A splitter's guide to the higher taxa of the flowering plants (Magnoliophyta) generally arranged to follow the sequence proposed by Thorne (1992) with certain modifications

61 Pages·1993·12.5 MB·English

by  James L Reveal

Phytologia(March 1993) 74(3):203-263.
A SPUTTER'S GUIDE TO THE HIGHER TAXA OF THE FLOWERING PLANTS
(MAGNOLIOPHYTA) GENERALLY ARRANGED TO FOLLOW THE SEQUENCE
PROPOSED BY THORNE (1992) WITH CERTAIN MODIFICATIONS
James L. Reveal
Department ofBotany, University of Maryland, College Park, Maryland
20742-5815 U.S.A.
ABSTRACT
Using the most recent system of classification for flowering plants
(Magnoliophyta) proposed by Thome (1992b) and generally following
his sequence ofnames but with the addition ofthe family names now
in current use, afamily systemis proposed that purposefully splits the
subclasses, superorders, orders, and families into snuiU units thereby
constructing a "splitter's" guide to the higher taxa ofMagnoliophyta.
Theresultingclassificationrecognizes 14subclasses, 63superorders,248
orders, zind 685 families.
KEY WORDS: Magnoliophyta, nomenclature, classification.
INTRODUCTION
Since 1959, considerable attention has been given to the classification of
the flowering plants, with a large number oftaxa recognized (Bedell & Reveal
1982a, b; Benson 1957; Boivin 1956; Brummitt 1992; Cronquist 1957, 1961,
1968, 1981, 1988; Cronquist et al. 1966; G. Dahlgren 1989a, b; R. Dahlgren
1975, 1980, 1983; R. Dahlgren k Bremer 1985; R. Dahlgren & Clifford 1982;
R. Dahlgren et al. 1981, 1985; Deyl 1955; Dostal 1957; Ehrendorfer 1983; En-
gler 1964; Erdtman 1952, 1966; Gibbs 1974; Goldberg 1986, 1989; Gunn et
al. 1992; Heywood 1978; Huber 1969; Hutchinson 1959, 1969, 1973; Kimura
1953, 1956; Mabberley 1987; Novak 1954, 1961; Pulle 1952; Rouleau 1981;
Soo 1953, 1961, 1967; Stebbins, 1974; Takhtajan 1959, 1970, 1973, 1980, 1983,
1985, 1986, 1987; Thorne 1974, 1976, 1977, 1981, 1983, 1992a, 1992b; Willis
1973). Concomitant withthis attentionhas beentherealization that botanical
203
PHYTOLOGIA
204 volume 74(3):203-263 Marcii 1993
nomenclatureought to be stabilized as much as possible insofar as nomenclat-
uraJ matters are concerned. As a result, the botaniczJ community is now
considering the concept of "names in current use" (NCU), and to that end,
lists of proposed names have been circulated for comment in anticipation ofa
formal publication ofprotected names in early 1993.
My own efforts on this task have concentrated on vascular plant family
names in collaboration with Dr. Ruurd D. Hoogland of the Laboratoire de
Phanerogamie, Museum National d'Histoire naturelle in Paris. To ascertain
the taxonomicimpact ofprotectingfamilynames, we haveworked closely with
Richard K. Brummitt (K), the late Arthur Cronquist (NY), Aaron Goldberg
(US),ArmenL. Takhtajan(LE),Robert F.Thorne (RSA),andJohn Wiersema
(USDA). To that end we have been able to resolve potential nomenclatural
conflicts. The need to resolve the nomenclaturaJ morass above the rank of
family is no less important than that at the family rank and below, and while
priority is not required above the rank of family, the need to have Aralidly
published names is mandated (Reveal 1992a, 1992b, 1993) and the need for a
stabilized nomenclature even at these higher ranks is gradually being recog-
nized.
In the following summary ofthe flowering plants (Magnoliophyta), I have
adopted thegeneral schemeproposed by Thorne (1992b), and included therein
ail of the proposed NCU family names. To this I have added class, subclass,
superordinal, and ordinal names. I have adopted 1966 as the starting date
for subclass and superordinal names, and 1789 as the starting date for ordinal
and family names. To Thome's linear arrangement of superorders, orders
and families, I have added the rank ofsubclass used by Cronquist (1981) and
Takhtajan (1987). Furthermore,as thefamilylistaccountsforallfamilynames
now in current use, Ihave also attempted to includeaJl subclass, superordinal,
and ordinal names now in current use. The philosophy behind the application
ofpriority to these higher names is outlined elsewhere (Reveal 1993).
The consequences of adopting, or not adopting, the proposal put forth by
Greuter(1991) regarding "namesincurrentuse" (NCU)arenot allthat serious
insofar as flowering plant fsimily names because so many of these names are
already protected (Reveal & Hoogland, 1991). Conservation, as traditionally
appliedto floweringplant family names, has been to ensurethat certain names
areusedinsteadofothers,but not authorshiporplaceofpublication. Greuter's
proposal would now protect the bibliographic information as well. Should the
NCU proposals fail, we will have to editorially correct the authorship and/or
bibliographic references of nearly 125 currently conserved family names on
Appendix IIB of the Code (Greuter et al. 1988). And, as may be seen from
the catalogue below, ifthe NCU proposals fail there will be a need to conserve
at least ten additional names just to retain certain nonconserved names now
in current use.
Reveal: Guide to higher taxa ofMagnoliophyta 205
CATALOGUE OF THE HIGHER TAXA OF MAGNOLIOPHYTA
Magnoliophyta Cronquist, Takhtajan, k Zimmermann, 1966
Vlagnoliopsida Cronquist, Takhta- Monodoraceae3 Agardh, 1858
.
jan, & Zimmermann, 1966 6. AristolochialesDumortier, 1829
I. Magnoliidae Novak ex Takhta- Asarales Burnett, 1835
jan, 1967 11. AristolochiaceaeA.L.Jussieu,
A. Magnolianae Takhtajan, 1967 1789, nom. cons.
1. Winterales A.C. Smith exRe- Asaraceae Ventenat, 1799
veal 12. Myristicaceae R. Brown,
1. Winteraceae R. Brown ex 1810, nom. cons.
Lindley, 1830, nom. cons. 7. Canelljdes Cronquist, 1957
2. Takhtajaniaceae (J. Leroy) 13. Canellaceae C. Martius,
J. Leroy, 1980 1832, nom. cons.
2. Illiciales H.-H. Hu ex Cron- WinteranaceaeWarburg, 1895
qtiist, 1981 8. Austrobaileyales Takhtajan
3. Illiciaceae (Candolle) A.C. ex Reveal, 1992
Smith, 1947, nom. cons. 14. Austrobaileyaceae(Croizat)
4. SchisandraceaeBlume, 1830, Croizat, 1943, nom. cons.
nom. cons. 9. Monimiales Dumortier, 1829
3. Magnoliales Bromhead, 1837 15. AmboreilaceaePichon, 1948,
5. MagnoliaceaeA.L.Jussieu, nom. cons.
1789, nom. cons. 16. Trimeniaceae (Janet R.
6. Liriodendraceae Barkley, Perkins& Gilg) Gibbs, 1917,
1975 nom. cons.
7. Degeneriaceae I. Bailey &; 17. Hortoniaceae (Janet R.
A.C. Smith, 1942, nom.. cons. Perkins) A.C. Smith, 1971
8. HimantandraceaeDiels, 1917, 18. MonimiaceaeA.L.Jussieu,
nom. cons. 1809, nom. cons.
4. Eupomatiales Takhtajan ex 19. Atherospermataceae R.
Reveal, 1992 Brown, 1814
9. Eupomatiaceae Endlicher, 20. Siparunaceae (A. de Can-
1841, nom. cons. dolle) Schodde, 1970
5. Annonales Lindley, 1833 21. Gomortegaceae Reiche,
10. Annonaceae A.L. Jussieu, 1896, nom,. cons.
1789, nom. cons. 10. Calycanthales C. Martius,
Hornschuchiaceae J. Agardh, 1835
1858 22. IdiospermaceaeS.T. Blake,
A
206 P HYT OL OGI volume 74(3):203-263 March 1993
1972 veal, 1992
23. Calycanthaceae Lindley, 18. NymphaealesDumortier, 1829
1819, nom. cons. Eurydales H.L. Li, 1955
ChimonanthaceaePerleb, 1838 35. Cabombaceae A. Richard,
Butneriaceae Barnhart, 1895, 1828, nom. cons.
nom. illeg. Hydropeltidaceae(Candolle) Du
11. Laurales Perleb, 1826 mortier, 1822
24. Lauraceae A.L. Jussieu, 36. Nymphaeaceae R.A. Sal-
1789, nom. cons. isbury, 1805, nom.. cons.
Perseaceae Horaninow, 1834 37. Euryalaceae J. Agardh,
25. Cassythaceae Battling ex 1858.
Lindley, 1833, nom. cons. 38. BarclayaceaeH.L. Li, 1955
26. HernandiaceaeBlume, 1826, D. Rafflesianae Thorne exReveal,
nom. cons. 1992
Illigeraceae Blume, 1833 19. Hydnorales Takhtajan ex
12. Gyrocarpales Dumortier, 1829 Reveal, 1992
27. GyrocarpaceaeDumortier, 39. Hydnoraceae C. Agardh,
1829. 1821, nom. cons.
13. Chloranthales Conzatti & 20. Mitrastemonales Makino,
L.C. Smith ex Reveal, 1992 1911
28. Chloranthaceae Blume, 40. MitrastemonaceaeMakino,
1827, nom. cons. 1911, nom. cons.
Hedyosmaceae Caruel, 1881 21. Rafflesiales Oliver, 1895
14. Lactoridales Takhtajan ex Cytinales Dumortier, 1829
Reveal, 1992 41. Cytinaceae (Brongniart)
29. LactoridaceaeEngler, 1888, A. Richard, 1824
nom. cons. 42. Apodanthaceae(R. Brown)
15. PiperaJes Dumortier, 1829 Tieghem exTakhtajan, 1987
30. Saururaceae Richard ex 43. Rafflesiaceae Dumortier,
E. Meyer, 1927, nom. cons. 1829, nom. cons.
31. Piperaceae C. Agardh, II. RanunculidaeTakhtajan exRe-
1824, nom. cons. veal, 1992
32. PeperomiaceaeA.C. Smith, E. RanunculanaeTakhtajan exRe-
1981 veal, 1992
B. Nelumbonanae Takhtajan ex 22. Paeoniales Heintze, 1927
Reveal, 1992 44. Paeoniaceae F. Rudolphi,
16. CeratophyllalesBischoff, 1840 1830, nom. cons.
33. CeratophyUaceae Gray, 23. Glaucidiales Takhtajan ex
1821, nom. cons. Reveal, 1992
17. Nelumbonales Burnett. 1835 45. Glaucidiaceae Tamura,
34. Nelumbonaceae(Candolle) 1972
Dumortier, 1829, nom. cons. 24. Menispermales Bromhead,
C. Nymphaeanae Thorne ex Re- 1838
Reveal: Guide to higher taxa ofMagnoliophyta 207
46. MenispermaceaeA.L.Jus- 1847
sieu, 1789, nom. cons. 61. PapaveraceaeA.L.Jussieu,
Pseliaceae Rafinesque, 1838 1789, nom. cons.
47. LardizabidaceaeDecaisne, 62. Platystemonaceae (W.R.
1839, nom. cons. Ernst) A.C. Smith, 1971
48. Sargentodoxaceae Stapf 63. Pteridophyilaceae(Murbeck)
ex Hutchinson, 1926, nom. Nakai exReveal& Hoogland,
cons.
1991
25. Podophyllales Dumortier, 64. HypecoaceaeH.M.Willkomm
1829 k J.M.C. Lange, 1880
49. Podophyllaceae Candolle, 65. FumariaceaeCandolle, 1821,
1821, nom. cons. nom. cons.
DiphylleiaceaeSchultz-Schult- CorydalaceaeGiseke, 1792, nom.
zenstein, 1832
illeg.
50. Leonticaceae Berchtold & in. CaryophyllidaeTakhtajan, 1967
J. Presi, 1820 F. CaryophyllanaeTakhtajan, 1967
26. BerberidalesDumortier, 1829 29. Caryophyllales Perleb, 1826
51. BerberidaceaeA.L.Jussieu, ScleranthaksDumortier, 1829
1789, nom. cons. SUenales Lindley, 1833
52. Nandinaceae Horaninow, Dianthales Burnett, 1835
1834.
66. AlsinaCeae(Candolle)Bart-
27. RanunculalesDumortier, 1829 ling, 1825, nom. cons.
Hellehorales Nakai, 1949) Stellariaceae Dumortier, 1822
53. HydrastidaceaeMartinov, 67. niecebraceae R. Brown,
1820.
1810, nom. cons.
54. Thalictraceae Rafinesque, Paronychiaceae A.L. Jussieu,
1815
1815
55. RanunculaceaeA.L.Jussieu, Scleranthaceae Berchtold& J.
1789, nom. cons. Presl, 1820
Anemonaceae Vest, 1818 HemiariaceaeMartinov, 1820
ClematidaceaeMartinov, 1820 68. CaryophyllaceaeA.L.Jussieu,
56. Helleboraceae Vest, 1818 1789, nom. cons.
Calthaceae Martinov, 1820 Cerastiaceae Vest, 1818
Actaeaceae Rafinesque, 1828 Dianthaceae Vest, 1818
Nigellaceae J. Agardh, 1858 Ortegaceae Martinov, 1820
57. CircaeasteraceaeHutchin- Telephiaceae Martinov, 1820
son, 1926, nom. cons. Saginaceae Sprengel ex Wein-
58. Kingdoniaceae A.S. Fos- mann, 1824
ter ex Airy Shaw, 1965 Silenaceae (Candolle) Bartling,
28. PapaveralesDumortier, 1829 1825
59. Chelidoniaceae Martinov
30. PortulacalesDumortier, 1829
1820
69. PortulacaceaeA.L.Jussieu,
60. EschscholtziaceaeSeringe, 1789, nom. cons.
PHYTOLOGIA
208 volume 74(3):203-263 March 1993
Montiaceae Rafinesque, 1820 1830, nom. cons.
70. Hectorellaceae Philipson FicoideaceaeA.L.Jussieu, 1789
& Skipworth, 1961 Galeniaceae Rafinesque, 1819
71. BasellaceaeMoquin-Tandon, Sesuviaceae Horaninow, 1834
1840, nom. cons. 83. MesembryanthemaceaeFena
Anrederaceae3 Agardh, 1858 1836
.
Ullucaceae Nakai, 1942 MesembryaceaeDumortier, 182S
72. DidiereaceaeDrake, 1903, nom. illeg.
nom. cons. 84. TetragoniaceaeNakai, 1942,
31. Cactales Dumortier, 1829 nom,. cons.
Opuntiales Willkomm, 1854 85. Halophytaceae Soriano,
73. Cactaceae A.L. Jussieu, 1984
1789, nom. cons. 86. Molluginaceae Hutchin-
Opuntiaceae Martinov, 1820 son, 1926, nom,. cons.
Cereaceae CandoUe & Spren- PhamaceaceaeMartinov, 1820
gel, 1821 Corrigiolaceae(Dumortier)Du-
32. NyctaginalesDumortier, 1829 mortier, 1829
Petiveriales Lindley, 1833 Glinaceae Dumortier, 1829
74. PhytolaccaceaeR.Brown, Adenogram.m,acea€ (Fenzl)Nakai
1818, nom. cons. 1942
Sarcocaceae Rafinesque, 1837 Polpodaceae(Fenzl)Nakai, 1942
75. Gisekiaceae (Endlicher) 33. Chenopodiales Dumortier,
Nakai, 1942 1829
76. Petiveriaceae C. Agardh, Atriplicales Horaninow, 1847
1824 87. DysphaniaceaePax, 1927,
Rivinaceae C. Agardh, 1824 nom.. cons.
Hilleriaceae Nakai, 1942 88. Chenopodiaceae Vente-
Seguiertaceae Nakai, 1942 nat, 1799, nom. cons.
77. AgdestidaceaeNakai, 1942 AtriplicaceaeA.L.Jussieu, 1789
78. Barbeuiaceae Nakai, 1942 Corispermaceae Link, 1829
79. AchatocarpaceaeHeimerl, Betaceae Burnett, 1835
1934, nom. cons. Blitaceae Adanson ex Post &
80. Stegnospermataceae (A. Kuntze, 1903
Richard) Nakai, 1942 89. Salicorniaceae Martinov,
81. NyctaginaceaeA.L.Jussieu, 1820
1789, nom. cons. 90. SalsolaceaeMoquin-Tandon,
Jalapaceae Batsch, 1802 1849
Allioniaceae Horaninow, 1834 34. Amaranthales Dumortier,
Bougainvilleaceae J. Agardh, 1829
1858 91. AmaranthaceaeA.L.Jussieu,
Pisoniaceae J. Agardh, 1858 1789, nom,. cons.
MirabilidaceaeW. Oliver, 1936 Celosiaceae Martinov, 1820
82. Aizoaceae F. Rudolphi, AchyranthaceaeRafinesque, 1837
Reveal: Guide to higher taxa ofMagnoliophyta 209
GomphrenaceaeRalinesque, 1837 Planchon) L. Beauvisage ex
DeeringiaceaeJ.Agardh, 1858 Bullock, 1959
IV. Dilleniidae Takhtajan ex Re- 102. Chrysobalanaceae R.Brown,
veal k Takhtajan, 1993 1818, nom. cons.
G. Dillenianae Takhtajan ex Re- Licaniaceae Martinov, 1820
veal &; Takhtajan, 1993 Hirtellaceae Horaninow, 1847
35. DillenialesHutchinson, 1924 103. SymplocaceaeDesfontaines,
92. Dilleniaceae R.A. Salis- 1820, nom.. cons.
bury, 1807, nom. cons. 104. Caryocaraceae Szyszy-
Soramiaceae Martinov, 1820 lowicz, 1893, nom. cons.
HihhertiaceaeJ. Agardh, 1858 RhizobolaceaeC&ndoUe, 1824,
36. Actinidiales Takhtajan ex nom. illeg.
Reveal, 1992 105. Marcgraviaceae Choisy,
93. Actinidiaceae Hutchin- 1824, nom. cons.
son, 1926, nom. cons. 106. Oncothecaceae Kobuski
94. Saurauiaceae J. Agardh, ex Airy Shaw, 1965
1858, nom. cons. 39. Aquifoliales Senft, 1856
37. ParacryphialesTakhtajan ex 107. Aquifoliaceae Bartling,
Reveal, 1992
1830, nom.. cons.
95. ParacryphiaceaeAiryShaw, niicaceaeBerchtoldkJ. Presl,
1965 1820
H.TheanaeThome exReveal, 1992 108. Phellinaceae (Loesener)
38. Theales Lindley, 1833 Takhtajan, 1967
Camelliales Burnett, 1835 109. Sphenostemonaceae P.
96. StachyuraceaeJ. Agardh, Royen k Airy Shaw, 1972
1858, nom. cons. 40. Ochnales Hutchinson ex Re-
97. Theaceae D. Don, 1825, veal, 1992
nom,. cons. 110. LophiraceaeLoudon, 1830
Camelliaceae Candolle, 1816 111. Sauvagesiaceae (Gingins
TemstroemiaceaeMirbel exCan- exCandolle)Dumortier, 1829
dolle, 1816 112. OchnaceaeCandolle, 1811,
Gor(fomaceac(Candolle) Spren- nom. cons.
gel, 1826 GomphiaceaeCandolle exSchni-
MalachodendraceaeJ.Agardh, zlein, 1843-1870
1858, nom. illeg. Luxemburgiaceae Tieghem ex
98. Sladeniaceae(Gilg&Wer- Solereder, 1908
dermann) Airy Shaw, 1965 113. Quiinaceae Choisy ex
99. Asteropeiaceae (Szyszy- Engler, 1888, nom. cons.
lowicz)Takhtajan exReveal 114. Scytopetalaceae Engler,
k Hoogland, 1990. 1897, nom. cons.
100. TetrameristaceaeHutchin- Rhaptopetalaceae Tieghem ex
son, 1959
Solereder, 1908
101. Pellicieraceae (Triana k 115. Strasburgeriaceae En-
210 P HYT L GIA volume 74(3):203-263 March 1993
gler& Gilg, 1924, nom. cons. Airy Shaw, 1965
41. Medusagynales Brenan, 1952 126. Napoleonaeaceae A. Rich-
116. Medusagynaceae Engler ard, 1827
& Gilg, 1924, nom. cona. Belvisiaceae R. Brown, 1821,
42. Ancistrocladales Takhtajan nom. illeg.
ex Reveal, 1992 127. Lecythidaceae Poiteau,
117. Ancistrocladaceae Plan- 1825, nom. cons.
chon eiWalpers, 1851, nom. Gustaviaceae Burnett, 1835
cons. 128. Asteranthaceae Knuth,
43. Dioncophyllales Takhtajan 1939, nom. cons.
ex Reveal, 1993 K. Sarracenianae Thome ex Re-
118. Dioncophyllaceae (Gilg) veal, 1992
AiryShaw, 1952, nom. cons. 48. SarracenialesBromhead, 1838
44. HypericalesDumortier, 1829 129. SarraceniaceaeDumortier,
119. Bonnetiaceae (Bartling) 1829, nom. cons.
L. Beauvisage exNakai, 1948 129a. HeliamphoraceaeChrtak,
120. ClusiaceaeLindley, 1836, Slavikova, & Studicka, 1992
nom,. cons. L. Ericanae Takhtajan, 1967
Guttiferae A.L. Jussieu, 1789, 49. Ericales Dumortier, 1829
nom. cons.; nom. alt. Vacciniales Dumortier, 1829
Garciniaceae Bartling, 1830 130. Pentaphylacaceae En-
CambogiaceaeHoraninow, 1834 gler, 1897, nom. cons.
CalophyllaceaeJ. Agardh, 1858 131. ClethraceaeKlotzsch, 1851,
121. HypericaceaeA.L.Jussieu, nom. cons.
1789, nom. cons. 132. Cyrillaceae Endlicher,
Ascyraceae Plenck, 1796 1841, nom. cons.
45. Elatinales Nakai, 1949 133. Ericaceae A.L. Jussieu,
122. Elatinaceae Dumortier, 1789, nom^. cons.
1829, nom.. cons. RhododendraceaeA.L.Jussieu,
Cryptaceae Rafinesque, 1820 1789
AlsinastraceaeRuprecht exBu- Rhodoraceae Ventenat, 1799
bani, 1901 Azaleaceae Vest, 1818
I. Nepenthanae Takhtajan exRe- Ledaceae Link, 1821
veal, 1992 Menziesiaceae Klotzsch, 1851
46. NepenthalesDumortier, 1829 Salaxidaceae J. Agardh, 1858
123. NepenthaceaeDumortier, Diplarchaceae Klotzsch, 1860
1829, nom. cons. 134. VacciniaceaeCandoUe ex
J. LecythidanaeTakhtajan exRe- Gray, 1821, nom,. cons.
veal, 1992 Androm,edaceae(Endlicher)Schni
47. LecythidalesCronquist, 1957 zlein, 1843-1870
124. BarringtoniaceaeF. Rudol- SiphonandraceaeKlotzsch, 1851,
phi, 1830, nom,. cons. nom,. illeg.
125. Foetidaceae (Nidenzu) Arbutaceae J. Agardh, 1858
Reveal: Guide to higher taxa ofMagnoliophyta 211
Arctostaphylaceae J. Agardh, 147. Theophrastaceae Link,
1858 1829, nom.. cons.
135. Pyrolaceae Dumortier, 148. Myrsinaceae R. Brown,
1829, nom. cons. 1810, nom. cons.
136. Monotropaceae Nuttall, ArdisiaceaeA.L.Jussieu, 1810
1818, nom. cons. Embeliaceae J. Agardh, 1858
Hypopityaceae Link, 1829 149. Aegicerataceae Blume,
137. Epacridaceae R. Brown, 1833
1810, nom. cons. 56. Primulales Dumortier, 1829
StypheliaceaeHoraninow, 1834 Samolales Dumortier, 1829
138. Prionotaceae Hutchin- 150. Primulaceae Ventenat,
son, 1969 1799, nom.. cons.
50. Empetrales Nakai, 1930 Lysimachiaceae A.L. Jussieu,
139. EmpetraceaeGray, 1821, 1789
nom. cons. AnagallidaceaeBatsch eiBorck-
51. Fouquieriales Takhtajan ex hausen, 1797
Reveal, 1992 Samolaceae Rafinesque, 1820
140. FouquieriaceaeCandolle, 151. Coridaceae J. Agardh,
1828, nom. cons. 1858
52. Ebenales Engler, 1892 N. Plumbaginanae Takhtajan ex
Diospyraks Prantl, 1874 Reveal, 1992
141. EbenaceaeGiircke, 1891, 57. PlumbaginalesLindley, 1833
nom. cons. 152. PlumbaginaceaeA.L.Jus-
GuaiacanaceaeA.L.Jussieu, 1789 sieu, 1789, nom. cons.
Diospyraceae Vest, 1818 153. AegialitidaceaeLinczevski,
142. Lissocarpaceae Gilg, 1924, 1968
nom. cons. 154. LimoniaceaeSeringe, 1851,
53. Styracales Burnett, 1835 nom. cons. prop.
143. Styracaceae Dumortier, Staticaceae Cassel, 1817
1829, nom. cons. ArmeriaceaeHoraninow, 1834
Halesiaceae D. Don, 1828 0. Polygonanae Takhtajan exRe-
54. Sapotales J.D. Hooker, 1868 veal, 1992
144. SapotaceaeA.L.Jussieu, 58. Polygonales Dumortier, 1829
1789, nom,. cons. 155. Polygonaceae A.L.Jussieu,
Achradaceae Vest, 1818 1789, nom. cons.
Bumeliaceae Barnhart, 1895 Rumicaceae Martinov, 1820
145. BoerlagellaceaeH.J. Lam, Eriogonaceae(Dumortier) Meis-
1925 ner, 1841
146. Sarcospermataceae H.J. PersicariaceaeAdanson exPost
Lam, 1925, nom.. cons. & Kuntze, 1903
M. Primulanae R. Dahlgren ex 156. CalligonaceaeChaJkuziev,
Reveal, 1992 1985
55. Myrsinales Bromhead, 1838 P. Celastranae Takhtajan, 1967
HYTO OGIA
212 P L volume 74(3):203-263 March 1993
59. Celastrales Baskerville, 1839 Kuntze, 1903
157. Celastraceae R. Brown, 167. Huaceae A. Chevalier,
1814, nom. cons. 1947
Euonymaceae A.L. Jussieu ex 168. ElaeocarpaceaeA.L.Jussieu
Berchtold & J. Presl, 1820 exCandolle, 1824, nom. cons.
ChingithamnaceaeHandel-Maz- AnstoteliaceaeDumortier, 1829
zetti, 1932 169. PlagiopteraceaeAiryShaw,
158. Canotiaceae Airy Shaw, 1965
1965 170. Tiliaceae A.L. Jussieu,
159. HippocrateaceaeA.L.Jus- 1789, nom. cons.
sieu, 1811, nom. cons. SparmanniaceaeJ. Agardh, 1858
Salaciaceae Rafinesque, 1838 171. Monotaceae(Gilg) Maury
160. Siphonodontaceae(Croizat) ex Takhtajan, 1987.
Gagnepadn& Tardieu exTar- 172. DipterocarpaceaeBlume,
dieu, 1951, nom. cons. 1825, nom. cons.
161. Pottingeriaceae (Engler) 173. Sarcolaenaceae Caruei,
Takhtajan, 1987 1881, nom. cons.
162. Goupiaceae Miers, 1862 Schizolaenaceae Barnhart, 1895
163. Lophopyxidaceae (En- RhodolaenaceaeBullock, 1958
gler) H. Pfeiffer, 1951 174. DiegodendraceaeCapuron,
164. StackhousiaceaeR.Brown, 1964
1814, nom. cons. 175. SphaerosepalaceaeTieghem
Q. Malvanae Takhtajan, 1967 ex Bullock, 1959
60. Malvales Dumortier, 1829 Rhopalocarpaceae Hemsley ex
Tiliales Hutchinson, 1924 Takhtajan, 1987
165. Sterculiaceae (CandoUe) 176. BombacaceaeKunth, 1822,
Bart-ling, 1830, nom. cons. nom. cons.
TriplobaceaeRafinesque, 1838 177. Malvaceae A.L. Jussieu,
166. ByttneriaceaeR. Brown, 1789, nom. cons.
1814, nom. cons. PhilippodendraceaeEndlicher,
HermanniaceaeBerchtold&J. 1841
Presl, 1820 PugosiaceaeMartinov, 1820, nor
Lasiopetalaceae Reichenbach, illeg.
1823 Hibiscaceae J. Agardh, 1858
Dombeyaceae(Candolle)Bart- PlagianthaceaeJ. Agardh, 1858
ling, 1830 R. Urticanae Takhtajan ex Re-
PremontiaceaeJ. Agardh, 1858 veal, 1992
Helicteraceae J. Agardh, 1858 61. Ulmales Lindley, 1833
Melochiaceae J. Agardh, 1858 Ficales Dumortier, 1829
Theobromataceae3 Agardh, 1858 178. Ulmaceae Mirbel, 1815,
.
ChiranthodendraceaeA. Gray, nom. cons.
1887 179. Celtidaceae Link, 1831
Cacaoaceae Augier ex Post & 180. Moraceae Link, 1831,