A Revision of the Genus Boesenbergia Kuntze (Zingiberaceae) in Thailand

24 Pages·1992·1.1 MB·English

by  Sirirugsa, Puangpen

NAT. HIST. BULL. SIAM SOC. 40 :67 ・90，1992
A REVISION OFT HE GENUS BOESENBERGIA KUNTZE
(ZINGIBERACEAE) IN THAILAND
Puangpen Sirirugsa*
ABSTRACT
Revision of血e.genus Boesenbergia (Zingiberaceae) in百lailandresulted in血e
問cognitionof 13 species. Ak ey to出巴百laispecies with descriptions and illus回.tions，釦d
information on their dis.tribution，ec ology and vemacular names are presented.
INTRODUCTION
Boesenbergia is ag enus of small rhizomatous herbs，mo stly growing on the floor
of出eforest. About 40 species of the genus are distributed in tropical Asia. Theg enus has
been treated by: BAKER (1890) for India，in cluding two new species former1yd escribed by
WA LLICH (1830); SCHUMANN (1904) for Bomeo and Sumatra，a nd much later SMITH
(1982) added eight new species; GAG阻 PA別(1908)for Indochina; VA LETON (1918) for
Java and Malaysia; RIDLEY (1924) dHOLTTUM (1950) for the Malay Peninsu1a.
佃
Thet ype species of Boesenbergia is Gastrochilus pulcherrima Wall. According
to the Intemational Botanical Nomenclature Gastrochilus is invalid，be ing al ater homo-
nym of組 orchidaceousgenus (HOL'甘 UM，1950).
In Thai1and 13 species of Boesenbergia are now recorgnized. A key to the Thai
species，w ith descriptions，il lustrations，a nd notes on their dis凶bution，ecology and ver-
nacular names is presented.
Boesenbergia O. Kuntze
Rhizome usually short，fl eshy; leafy stem 20--60c m tall，ra rely up to 90 cm，b eing1t o
釘
8l eaves，u sually with af ew sheaths outside the leaf-sheaths. Leaves elliptic，o blong or
elliptic-oblong，ra rely ovate or lanceolate. Petiole 2-15 cml ong，ra rely sessi1e; ligule bi1obed.
I析Orescenceterminal，b etween出etwo innermost leaf-sheaths or directly from the rhi-
zome sep釘atelyfrom the leaf-sheaths，a xis of inflorescence be釘ing2-r創立edaltemate
bracts. Bracts oblong，el liptic or lanceolate，(2 -5)x(1-2) cm，en closing as ingle bracteole
and as ingle flower; bracteoles shorter，ei ther nToweror wider than the bract，bo th sides
釘
inflexed enfolding the flower. Calyx tubular，sh ort，3・ lobed;lobes佐uncateor acute，de eply
divided on one side. Corolla-tube slender，l onger than bract; lobes usually subequal，th e
dorsal one白e1釘gest.Labellum usually longer than corolla-lobes and staminodes，sa ccate，
白.eapex often irregularly crenate and crinkled，ra rely bi1obed. Staminodes obovate，el liptic
or oblong，ra rely ovate or oblanceolate，ab out the same length or shorter and al ittle wider
than the corolla-Iobes. Stamen: filament about as long as or either shorter or longer出an
*Department of Biology，Fac ulty of Science，Pri nce of Songkla University，Ha t Yai，百lailand.
67
68
PUANGPEN SIRIRUGSA
the anther; anther opening by slits or in B. curtisii by apical pores; connective either
produced beyond the thecae or no .tOvary glabrous or hairy，子locularor incompletely 3-
locular with axile placenta，1 -10cular with free-central placenta or in B. parvula with
parietal placenta; stylodes slender 3-6 mmo r up to 1c m long. Fruit ellipsoid or cylin-
drical-ellipsoid. Seeds arrillate，re latively large; ilwhite.
紅
Key to the Species in Thailand
1. Inflorescence basal directly from rhizome 2
1. Inflorescence terminal 3
2. Spike conicaI or oblong-conical，se parate from leaf-sheaths. Staminodes longer than
half the length of labellum 1. B. basispicata
2. Spike linear，wi thin leaf-sheaths. Staminodes shorter than half the length of labellum
2. B. prainiana
3. Anther crest not produced beyond thecae 4
3. Anther crest produced beyond thecae 8
4. Pollen sacs poricidal dehiscing. Inflorescence not exerted from leaf-sheaths. Label
晴
lum not saccate 3. B. curtisii
4. Pollen sacs longitudinal dehiscing. Inflorescence elongate. Labellum saccate 5
5. Leaf base cordate; petiole relatively longer，u p to 8c m 4. B. petiolata
5. Leaf base cuneate，pe tiole relatively shorter，se ssile or up to 5c m 6
6. Ligule and ovary hairy 7
6. Ligule and ovary glabrous 5. B. xiphostachya
7. Leaves sessile，n arrow，a bout 5t imes longer than wide. Labellum ovate
6. B. thorelii
7. Leaves petiolate，b road，a bout 2t imes longer than wide. Labellum obovate
7. B. long伊es
8. Leaves absent at anthesis 8. B. siamensis
8. Leaves present at anthesis 9
9. Inflorescence compact，in cIuded，us ually on1yf lowers exerted beyond leaf-sheaths
9. B. rotunda
9. Inflorescence elongate，ex erted from the leaf-sheaths 10
10. PeduncIe2 -4 cm long. Inflorescence 10・20cm long; two rows of bracts facing one
way 10. B. plicata
10. PeduncIer arely longer than 2c m. Inflorescence 3-12 cm long 11
11. Leaves lanceolate，ap ex long acuminate，ba se mostly truncate. Petioles usuaIly longer
than 15 mma nd up to 35 mm. Corolla-lobes elliptic-oblong or oblong，a pex acute
or mucronate. Connective glabrous 11. B. acuminata
11. Leaves otherwise. Petioles rarely up to 12 mm. Corolla-lobes oblong，a pex rounded.
Connective hairy 12
12. Plant with leafy stem rarely more than 16 cm tall. Leaves ovate to elliptic，2- 5 cm
long. Staminodes oblong，g labrous，a bout 10x4 mm. Labellum ovate-oblong，g la-
brous，e xcept the brownish-hairy base，2 0-24 x 15-20 mm. Ovary 1・loculate
12. B. parvula
A REVISION OFT HEG ENUS BOESENBERGIA 69
12. Plant with leafy stem 2050cm tall. Leaves elliptic or oblong，1 040cm long.
・ ・
Staminodes obovate，glandular"hairy，ciliate，about 20x13mm.Labellum ovate or
suborbicular，gl andular-hairy，c iIiate，a bout 3033x2 5-32 mm. Ovary 3loculate
・ ・
13. B. pulcherrima
1. Boesenbergia basispicata K. Larsen ex P. Sirirugsa，No rd. J. Bot. 7( 4) :1 987. Fig.l
(L-R) PlateI: 1
Leafy stem 40--60ー( 75)cmt aIl; sheaths up to 10 cml ong，gl abrous or glabrescent.
Leaves 3-6; blades oblong，of ten unequaI-sided，1 5-35 x4 ー-9.5cm，ap ex acuminate，ba se
cuneate or obtuse，de current，up per surface glabrous，lo wer surface glabrescent. Petiole (5)
7・15(18) cm long，g labrous; ligule-Iobes broadly仕iangularor oblong，3 -6 mml ong，
glabrous; leaf-sheaths 8-14 cm long，g labrous or glabrescent. Inflorescence bas ，Ia as-
cending from rhizome，no t enclosed by leaf-sheaths，co nical or oblong-conic ，Ia8-12 (-17)
cm long; flowers about 12; peduncle ca. 5m ml ong. Bracts dark reddish，m ottled with
pink，of ten incurved，ca . 3x 1. 9c m，gl abrous; bracteoles folded，ca . 1.4x 0 .8 cm，gl abrous.
CaIyx pinkish，ca . 4m ml ong，lo bes short，tr uncate，ca . 1m ml ong. Corolla: tube ca. 2.5
cm long; lobes elliptic，ap ex rounded，in curved，ca . 1.2x 0 .6 cm，gl abrous. Labellum ovate
or suborbicular，sa ccate，a pex irregularly crenate and crinkled，ca . 2.0 x1 .7c m，ha iry at
the base，fi nely cilia匂.Staminodes whitish，el liptic or ovate，ap ex rounded or biIobed，ca .
1.1 x0 .7 cm，l onger than half the leng出 oflabeIlum，fi nely hairy and ciliate. Stamen:
filament ca. 4m m，sp arsely brownish-hairy; therca. 5m m; connective almost glabrous;
叩
crest not prouduced beyond the thecae. Ovary ca. 4m ml ong，g labrous，i ncompletely 3
・
locular; stylodes linear，ac ute，ca . 5.5 mml ong. Fruit eIIipsoid，ta pering towards apex，ca .
1.7x 0 .8 cm. Seeds ca. 7，ar illate，7 -10 x2 -3 mm; aril whitish-hairy.
T ha i 1 a n d .ー-PENNINSULAR
Nakhon Si Thammarat (Khao Lu g，type) ，T rang，N arathiwat.
佃
D is t r i b u t i 0 n. -Endemic to Peninsular ThaiIand.
E c0 10 gy . -Common on damp slopes in evergreen forest，3 00・600m ms .l
V er n a c u 1 a r. -Krachai khao luang (問白川"II'1'1i1fl叩)
2. Boesenbergia prainiana (King ex Bak.) Schltr. in Fedd. Rep. 12:316.1913; Loes. in
Pflanzenf:n.2. 15 a: 570.1930; Holttum，G ard. Bull. Sing.13: 108. 1950. -Kaempferia
釘
prainiana King ex Bak. f. in F.lBr. Ind. 6: 220. 1890; Schum. in Pflanze .4.46:84.
町
1904.-Gastrochilus prainiana (King ex Bak.) Rid ，.lJ. Str. Br. As. Soc. 32:115.1899; FI.
Ma .lMat. Pen 2: 18.1907; F .lMa .lPen. 4: 248.1924. -Gastrochilus albosanguinea Rid ，.l
J. S江 Br.As. Soc. 32: 111.1899; Schum. in Pflanzenr. 4. 46:93. 1904; Rid ，.lMat.FI. Ma .l
Pen. 2:15.1907; F .lMa .lPen.4:247.1924. --Boesenbergia albo-sanguinea (Ridl.) Loes. in
Pflanzenfam.2. 15 a: 568.1930. Fig.2，P lateI: 2
Leafy stem 25-40c m taIl; sheaths pu中le，the longest up to 12 cm long，ha iry.
Leaves 1-4， ususIly 1o r 2; blades eIIiptic or eIIiptic-oblong，r arely oblanceolate，o ften
unequal sided，1 2-23 x5 -10 cm，ap ex short acuminate，ba se cuneate，ra rely rounded，a nd
decurrent，up per surface glabrous，lo wer surface sparsely hairy. Petiole 2.5-6.5(-9) cml ong;
Iigule-lobes broadly triangul訂， 2-4 mml ong，h airy; leaf-sheaths 4-10 cm long，h airy.
Inflorescence basal，as cending from rhizome within the innermost leaf-sheath and clasped
by sheaths，Ii near，1 1-25 cm long;peduncle 1-4 cm long. Bracts incurved，ca . 1.8-2.2 x
70
PUANGPEN SIRJRUGSA
I
I~ lmm
2mmOR
I em
~H
I
I em
0.(.. )
~·
G
A I em ...
0.
ffi ·
~
· .·,.
.
'
D I E
Figure I. A-K : Boesenbergia acuminara. A : Habit. B : Bract. C : Bracteole. D : Dorsal corolla-lobe. E : One
of lateral lobes. F: Labellum. G: Staminodes. H :Calyx and ovary. I: Anther from behind. J: Stigma.
K :Fruit (A, K from shimizu et al. T : 7917; B-J from Kerr 19738). L-U : B. basispicara. L : Habit.
M : Bract. N : Bracteole. 0 : Flower. P : Labellum. Q : Staminodes and stamen from the front. R :
Calyx and ovary. S: ovary in X.S. T: Ovary in L.S .. U: Seed. (after Sirirugsa, 1987).
A REVISION OF THE GENUS BOESENBERGIA 71
1mm
[
2mm
H
Figure 2. Boesenhergia prainiana. A : Habit B : Flower. C : Bract. D : Bracteole. E : Calyx and ovary. F :
Stamen from behind. G : Stamen from the front. H : Stylodes and ovary. l : Ovary in L.S. showing
seeds. (All from Charoenphol et al. 3987)
72 PuANGPEN SIRIRUGSA
1.5c m，s pselyhairy; bracteoles folded，1 .2-1.6 x0 .8 cm，s pselyhairy. Calyx tinted
釘 紅
with red spots，4-5mml ong;lobes tmncate，ca.1mm long.Corolla--tube ca.35cm long;
。
lobes lancelate-oblong，apexrounded，35-5x1.3mm.Labellum whitish with red blotch
at apex，ov ate，s位 onglysaccate，ap ex undulate and crinkled，ca . 2x1.7c m，br ownish-hairy.
Staminodes whitish，el liptic，ap ex rounded，ob tuse or mucronate，ca . 6.5 x3 .5 mm，sh orter
than half the length of labellum，gl abrous，ci liate near the base. Stamen: fi1ament ca. 6m m
long，fi nely hairy; anther ca. 5m ml ong; connective hairy; crest not beyond thecae. Ovary
ca. 3m ml ong，gl abrous，in completely 3locular;stigma trilobed; stylodes linear，ac ute，ca .
・
7m ml ong.
T ha i 1 a n d . -SOUTHWESTERN:Prachuap Khiri Khan;
田
PENINSULAR: Narathiwat，Pa ttani.
D is t r i b u t i 0 n. -Malay Peninsular，( Perak，K ing 726，ty pe)，In dia.
Ec 0 10 gy .-Common in evergreen fo!est，a long stream，5 0-400 m msl.
V er n a c u 1 a r. -Kathue daeng (n~'Vì叫附)
3. Boesenbergia curtisii (Bak.) Schltr. in Fedd. Rep. 12:316. 1913; Loes. in Pflanzenfam.
2. 15a: 571. 1930; Holttum，Ga rd. Bull. Sing. 13: 112. 1950; Baker & B北h.f.Fl.Java 3:
69. 1968.ー-Gastrochiluscurtisii Bak. in Hook.f. Bot.Mag.: t.7363. 1894; Rid ，.lJ. Str. Br.
As. Soc. 32: 113. 1899; Mat. Mal. Pen. 2: 18.1907; F .lMal. Pen. 4: 249.1924.--GJavanus
Schum. in Pflanzenr. 4. 46: 95. 1904，Va let.; Bull. Jard. Bot. Buitenzorg，se r. 2. 27: 92.
1918.--G.acuta Ridl. J. S包 Br.As. Soc. 59:202. 1911;日.Mal.Pen. 4: 249. 1924.-
Boesenbergia javanum (Schum.) Schltr. in Fedd. Rep. 12 :3 16. 1913; Loes. in Pflanzen-
fam. 2. 15a: 571. 1930. Fig.3，P lateI: 3
←
Leafy 批 m40-5090) cm tall; sheaths 7-13 (-18) cm long，ha iry. Leaves 4-5;
blades elliptic，el liptic-oblong to oblong，1 2-40( -48) x3 -16 (-18) cm，a pex acuminate，
base rounded or cuneate，m ore or less decurrent，so metimes subcordate. Petiole ι10( -16)
cm long，sp arsely hairy; ligule-lobes lanceolate，ac ute，5 -12 mml ong，ha iry; leaf-sheaths
7-13 cml ong，sp arsely hairy.lnflorescence terminal，en closed by the upper two leaf-sheaths，
5-9 cm long; rachis short; peduncle 0.5-1.5 cm long. Bracts ca. 4.5 x0 .1 cm，ha iry with
red streaks; bracteoles ca. 4.2 x1 .2c m，ha riy，wi th red streaks. Calyx ca. 1.8-2.5 cml ong，
hairy; lobes acute，ca . 3c m long，ci liate. Corolla: tube ca. 7c m long，br oadened upwards;
lobes whitish，o blong，a cute，1. 5-1.7c m long. Labellum yellowish with pink streaks on
either sides at the base，el liptic or obovate，ap ex lobed and irregularly crenate，re flexed，ca .
1.6xO.9 cm，ha iry along at the median below the middle downwards.
Staminodes whitish，o blong，a pex rounded or slightly lobed，ca . 1.5 x0 .5 cm，g labrous，
with red blotches at the base. Stamen:・filamentca. 4m ml ong，ha iry at the back; anther
ca. 3m ml ong; connective pubescent; pollen sacs poricidal opening，p ore ci1iate; crest
bilobed，no t produced beyond白ecae.Ovary ca. 5m ml ong，gl abrous，3 locular;ovules 4-
・
5i n each locule; stigma lobed，ci liate; stylodes linear，8 -10 mml ong.
Th a i 1 a n d . -SOUTH-EASTERN :C hon Buri; SOU1百-WESTERN:Kanchanaburi，
Ratchaburi，Pr achuap Khiri Khan; PEN別SULAR:Chumphon，Su rat Thani，Ph angnga，Na khon
Si Thammarat，T rang，P hattalung，S ongkla，Sa tun.
D is t r i b u t i 0 n. -Malay Peninsula (Lankawi，Cu rtis 2896，type)，Ja va，In dia.
。
E c0 10 gy . -Onl imestone hi1l in evergreen or mixed deciduous forests，
-300m msl.
A REVISION OF THE GENUS BOESEN BERG/A 73
',,
I '
'' ,·
,.
·'
'
D
I
2mm
Figure 3. Boesenbergia curtisii. A : Habit. 8 : Flower. C : Bract. D : Bracteole. E : Calyx and ovary. F: Stylocles.
G: Ovary in X.S. (A, from Charoenphol et al. 3643; 8, C, D, from Kerr I 0588: E, F, G, from Larsen
10458).
74
PUANGPEN SiRIRUGSA
,,,,r
•' '' '
I I ~
'···
' '' I em
I em
'; p
lmm
1 J
em
Figure 4. A-1 : Boesenbergia pe1iola1a. A : Habit. B : Bract. C : Bracteole. D : Flower. E : Staminocles and
stamen from the front. F : Labellum. G : Stamen from behind. H : Ovules. I : Seed. (All from
Maxwell 74-631). J-Q :B. siamensis. J: Habit. K: Dorsal corolla-lobe. L: Staminodes. M: Labellum.
N : Stamen from the front. 0 : Stamen from behind. P : Stigma. Q : Ovary in L.S. (All from G. den
Hoed 238). (after Sirirugsa, 1987)
A REVISION OFT HEG ENUS BOESENBERGIA 75
4. Boesennbergia petiolata P. Sirirugsa，N ord. J. Bot. 7( 4): 423. 1987. Fig.4 (A-I)
Leafy stem ca. 50 cm taIl; sheaths ca.ι12 cm long，gl abrous. Leaves 4-5; blades
eIliptic or oblong，r arely ovate，7 -19 x2 .5-6 cm，a pex acuminate，b ase cordate，r arely
truncate，b oth surfaces glabrous. Petiole 3( 4)-6 (-8) cm; ligule-lobes broadly triangul
釘
or oblong，a pex obtuse，7 -15 mml ong，m embranous，g labrous; leaf-sheaths up to 20 cm
long，gl abrous，s ometimes with gland-dots. lnflorescence terminal，b etween the upper two
lea凶.fsheaths，later clasped by the uppermost leaf-sheath，ca . 8c m long. Bracts ca. 3x 0 .8
cm，ha iry; bracteoles ca. 2.3 x0 .3 cm，ha iry. Calyx ca. 4m ml ong; lobes broadly trian-
gular，I-1.5m ml ong; lower pa口ofthe tube sparsely reddish-brown-hairy. Corolla: tube
ca. 1.6 cm long，b roadened upwards; lobes elliptic，a pex rounded，in curved，ca . lxO.5 cm，
with gland-dots. Labellum obovate or suborbicular，s trongly saccate，a pex irregularly cre-
nate dcrinkled，ca . 1.5x 1 .5c m，br ownish-hairy，w ith gland-dots. Staminodes obovate，
加
apex rounded，s ometimes iegularlylobed，ca . 8x 5 m m，gl andularhairy，with gland-dots.
汀
帽
Stamen:・filamentca. 4m m，gI andular-hairy; anther ca. 5m m; connective glandular-hairy;
crest bilobed，no t produced beyond thecae. Ovary ca. 3-4 mml ong，r eddish-brown-hairy;
stigma lobed; stylodes linear，a cute，ca . 2.5 mm; placenta free-centraI; ovules ca.6. Fruit
ellipsoid，ca . 10x3 mm，s parsely reddish-hairy near the top and base. Seeds ca. 4，cy lin-
drical-Ianceolate，ca . 6.5xl mm，fi nely hairy，w ith gelatinous cap，ar illate; aril whitish.
Th a i 1 a n d .一一NORTHERN:Chiang Mai，p hitsanulok，E ASTERN: Chaiyaphum，
Nakhon Ratchasima; CENTRAL: Saraburi.
D is t r i b u t i 0 n.一一Endemicto Thailand.
Ec 0 10 gy . -In mixed deciduous and evergreen forests.
5. Boesenbergia xiphostachya (Gagnep.) Loes. in Pflanzenfam. 2. 15a: 568.1930.
-Gastrochilus 卯 hostachyumGagnep.，B uIl. Soc. Bot. Fr. 53: 147. 1906.
Leafy stem up to 70 cm tall，be aring 4-8l eaves; sheaths about 3，th e longest up
to 18 cm long. Leaves lanceolate，1 3-20 (-30) x3 -5 (8.5) cm，a pex acuminate，b ase
cuneate，b oth surfaces glabrous. Petiole 2-5 cm long，gl abrous; ligule-lobes oblong，a pex
rounded，0. 1ー1.5x0 .3-0.5 cm，g labrous; leaf-sheaths up to 27 cm long，g labrous. lnflo-
rescence terminal，u suaIlyp eduncle enclosed by the uppermost Ieaf-sheaths，e xtended to
25 cm long; flowers up to 20 or more; peduncle 2-6 cm Iong. Bracts ca. 5x 1.4 cm，
glabrous not concave; bracteoles folded，ca . 3x 0 .9 cm，ha iry. Calyx ca. 5m ml ong，lo bes
truncate. Corolla: tube extended，di lated to the top; Iobes oblong，a pex rounded，gl abrous，
ca 1.2 x0 .5 cm. Labellum eIliptic or ovate，sa ccate，a pex iηegularly crenate and crinkled，
ca 1.3 x0 .8 cm，g labrous with conspicuously gland-dots. Staminodes oblanceolate，g la-
brous with gland-dots. Stamen: filament 2-3 mml ong，h airy; anther ca. 7.5 mml ong，
connective glandular-hairy，cr est bilobed，n ot beyond thecae. Ovary ca. 4m ml ong，gl a-
brous，l -Iocular; stylodes linear，o btuse，ca . 3m ml ong; placenta free-central.
T ha i 1 a n d .一一NORTH-EASTERN:S akon Nakhon.
D is t r i b u t i 0 n. -Indo-china (type).
Ec 0 10 gy . -In evergr，田nforest.
6. Boesenbergia thorelii (Gagnep.) Loes. in Pflanzenfam. 2. 15a: 568.1930. -Gastrochilus
thorelii Gagnep.，B ull. Soc. Bot. France 14: 163. 1907.
76 PUANGP印 SIRIRUGSA
Leafy stem up to 60 cm tall; sheaths ca. 3，u p to 17 cm long，sp arsely hairy or
glabrous. Leaves 5-8， sessile; blades elliptic-oblong，1 ι18 x2 -3.5 cm，s pex acuminate，
base cuneate，bo th surfaces glabrous or almost glabrous; ligule-lobes oblong，ap ex rounded，
ca.6θmm long，hairy，ci liate. lnflorescence terminal，e nclosed by血euppermost leaf-
sheaths，5 -8 cml ong; peduncle ca. 1-1.2c m long. Bracts ca. 3.4-4 x0 .6 cm，ha iry，ap ex
ciliate; bracteoles ca. 2.8 x0 .7 cm，ha iry，a pex ciliate. Caか'xca. 6m ml ong，g labrous;
lobes truncate，u nequal，t he longest one ca. 1.5 mm. Corolla: tube ca. 2.4 cm long，
broadened towards the top，gl abrous;lobes elliptic，a pex rounded or short mucronate，1 0
-16 x5 -8 mm，gl abrous，w ith gland-dots. Labellum ovate，st rongly saccate，a pex irregu-
larly crenate，ca . 2.5 x2 .2 cm. Staminodes glabrous，w ith densely gland-dots. Stamen:
filament ca. 6.5 mm，sh ort brownish hairy; anther ca. 6m ml ong; connective glandular-
hairy; cr，回t2・lobed，not beyond血ecae.Ovaηca.2-6 mml ong，gl abrous; style upper p紅t
hairy; stigma ciliate; stylodes linear，ac ute，ca . 2-4 mml ong; placenta合'ee-central，ovules
ca 15.
Thailand.一ーCENTRAL:Saraburi; EASTERN; Nakhon Rachasima
D is t r i b u t i 0 n. -Laos (Bassac，百lOrels.n. type，P)
E c0 10 gy .ー Inbamboo-hardwood evergreen forest，ca . 200 m msl.
7. Boesenbe棺ialongipes (King & Prain) Schltr. in Fedd.Rep. 12: 316.1913; Loes. in
Pflanzenfam. 2. 15a: 571.1930; Holttum，G ard. Bull. Sing. 13: 114.1950. -Gastrochilus
longipes King & Prain ex Ridley，J. Str. Br. As. Soc. 32:113.1899; Mat.日.Mal. Pen. 2:
一
17.1907. Kaempferialongipes (King & Prain) Schum. in Pflanze町 .4.46:79.1904. Fig.5
Platel:4
Leafy stem ca. 50 cm tall; sheaths ι10 cm long，pu rple，sp arsely hairy. Leaves
2-5，ra rely 1o r 6; blades elliptic or elliptic-oblong to oblong，m ostly unequal sided，1 0
・
12 x4 ー7cm，a pex acuminate，ba se cuneate，up per surface glabrous，lo wer surface hairy.
Petiole 3-4 cm long，s parsely hariy; ligule-lobes凶angularor lanc泡olate，1-2 cm long，
hairy; leaf-sheaths 11-17 cm long，ha iry. lnflorescence terminal，be tween the uppermost
leaf-sheaths，ca . 10 cm long; peduncle 2-3 mml ong. Bracts ca. 3 x 0.7 cm，h airy;
bracteoles folded，ca . 2x 0 .8 cm，ha iry. Caか'X5-7 mml ong; lobes short加 dbroaded，ap ex
truncate，ca . 2m ml ong，ha iry. Corolla: tube ca. 2.4 cml ong; lobes elliptic，ap ex rounded，
incurved，ca . 1x 0 .7 cm，gl abrous. Label/um whitish with red blotches at the middle，ob ovate，
saccate，a pex ref1exed and crinkled，ca . 2-3 xO .ι1 cm，o utside short brownish hariy.
Staminodes whitish，ob ovate，ap ex 2・lobed，ca. 9x 7 m m，ou tside short brownish hairy at
base. Stamen:・filamentca. 3m ml ong，sh ort brownish hairy; therca. 6m ml ong; con-
佃
nective glandul hairy;crest bilobed，no t produced beyond thecae. Ovary ca. 4m ml ong，
釘
sparsely hairy，tr ilocular; stigma inconspicuously lobed，no t ciliate; stylodes cylindrical，ca .
4m ml ong; placenta axile.
Thailand.ーP聞 INSULAR:Nakhon Si Thammarat，T rang.
D is t r i b u t i 0 n. -Malay Peninsula (Perak，W ray 4220，ty pe).
E c0 10 gy .ー Inshaded evergreen forest，ca . 150 m msl.
V er n a c u 1 a r. -Nila phusi (UfUJ世)(Southern)
8. Boesenbergia siamensis (Gagnep.) P. Sirirugsa白，Nord.J. Bot. 7(4): 425， 1987.
-Gastrochilus siamense Gagnep.，N otul. Syst. 14. 1: 36.1950. Fig. 4( J・Q)