps.), department,province, or state(ita-
lics), collector or source, date, numberand sex, deposition.
SPECIES KEYTOEUTHOREMALES
The penis horns of 5 species figured by KENNEDY (1919) as well as the
horns of2 figured by DE MARMELS (1988a) are similar. Ourexaminationof
all 8 species showed thepenis tobe similar, the apical horns ofmost measuring
0.20or0.25mm. We donot considerthepenis a good diagnosticcharacter.Also,
because species differenceswere not found in either abdominal appendages or
body colourpattern, the key is basedentirely on wing colorpattern. Determina-
tions shouldbe confirmedby species accounts, figures andtables.
I Wingshyalineexcept forayellow tintalongthe costa hyalina
1’ Wingseitherconspicuouslyorange-redortransversely bandedwithbrown-black 2
2 Eachwingwith anextensive orange-red triangulararea mirabilis
Study ofPolythoridae 277
Figs 1-8. Diagramsofwingcolor patterns in darkbanded speciesofEuthore: (1)h.wfasciata,form
"fasciata”; - (2)h.w.fasciata,form ’’plagiata”; - (3)f.w.fastigiata;— (4)h.w.fastigiata; -
(5)h.w. inlactea; —(6)h.w.fassli;— (7)h.w. leroii; —(8)h.w. meridana.[Dashes: basaltranspa-
rency; — clear; opaque pale; — black: dark brown orblack; - dots:apicaltransparency].
2’ Each wingwitheitheratransverse darkbandora narrowapicaldark area 3
3 Dark wingband bordered proximallyby anopaque white-yellowor orange area 4
3’ No opaquepaleareaproximadofthe dark band (Fig. 5) inlactea
4 Wing transparentgoldenbrown withanarrowopaqueorangebandbordered
distally byanarrow darkone(Fig.6) ,fassli
4’ Wingnot goldenbrown but withan opaque white-yellow and adark area 5
5 Front wingwith dark areaonly 1.5-2.5 mm wide,usuallyonly anarrow
apicalcrescent (Fig. 3) fastigiata
5’ Front wingwith adistinct dark band which isnevermerelyonly anapical
crescent 6
278 G.H. Bick & J.C. Bick
6 Hind wingdarkband beginning 9-12 mm distad ofnodus (Fig. 8) meridana
6' Hindwingdark bandbeginning1-7mm distadofnodus 7
7 Hindwing with a3-8 mm apicaltransparent areadistad ofthe darkband(Figs 1,2) .. fasciata
T Hindwingwithoutanapicaltransparent area,thedarkbandreachtingtheapex(Fig.7) Ileroii
EUTHOREFASCIATA (HAGEN IN SELYS)
Figures 1.2; Tables I.II
Thorefasciata HAGEN in SELYS. 1853: 70 (Venez.; d, 2 descr.); - SELYS,
1854: 259 (Col.. Venez.: Puerto Cabello; addit. d, 2 descr,).
Euthorefasciata:SELYS. 1869: 32(Venez,;addit.â. 5 descr.,genusestab„/a.«70/a
typesp.); - McLACHLAN, 1878:86(Ec.; 2 6 descr.); - RIS, 1918: 27, 188(Col.:
Rio Negro.Casanara, Sosomuco, Medina, Villavicencio, 450-1300 m; <3, 9 descr.,
key); - KENNEDY, 1919: figs 21,22 (Col.; "genotype”,penis); - MUNZ, 1919:
fig. 3 (h.w. 9); RACENIS, 1953: 28 (Venez.:Puerto Cabello; fig. 9. 6 wings,
venation only); - MONTGOMERY, 1967: 125, 127, 136, 147(Venez.;key, type 6,
9,No. 12116,MCZ); - DEMÄRMELS, 1981: 26,28(Venez.;ecol.,color photo<3).
Euthoref.fasciata: RACENIS, 1958: 202 (Venez.:Cumbre de Choroni, Rancho
Grande, 1000-1400 m); - DE MARMELS, 1982: 39 (Venez.: Aragua,Fed. Disk,
Miranda, Zulia, 1000-2000 m; ecol., behavior); —l9BBa:93 (Venez.:Cumbre de
Choroni. 1200 m; fig. 5,penis, fig.6, photowhole insect); —l9BBb: 380(Venez.:
RanchoGrande; fig. 3, superiorappendage,fig. 6, penis); - 1990: 335 (Venez.).
EuthoreplagiataSELYS, 1873: 37 (’’raced*E.fasciata?”) Rio Negro,Amérique
équatoriale; 1 $ descr., 6 unknown); 1879: 56 (Ec.;probably âofraceplagiata,
1S descr.); - MONTGOMERY, 1967: 125, 148(Rio Negro,Brazil;type 5BMNH;
syn. ofE. fasciâta)\ - KIMMINS. 1970: 200 (holotype 2,BMNH).
Euthoref.plagiata: DE MARMELS, 1988a: 93(Venez.: Loma de Rio, Palmira;
fig. 4, penis, fig. 6, photowhole insect, cf.E. f.fasciata; McLACHLAN’s<1878),
RIS’(1918)specimensassignedhere; MONTGOMERY’S(1967)Brazilprobablyinac-
curate).
Euthorefasciataformsulfurata:DEMARMELS, 1982: 40(Venez.: CoastalCordil-
lera; d with butter yellowwingbars).
Materialexamined
Form fasciata: — COLOMBIA,Cundinamarca, Bogota, P.Mays, VII1-I983, 2 d, RG; - I d,
ANSP; - Lindig. 1862,I d, MCZ; - ’’Appun", I d,MCZ; - COLOMBIAonly, FO. 16d,2
2.FSCA.— ECUADOR,Manabi,Chone, VIII-1939. 1 d.FSCA; - Morona-Santiago,RioBlanco,
WCM.V-1936. I d. X-1939, 2 d. UMMZ; - Rio Napo.WCM, 1 d,UMMZ; - Rio Pastaza,
WCM,X-1936. I d,UMMZ; - Rio Upano,L. Alonzo, I d.UMMZ;- Napo,Cotundo, VI-1975,
R. LaFebre, I d,CC. - VENEZUELA, Aragua,CumbredeChoroni,Rosales,V-1953, I 2,FSCA;
- T.Donnelly. VII-1989. 2 d, J.Michalski; - F. Test,VII-1955, 1 d, 1 2,UMMZ;"CFR",V-
1969. 1 d.FSCA; - Carabobo,Puerto Cabello,? Thoony,3 d, MCZ; - DE, Los Vemados, IV-
1939, 1 d,"CJR", X-1958. 2 d.FSCA; - Miranda, Curupas.G.Vivas. 1X1937,I 2, FSCA; -
Yaracuy, La Puerta. F. Fernandez, VII1-I945, I d, CC: — J. Osario, XI-1981, I d, FSCA; —
VENEZUELA only, I 6.2 2,MCZ.
Formplagiata:- COLOMBIA.Casanare,Fassl, 11-1911,I d.I 2,UMMZ; — Cauca,Popayan,
III-1912. 1 2, FSCA; — Meta,Villavicencio, VIII-1913, 2 d, 1 2, UMMZ; — "Monitz", I d.
Study ofPolylhoridae 279
MCZ; - COLOMBIAonly, FO, 9 9, FSCA. - ECUADOR, Morona-Santiago,Mangosisa.L.
Alonzo, 2 (J, I 9,UMMZ. - VENEZUELA, Aragua,Colonia Tovar.RG,IX-1980. 2i.RG; -
Tachira, SanCristobal,Y.Belandia,IV-1987, 2 6, J.Michalski. - COUNTRY ?. 1 6. UMMZ, I
6.BMNH.
Form sulfurata: - VENEZUELA,Miranda,QUEBRADA PASQUIRE.DM. X-1980, 1 6.RG;
- AVILA, DM, VI-1981, 1 6, X-1981, 1 <5, RG; - DM, X-1983. 1 <J. UMMZ.
Type data —E.fasciatalectotype6 No. 12116in MCZ.Specimenswellpreserved,wingpattern
definitebutbody colorabnormally dark. Detailsofwings given inTable I.
Head dorsally black except for4small palespots. Prothorax black, a meresuggestion ofpale
spots on middlelobe, Pterothorax black with the usual5 palestripes, anterior onesnarrow,others
scarcely detectable.Legs,abdomenandappendagesentirely black.
E.fasciolaparatypc 9No. 12116in MCZ.Abdomenmissing,head andthoraxabnormallydark.
Front wings completelyhyaline,notbanded asin plagiata.Detailsofwings givenin Table 1.
Headblackdorsally with4smallyellowspots.Midlobeofprothorax withlarge pairedpale spots
narrowlyseparatedbyblack. Pterothorax withnarrow, scarcely visiblepalestripes.Legsblack.
E. plagiataholotype $ in BMNH. Specimen in good condition, wingcolor pattern definite,
abdomenrepaired.Wingsbanded, frontonesconspicuouslysoincontrastwith entirelyhyalineones
offasciata;opaquewhitebegins proximadofthenodus, notat itas infasciola.Otherwingdetails
given inTableI.
Head dull black dorsallywith 4small yellowspots.Middle lobe ofprothoraxwith 2 largedorsal
yellowareasseparatedbyblack. Pterothoraxblack with theusual 5 yellow stripesand anadditional
TableI
Characteristics (mm)ofwingsofEuthorefasciata(lectotype 6,paratype 9,MCZ) and E.plagiata
(holotype9, BMNH)
Wle..tntcsisgntghfi Bn-oWta8doasouies$ |Ns~ttoSI3i“dgoumsa |An|tecnodals PoIVs|tnocda.ls .S-£t1igma B-tari8Ia3sn:,asl- pICarency 2IWw-Bihdsitthe Bb.|ero>Sgw1inn1s1bneo&Cydounsd ob-W£froCiIocdwitnfh tc8aorsta -A—triICgapni3sc-.al p>Iare*ncy
S03p1?eOcioe'sy. 03 Z < cu w ffla cq^^-a;t<q.
fasoiclia|taCi
rlTecntotypTDen> 6o*
f.iws. O3J1.6O' £14.6O' 1u3>b.0 U3)800 44^O0 U3»b.0 44^b.0 1144..00 4.b0 1t2o.t2o to2.1
hO'.iws. 3U)0O.7 1to2b.0 1u3>.7 t2o800 44^O0 2JO.7 44ob.0 1y3j.1 '-/5>b.0 11.O6' t2o.1
fas»)cSia'tsa
p■aaarraattyyppee 99
f-ti.,ws. U3)0.8oc 1t2oL.3*j 1U3»b.0 U3)t2o 3OJ9sC u3>.1
hO'.iws. t2o9sC.1— -11.O6' 1U3>L.5/t t2o7-o O3J9sO O3J.b0 11.O6' ■4P*.b.0 44ob.0 44^.L3*j 080.l/5i
p-3llaaggiiaatlaa
h3-oolloottyyppee 99
f.w. to2O6'.b0 10.O6' 1to2.b0 t2o7-0 U3>1 t2o.7 8ocb.0 '5-/I.b0 t2o.L5ti U5»b.0 p8c.L5/i
=hr.jwS. tO2O6'.7 9)C.l5/> 1t2o.I5a to2U3» U3)2to t2o.7 7.l5/l 7.Ö0 44^.b,0 6o.b0 6O'.7
280 G.H. Bick & J.C, Bick
half-lengthmesepisternalone.Alllegsmissing.Abdomen black,yellowlateral stripesonsegments
2 and 3,4unmarked. 3 and 5 each with asmall lateral basal yellowspot.
SELYS (1873, 1879)stated thatthefrontwingsofplagiatafemales are banded,those offasciata
hyaline.This differenceis obvious in thetypesofthe taxa which westudied, but MONTGOMERY
(1967)statedthatthesefemaletypes,presumablythe samespecimens,wereidentical andthatplagiata
isasynonym offasciala.SELYS(1873, 1879)gaveverybriefnotes forhis Imaleplagiatasurmising
thatitwasonlyavarietyoffasciala.There isnomaletype ofplagiatabutwe studiedthe maletype
offasciala. DE MARMELS (1982, 1988a, 1990)assigned Venezuela specimensto E. f.fasciata
(northern).E.f.plagiata(southern)and toformsulfurata(2 small northernpopulations). However,
all 3 ofhis”plagiata”females hadhyaline frontwings (DE MARMELS, 1988a,p. 93) as inthe
fasciatatype.
Our materialofE.fasciata included2 kinds ofreadily distinguished females:
those with hyaline front wings as in thefasciata type, those with banded ones
as in the plagiata type. The hind wings of both kinds were very similar and
overlapped in all measurements.
Specimens ofE.fasciala includedmaleswhichdifferedfromall othersby the
opaque yellow or cream wing color andmales in which theopaque wing color
was whiteand the wing color patterns very similar. Distinction between these
latterwasmanifestonly whenmaleswere foundassociatedinthesamecollection
with the 2 very differentkinds offemales. One kindofmalewas collectedwith
femaleshaving frontwings as inthefasciata type,theother withfemaleshaving
front wings as in theplagiata type. Measurementsof hind wing color patterns
ofvarious populations (Tab. II), showedthat these2 kinds ofmalesoverlapped
in mostcharacters. Thus,E.fasciata is polymorphic with 3 forms.They are only
Table II
Acomparisonin mm ofhind wing color pattern of3forms ofEuthorefasciatamales
Form Area No. Width of Dark begins Width of Widthof
opaque disladof dark apical
pale nodus at costa transparency
"SULFURATA" Venezuela 3 8-10 3-4 9-11 3-4
"FASCIATA" Colombia 19 5-9 3-6 6-9 3-5
Venezuela 5 7-11 2-4 8-11 3-5
Ecuador 5 4-6 2-3 9-10 3-6
Associated with
hyalinef.w. 5 Venez. 1 7 2 9 5
"PLAGIATA"
Associated with
banded f.w. 2 Colombia, Ecuador 2 6-7 3-3 5-9 5-5
Venezuela, southern 2 6-6 1-2 8-8 8-8
Venezuela, northern 2 6-7 l-l 9-9 6-7
StudyofPolythoridae 281
formsbecausewedidnot detectgeographical orecological isolationnorstructural
differencesand because ofoverlap (Tab. II) in so many wing characters.The 3
formsaredesignated:.)1)sulfurata,following DE MARMELS(1982),(2)fasciata
because ofassociation withfemales having colorless front wings as inthe type,
(3) plagiata from associationwith females having bandedfront wings as in that
type.
Formsulfurata is represented by males only, all fromMirandaState,northern
Venezuela.Theusually pale wing area is sometimes clearly yellow as in the DE
MÄRMELS(1981) photo, sometimesonly cream-colored.
Formsfasciata andplagiata occur inVenezuela,ColombiaandEcuador,both
forms having been collected from the same localities. Because the male hind
wing color patterns overlapped in most characters (Tab. II), some specimens
could not be assigned confidently to either form. Indeed, without associated
females, placement ofmost males in either form is risky. The best diagnostic
character isthe slightly larger(Figs 1,2; Tab. II) apical transparency in thehind
wings of some specimens offormplagiata.
Despite theobviously greatdifferenceinfrontwings,thehindwings offemales
ofthe 2 forms were very much alike, overlapping in every wing color pattern
studied. This hind wing similarity in both sexes supports our hypothesis that
neitherform is a separate species or subspecies.
In each of2 collections(Colombia,Ecuador) a male was associatedwith a
bandedfront wing female.Inanother fromVenezuelaamalewas associatedwith
a hyaline front wing female. The fact that all 3 males (Tab. II) are essentially
alikeshows thatvery similarmalesoccur closely associatedwith2 very different
kinds offemales.Twokinds of femaleswithinwhatseemstobethesamespecies
is similarto theconditiondescribedforPolythoreproceraSelys (BICK& BICK,
1985)and reported for Calopteryx spp. (DE MARCHI, 1990).
EUTHOREFASSLIRIS
Figure6; Table III
EuthorefassliRIS, 1914: 282 (Col.:Monte Socorro,Santa Margarita2300 m;6 6
descr., wingsfig. 1 photo); — 1918: 27, 188 (Col.;key, 1914photoinaccurate); —
MONTGOMERY, 1967: 125, 136, 147(Col.;key, lectotype SNo.417SMF).
Materialexamined. — COLOMBIA,Amazonas,Leticia,C.Farrell, I 6, RG.
This specimen was collectedfareast ofthose reported by RIS (1914, 1918).
The opaque pale band is orange, exceptionally narrow (Tab. Ill) and almost
pointed at itsanal end.
Thefemaleis unknown.
282 G.H. Bick & J.C. Bick
EUTHOREFASTIGIATA(SELYS)
Figures3,4;Table III
ThorefastigiataSELYS. 1859: 450 (Bogota; 3 descr.. 9 unknown).
Euthorefastigiata:SELYS, 1869: 33 (Bogota; 3 descr.); - RIS. 1918: 28. 188
(Col.: Bogota. Rio Negro.Sosomuco, Medina, Quetoma. 800-1600 m; key. 3. 9
descr..fig. II 3 wings,ecol.); — KENNEDY, 1919: figs 23. 24 (Col: Tosomorio;
penis); - MONTGOMERY. 1967: 125, 136, 147 (Col.;key, types 5 3 IRSN, I 3
BMNH).
Euthoref.fastigiata:DEMARMELS, 1988a:93,fig.6(Venez:Tachira.Los Delicias,
17-1800m;photo,cf. E.f. meridana,Bogota illogicaltype locality); - 1990: 335
(Venez.),
Material examined. - COLOMBIA,Cundinamarea ,Bogota, 2 6 ,ANSP; — Lindig, 1862,
4(5.MCZ;—Meta,Villavicencio,VI11-I913, I 3,UMMZ; — RioMeta,3 3.BMNH; — Sosomuco,
Fassl. III-19II,2 3.UMMZ. - COLOMBIAonly, FO, II 3.FSCA, I 3. UMMZ; - Sjosted,2
3. UMMZ. - VENEZUELA, Tachira,Rubio-Delicias. IX-1966, 1 3.FSCA.
DE MARMELS (1988a)disagreed with Selys’ designation ofBogota as the
type locality, but we do have 6 specimens from there.
E. meridana has been designated a race or subspecies offastigiata (SELYS,
1879; RACENIS, 1958;DE MARMELS, 1988a). The hind wings of both are
similar(Figs 4, 8; Tab. Ill), but we judge each to be a separate species (as did
TableIII
Acomparisonin mm ofhind wingcolor patterns ofmales of5 species ofEuthore having dark-
bandedwings
B
costa
sz opaque b.eSgins.no2dus P t/5 s£tigma
ö leWnggt»h tr|acnsparencoy « of _at *en§ds.2Î* Ictransparency
S05p1e?ncine>'sSO Ga?r3eeoÛoa3rgrora■p33h-ioc’ iNzu$m£ber 5Wÿing B_caSgsal §ag. 5Wx«id:th jpaajle. BarEoowänb1e£yon:d SiW:i£dlth bJr>oow5n Bm>orownb£e£fore A-<«pgicalSac.
fÖa*sSotigSia’sta nCol5o*m3bgi;a5’ 1o6
r3an3gfroe«D 2288--3333 800-11 313j-1800 080-1£4 •4f-t6O' 1--k4 o0-1
Imea33n 3300..44 9■pk.2) 1’-5O.44^ 1p0b.8o 5Oi.44^ 1‘v.O9 p0.1
ImerIidanSa V£e3nne>NzuÎeLl0a3 9sC
r3an3g0Cea 2299--3322 99--1100 1U3»-1L7j '9P-1K2) ’5-/t--L7i 0o 0o-1
Imea33n -31.1 9sO.1 1y5*.1“ 510U.3) 6O'.44^ 0O 0o.6O'
f/ausssi/li CnolÔom3bg;i3a‘ I- K2)s9C 1L3k> h2J.ly5» 1k.7j 44^ 6O'.k7l 1o0
inlSa’ctea P-ooeruc 1- K2>6O' 11 0o 1.44^ 9vO 1.44- 6O'
leroii Cooloo'm3bgi;5a’ 1- 3o0 6O' 13 7-J.0 1K2) 0 0o
SludyofPolylhoridac 283
MONTGOMERY, 1967)because ofthe great and nonoverlapping differencein
widthofthefrontwing black: 1.5-2.5mmfastigiata; 4.5-5.5 mmmeridana.Also
the widths of dark in meridanafront and hind wings are similar whereas these
widths differgreatly infastigiata.
R1S (1918) stated that the female front wing is hyaline, the hind,banded.
EUTHORE HYALINA (SELYS)
ThorehyalinaSELYS, 1853: 71(Bahia; 3 3,descr.);— 1854:261 (expandeddescr.).
Euthore hyalina: SELYS, 1869: 33 ("Bahia ??.”, Bogota; note on 2); — RIS,
1918: 25, 26, 188 (Col.: Muzo, Pacho. Bogota;Peru: Cuzco, 400-2300m; key, 3, 2
descr.); - KENNEDY, 1919: figs. 25,26(Bogota;penis); - RACENIS, 1959: 487
(Peru: Santa Ana); — MONTGOMERY, 1967: 125, 127, 135, 148(Brazil;key, lecto-
type 3 IRSNdesig,).
Material examined. — COLOMBIA, Cundinamarca, Bogota, 1911, 2 3, UMMZ; I d.
I 2,ANSP;Pacho,22.BMNH; - COLOMBIAonly.FO, 1 d.FSCA;1 6,BMNH. - GUYANA,
Demarara, E.Klots, IV-1937, 1 3, FSCA. - "NEWGRANADA", 2 <5, BMNH. 1 3,ANSP. -
VENEZUELA, Lara, Cubiro,Y.Fernandez, 1-1954, I 2,FSCA.
SELYS (1853, 1854)listedBahiabutin 1869questioned that locality. MONT-
GOMERY (1967), possibly thinking ofBahia state,gave Brazil. Ifthisis indeed
the Selys locality, it wouldbe far east ofall records ofthe genus. The correct
locality maybeBahia deCaraquez on thePacificcoastofEcuador. Nevertheless,
E. hyalina is widespread, reaching Guyana andsouthern Peru.
EUTHOREINLACTEA CALVERT
Figure5; Table III
Euthore fasciata inlactea CALVERT, 1909: 91 (Peru: Piches, Perene Valleys, 2-
3000’; 2 3 descr.); - KENNEDY, 1920: figs24, 25 (penis); - RACENIS, 1959:
486(Peru: Piches, Perene).
Euthoreinlactea: KENNEDY, 1919: figs27, 28(Peru;penis); — MONTGOMERY.
1967: 125, 136, 148(Peru;key, type, USNM, 1 d paralype ANSP).
Material examined. — PERU, Loreto, Balsapuerto,P. Nagel, VI-1933, 1 3. FSCA. (S.W.
Dunkle(pers.comm.) examineda type 3 in USNM|.
This new locality, the first since the original, is farto its north. E. inlactea
(Fig. 5) may suggestfasciata (Figs 1,2)except fortheabsence ofwhite andthe
greaterwidthof black in hind wings oftheformer.
The female is unknown.
284 G.H.Bick & J.C. Bick
EUTHORE LEROIIRIS
Figure7; Table III
Euthore leroii RIS, 1918: 28, 188 (Col.: Canon del Tolima, 1700 m; 2 6. I 2
desc.,key); MONTGOMERY, 1967: 126, 136, 148(Col.; key, type No.418 6, No.
4202, SMF),
Material examined. — COLOMBIA,Caldas, Pensilvania, 1913, 1 â, UMMZ.
TheCaldaslocality, the only one additionaltothetype, is atno greatdistance
from theoriginal. Wings ofleroiihave beenconsidered(RIS, 1981;MONTGO-
MERY, 1967)exceptionally broad,but those of some specimens offasciata are
almostas wide. The2 species are readily distinguished: the hindwing darkband
of leroii (Fig. 7) reaches the apex, whereas fasciata (Figs 1, 2) has a 3-8 mm
clear apical area.
RIS (1918)stated thatboth frontandhind wings ofleroiifemalesarebanded.
EUTHORE MERIDANASELYS
Figure 8;Table III
Euthore meridana SELYS, 1879: 58 (Venez.: Merida; o adulte, jeune descr., V
unknown,prob,race offastigiata); MONTGOMERY, 1967: 126, 136, 148(Venez.;
key, type 6 IRSN); DEMARMELS, 1982:39 (Venez.: Merida, Lara; ecol.).
Euthorefastigiatameridana:RACENIS, 1958: 202(Venez., Lara: Cubiro,Sanare,
Sierra Terepayma, 1300-1600 m); - DE MARMELS, 1988a:94 (Venez.: Merida,
Barinas, Lara; fig. 6, photoentire insect); - 1990: 335 (Venez.).
Material examined. — VENEZUELA, Barinas. San Isidro, J.S.VII-1975, 3 6, 1 2, FSCA;
— Lara,Carret, Humocaro Alto, 11-1980,2 <J, CC; — Sanare, Gutierrez & Gonzalez,VII-1980, 1
d,FSCA; - J.Osorio, III-198I,3 <J, I 2,FSCA; - Merida,San Jacinto,Adams& Bernard, VI-
1975, 1 <J, FSCA.
Although meridanasometimeshasbeen consideredasubspecies offastigiata,
thereis littlepossibility ofconfusing thetwo becauseofthe muchmore limited
extent ofthefront wing dark infastigiata. E. meridanadiffersalso from leroii
andfasciata: in meridanathehind wing (Tab. ill) dark band is much narrower
(5-7 mm) than in leroii(12) and inmeridanathe widthof white(13-17 mm)is
greaterthan infasciata (4-11).
Ourspecimens include2femalescollectedwithmales.ThefemalefromBarinas
has hyaline frontwings, whiteanddark-bandedhindones. Thesomewhatimma-
ture Larafemalehas hyaline frontwings, only amere traceofwhite inthehind.
EUTHORE MIRABILISMcLACHLAN
EuthoremirabilisMcLACHLAN 1878: 87 (Ecuador: Intag;6 d,6 9, descr.); —
SELYS, 1879: 57(Intag; d,9 descr.); - KENNEDY, 1919(Ec.; figs 29, 30,penis.