A revision of the genus Acanthotyla stål with the description of five new species and synonymical note (Heteroptera: Coreidae: Colpurini)

2005·11.8 MB·

by  Harry Brailovsky

PROC. ENTOMOL. SOC. WASH.
107(2), 2005, pp. 279-302
A REVISION OF THE GENUS ACANTHOTYLA STAL WITH THE
DESCRIPTION OF FIVE NEW SPECIES AND SYNONYMICAL NOTE
(HETEROPTERA: COREIDAE: COLPURINI)
Harry Brailovsky
Departamento de Zoologia, Instituto de Biologia, U. N. A. M., Apdo Postal No. 70153,
Mexico, 04510 D.F. Mexico, (e-mail: [email protected])
—
Abstract. The genus Acanthotyla Stal is revised. The following new species are de-
scribed: A. kaloboana, A. kiungala, A. nabirenia, and A. protenta all from New Guinea
and A. borneana from Borneo. Acanthotyla aruana Horvath is synonymized with A.
fasciata (Walker) (n. syn.); new records for the previously known species are added.
Habitus illustrations and drawings of the male and female genitalia, as well as head and
pronotum, are provided, and a key to the known species is presented.
Key Words: Hemiptera, Heteroptera, Coreidae, Colpurini, revision Acanthotyla, new
species, Borneo, New Guinea, synonymical note
The genus Acanthotyla was proposed by genus is characterized by having the tylus
Stal (1873) to include only the species Cle- projecting in front ofjuga, upturned to form
tusfasciatus Walker (1871) from Mysol Is- a horn at the apex, the mandibular plates
land. Cletus crassus Walker (1871), de- with prominent tubercle, the antennal seg-
scribed in the same paper, was transferred ment III longer than IV, the antenniferous
to Acanthotyla by Blote (1936), but later tubercles at outer corner projected forward,
Brailovsky (1995) moved it to his new ge- calli conspicuously convex, and posterior
nus Brachylybastella as the type species. lobe of pronotal disk with a transverse
Breddin (1900) included Acanthotyla in his wrinkle. The specific differences are chiefly
key to the genera of Colpurini. The second in the development of the humeral angles
known species, A. aruana Horvath (1919), and the intercallar space of the pronotum;
is synonymyzed in this paper under A. fas- the shape of the posteroventral edge of the
ciata. The last group of species was de- male genital capsule, and shape of the fe-
scribed by Blote (1936) from New Guinea male genital plates.
under the binomial names Acanthotyla dis- Members of this genus are distributed
tinguenda and Brachylybasflexiiosus. Brai- from Borneo, Aru Island, Mysol Island. Iri-
lovsky (1993) revised the Colpurini ofAus- an Jaya. Papua New Guinea, and Australia
tralia and recorded A. fasciata for the first (Brailovsky 1993).
time from that region, and later Brailovsky Previously, only three species of Acan-
and Martinez (1994) transferred B. flexu- thotyla, A. distinguenda, A.fasciata, and A.
osus to Acanthotyla. flexuosa, were known. In this contribution
Acanthotyla belongs to the group of Col- the genus is redescribcd. new records forA.
purini with abdominal sternite VII of the distiiii^ncnda, A. fasciata, and A. flexuosa
female without plica or fissura, and has not are given, and five new species, from Bor-
been previously reviewed or revised. The neo. Irian .lava, and Papua New Guinea are
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280 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON
described. Acanthotyla aruana Horvath is transverse wrinkle, calli conspicuously con-
synonymyzed under A. fasciata. vex, femora unarmed or with double row of
The following abbreviations indicate in- tiny tubercles, and male genital capsule
stitutions where specimens are deposited or usually with small to large median projec-
from which material was generously lent: tion (Figs. 11-25). In Agathyrna, antennal
The Natural History Museum, London segments III and IV are subequal in length,
(BMNH); Bernice P. Bishop Museum, Hon- the head dorsally weakly convex, the anten-
olulu, Hawaii (BPBM); California Acade- niferous tubercles at outer apex obtuse, not
my of Sciences, San Francisco, California armed, posterior lobe ofpronotal disc with-
(CAS); Hungarian Natural History Muse- out transverse wrinkle, calli almost flat,
um, Budapest (HNHM); Natural History femora ventrally armed, and male genital
Museum ofLos Angeles County, California capsule obtusely rounded at apex, and never
(LACM); Queensland Museum, Brisbane, projecting into spine.
Australia (QMBA); Rijksmuseum van Na- Generic redescription. Head: Width
turlijke Histoire, Leiden, Netherlands across eyes greater than head length; qua-
(RNHL); Forschungsinstitut und Naturmu- drangular, dorsally convex to moderately
seum Senckenberg, Frankfurt am Main, globose; tylus extending anteriorly to juga,
Germany (SMFD); Coleccion Entomologi- apically upturned to form a sharp median
ca del Institute de Biologia, Universidad horn; juga unarmed, thickened, shorterthan
Nacional Autonoma de Mexico (UNAM); tylus; antenniferous tubercles armed with a
National Museum of Natural History, short and robust lobe; sides ofhead in front
Smithsonian Institution, Washington D.C. ofeyes unarmed, straight; antennal segment
(USNM); Zoologisches Museum, Hum- I robust, thickest, slightly curved outward;
boldt Universitat, Berlin, Germany segments II and III cylindrical, slender, and
(ZMHB); Zoologische Staatssammlung segment IV fusiform; antennal segment II
Miinchen, Germany (ZSMC). longest, IV shortest, and III longer than I;
All measurements are given in millime- ocelli weakly elevated, with a deep circular
ters. pit in front of each; eyes substylate, protu-
berant; postocular tubercle moderately pro-
Acanthotyla Stal tuberant; buccula short, elevated, angulate
behind anterior emargination, not projected
Acanthotyla Stal 1873: 68; Breddin 1900:
beyond antenniferous tubercles, with sharp
194; Blote 1936: 50; Brailovsky 1995:
93. Type species: Cletusfasciatus Walker posterior projection; rostrum reaching an-
1871: 196-197. Designation by Mono- terior third of abdominal sternite III or IV;
mandibular plates directly below apices of
typy.
juga, each with prominent tubercle.
—
Diagnosis. Like Agathyrna Stal, in this Thorax: Pronotum wider than long, tra-
genus the tylus projects in front of juga, peziform, moderately to strongly declivent,
upturned to form a small horn at its apex, and bilobed; anterior lobe shorter than pos-
the mandibular plates are directly below terior lobe; collar wide; frontal angles pro-
apices ofjuga and with small prominent tu- duced forward as small conical teeth some-
bercle, buccula short, raised and angulate times difficult to see; anterior half of an-
behind anterior emargination, and abdomi- terolateral margins convexly rounded, and
nal sternite VII of female entire, without posterior half obliquely straight; humeral
plica or fissura. In Acanthotyla, antennal angles rounded to obtuse, moderately to
segment IV is shorter than III, the head dor- conspicuously directed upward, and hardly
sally strongly convex, antenniferous tuber- prominent; posterolateral and posterior
cles at outer apical comer projected for- margins straight; calli conspicuously con-
ward, posterior lobe of pronotal disc with a vex to globose, separated along midline by
VOLUME NUMBER
107, 2 281
(^ C C Ct C C^,L\ , ^ c f
Figs. 1-10. Acanthotyla spp. 1-4. Pronotum. 1, A. fasciata. 2, A.flexuosa. 3. /\. kiilohoniia. 4. .4. kiiiiii^ala.
5-10, Parameres. 5-6, /\. protenta. 7-8, v4. distinguenda. 9-10. A. fasciata.
PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON
282
a longitudinal furrow, entirely flat, or sep- Features in Common for Species
arated by a convex longitudinal expansion; Described
posterior margin with transverse wrinkle Coloration: 1, Antennal segment I yel-
(Figs. 1-4); anterior lobe of metathoracic low speckled with pale brown or pale or-
peritreme reniform, posterior lobe sharp, ange irregular spots. 2, Hemelytral mem-
small; mesosternum with longitudinal fur- brane pale to dark brown with yellow veins.
row; propleura laterally convex to hemi- 3, Connexivum reddish brown with anterior
spheric, in dorsal view visible. Legs un- third and posterior margin yellow. 4, Pro-
armed; femora usually tuberculate; tibiae sternum, mesosternum, and metasternum
cylindrical, sulcated. black to reddish brown. 5, Anterior and
Scutellum: Triangular, longer than wide, posterior lobe of metathoracic peritreme
apically rounded to subacute; disc basally pale yellow.
markedly globose, and distally before the
apex remarkably depressed. Key TO Species of Acanthotyla
Hemelytron: Macropterous, almost
reaching apex of last abdominal segment; 1. Head in dorsal view almost entirely black to
reddish brown 2
apical margin sinuate to obliquely straight; - Head in dorsal view yellow with punctures
apical angle reaching almost middle third of reddish brown to chestnut orange 8
hemelytral membrane; costal margin emar- 2. Male 3
ginate. - Female 6
Abdomen: Connexivum higher than ter- 3. Posteroventraledgeofmalegenital capsuleat
middle third with a stout or slender and elon-
ga; connexival margin entire, and posterior gate projection obliquely directed upward
angle blunt, and not extended into spine; (Figs. 15-1.6, 19-20) 4
abdominal sterna with median longitudinal - Posteroventral edge of male genital capsule
furrow running to posterior third of sternite without stout or elongate projection (Figs.
13-14, 21-22, 25) 5
IV.
4. Posteroventral edge ofgenital capsuleatmid-
Male genital capsule: Usually with
dle third with a stout spinous conical projec-
small to large median projection (Figs. 11- tion; lateral angles of genital capsule with
15). Parameres simple, elongate, with apical short blunt processes (Figs. 15-16); humeral
third slightly curved (Figs. 5-10). angles broad, rounded, and not exposed . . .
Female genitalia: Abdominal sternite A. distinguenda Blote
- Posteroventral edge ofgenitalcapsuleatmid-
VII complete, without plica or fissura (Figs.
dle third with a relatively slender, elongate
26-35). conical projection, apically acute; lateral an-
Integument: Body surface rather dull, gles ofgenital capsule elevated, exposed, and
with short, decumbent to suberect indistinct apically quadrate (Figs. 19-20); humeral an-
pubescence; abdominal sterna with few gles weakly exposed, and narrowed ....
A. protenta, n.sp.
moderately long, and conspicuous bristle 5. Posteroventral edgeofgenital capsule atmid-
like hairs; pubescence ofantennae and fem- dlethirdwithtinyprojectiondirectedupward,
ora short, mainly suberect; pubescence of and sometimes hard to see (Figs. 13-14, 25);
tibiae and tarsi longer and rather dense; dor- intercallar space broad and flat; humeral an-
gles rounded (Fig. 1) A.fasciata (Walker)
sal and ventral surface of head, pronotum, - Posteroventral edge ofgenitalcapsuleatmid-
scutellum, clavus, corium, propleura, me- dle third broad, thick, without stout spinous
sopleura, metapleura, abdominal sterna, and projection directed upward (Figs. 21-22); in-
exposed parts ofgenital plates ofboth sexes tercallar space narrow, with a weakly longi-
strongly punctate, and each puncture deeply tudinal carina; humeral angles laminate ....
A. nabirenia, n.sp.
excavated; prosternum, mesosternum, and
6. Gonocoxae I with mesial marginbroadlycon-
metasternum smooth; antenna and legs mi- tiguous, and not emarginate; paratergite IX at
nutely granulate to tuberculate. lower third exposed, and distally bifurcate
VOLUME 107, NUMBER 2 283
Figs. 11-25. Male genital capsule ofAcanthotyla spp. 1 I. 13. LS. 17. 19. 21. 1?^. 25. Lateral \ic\v. 12. 14,
16, 18, 20, 22, 24, Caudal view. 11-12, A. borneana. 13-14, 25, A.fasciatu. 15-16, A. disiinintciula. 17-18. ,4.
kiungala. 19-20, A. protenta. 21-22, A. nabirenia. 23-24, A. kaloboana.
—
.
284 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON
(Figs. 29-30, 33. 35) IV, 1.28. Pronotum: Length 2.16; maximum
A. dislingiienda Blole, width of anterior lobe 1.84; maximum
A. fasciata (Walker), and A. protenta, n.sp. width of posterior lobe 3.20. Scutellar
- Gcoonntoicguooxuase, Iawnidthctohnespmiecsuioaulslmyargeimnarbgrionaadtley; length 1.44; width 1.32. Body length 9.62.
paratergite IX with lower third not exposed Female: Head length 1.60; width across
and distally truncated (Figs. 26. 31) 7 eyes 1.98; interocular space 1.18; preocular
7. Mesial margin ofgonocoxae I with the space distance 1.26; interocellar space 0.44;
between the border and the margin strongly length of antennal segments: I, 2.14; II,
depressed throughout the entire plate (Figs.
28, 32) A. kaloboima, n.sp. 3.12; III, 2.16; IV, 1.28. Pronotum: Length
- Mesial margin ofgonocoxae I with the space 2.40; maximum width ofanterior lobe 2.00;
between the border and the margin shallowly maximum width of posterior lobe 3.38.
depressed A. nabirenia. n.sp. Scutellar length 1.60; width 1.52. Body
8. Male 9 length 10.48.
- Female 12
Male: Dorsal coloration: Head yellow
9. Posteroventral edge ofgenital capsule at mid-
dle third broad, thick, without a stout spinous with punctures chestnut orange; antennal
projection directed upward (Figs. 23-24) . . . segments II and III yellow, and IV dark
A. kaloboana, n.sp. chestnut orange with apex paler; anterior
- Posteroventraledgeofgenitalcapsuleat mid- lobe of pronotal disk dark yellow, with
dlethirdwithstoutandbroadorelongatepro-
jection (Figs. 11-12, 17-18) 10 transversal fascia behind calli pale yellow,
10. Posteroventral edge ofgenitalcapsule at mid- and punctures reddish; intercallar space
dle third with a relatively elongate spinous dark brown with orange reflections; poste-
conical projection, directed posteriorly and rior lobe ofpronotal disk pale orange brown
upward (Figs. 17-18)
with punctures reddish; scutellum yellow
A. kiungala, n.s.p.
- Posteroventral edge ofgenital capsule atmid- with punctures chestnut orange and basal
dle third with broad projection (Figs. 11-12) angle black; clavus and corium pale brown,
11 with punctures reddish brown to chestnut
11 Posteroventral edge ofmale genital capsuleat orange, and claval and corial veins, and
middle third with stout quadrate projection,
costal margin yellow; apical margin pale
apically bifid, and clearly directed upward . . brown with yellow marks; dorsal abdominal
A.flexitosa (Blote)
- Posteroventral edge ofgenitalcapsule at mid- segments dark orange with black marks.
dlethirdwithstout,roundedtubercle,directed Ventral coloration: Head yellow with
posteriorly and upward (Figs. 11-12) . . . punctures chestnut orange, and middle third
A. bonieana, n.sp.
with broad longitudinal black stripe; rostral
12. Mesial margin ofgonocoxae I with the space
between the border and the margin strongly segment I yellow with chestnut orange re-
depressed throughout the entire plate (Figs. flections, and segments II to IV chestnut or-
26, 31) A. kaloboana, n.sp. ange; propleura, mesopleura, and metapleu-
- Lowerandmesial marginofgonocoxaeIwith ra yellow with punctures chestnut orange;
the space between the border and the margin
coxae dark reddish brown; trochanters yel-
shallowly depressed (Figs. 27-28, 32 34)
A. borneana, n.sp., A.flexiiosa (B.l.ote), low; fore and middle femora yellow, speck-
andA. kiungala, n.sp. led with pale brown discoidal spots, and
with subdistal pale brown ring; hind femur
Acanthotyla borneana Brailovsky,
yellow, speckled with pale brown discoidal
new species spots, and with two pale brown rings, one
(Figs. 11-12, 27, 34) near middle third and the other subdistal;
Description. Measurements: Male: tibiae pale brown with two yellow rings,
Head length 1.50; width across eyes 1.88; one subbasal, the other near middle third;
interocular space 1.12; preocular distance tarsi pale chestnut yellow; middle third of
1.22; interocellar space 0.43; length of an- abdominal sterna dark reddish brown, and
tennal segments: I, 2.08; II, 2.96; III. 2.08; laterally yellow with punctures reddish
VOLUME 107, NUMBER 2 285
£K
^T^
Figs. 26-35. Female genital plates ofAcanthoryla spp. 26-30, Caudal view. 31-35, Lateral \ic\\. 26. 31. .\.
kaloboana. 27, 34, A. borneana. 28, 32, A.flexiiosa. 29, 33, A. dislingiteiula. 30, 35, A. protvnta.
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286 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON
brown to chestnut orange; genital capsule shape of the male genital capsule is distinct
reddish brown. (Figs. 13-16, —25).
Head: Rostrum reaching anteriorthird of Etymology. Named for its occurrence
abdominal sternite IV. in Borneo.
Thorax: Humeral angles subacute, ele-
vated, slightly laminate, and higher than Acanthotyla distinguenda Blote
posterior pronotal disk; intercallar space (Figs. 7-8, 15-16, 29, 33, 36)
narrow with prominent carina. Acanthotyla distinguenda Blote 1936: 50.
Genital capsule: Posteroventral edge
transversely concave; lateral angles straight Redescription. Measurements: Male:
not expanded; middle third with stout and Head length 1.40-144; width across eyes
rounded tubercle, directed posteriorly and 1.97-2.01; interocular space 1.24-1.34;
clearly upward (Figs. 11-12). preocular distance 1.08-1.12; interocellar
Female: Color similar to male. Connex- space 0.38-0.44; length of antennal seg-
ival segments VIII and IX dark chestnut or- ments: I, 1.62-1.64; II, 2.40-2.52; III,
ange with reddish brown reflections; dorsal 1.76-1.97; IV, 1.20-1.32. Pronotum:
abdominal segments VIII and IX dark Length 1.76-1.84; maximum width of an-
chestnut orange; genital plates yellow with terior lobe 1.40-1.76; maximum width of
punctures chestnut orange to reddish posterior lobe 2.64-2.76. Scutellar length
brown, and with middle third of gonocoxae 1.42-1.44; width 1.16-1.24. Body length
I and outer margin ofparatergite IX reddish 8.45-9.30. Female: Head length 1.52-1.56;
brown. width across eyes 2.12-2.14; interocular
Genital plates: Gonocoxae I enlarged space 1.33-1.36; preocular distance 1.16-
dorsoventrally; mesial margin broadly con- 120; interocellar space 0.40-0.46; length of
tiguous and weakly emarginated; parallel to antennal segments: I, 1.60-1.68; II, 2.48-
each margin at lower and middle third shal- 2.50; III, 1.79-1.84; IV, 1.1.30-1.32. Pron-
lowly depressed, with upper third overlap- otum: Length 2.08-212; maximum width of
ping; paratergite VIII small, triangular, with anterior lobe 1.44—1.80; maximum width of
spiracle visible; paratergite IX not overlap- posterior lobe 2.82-2.84. Scutellar length
ping at middle third, and larger than para- 1.52-1.54; width 1.30-1.32. Body length
tergite VIII (Figs.—27, 34). 10.15-10.38.
Type material. Holotype: S, Borneo Male: Dorsal coloration: Black with ty-
(British N.), Sandakan Bay (SW), Sapagaya lus and postocular tubercle yellow; antennal
Lumber Camp, 2-20 m, 4 November 1957, segments II and III pale yellow, and IV pale
J. L. Gressitt (BPBM). Paratype: 1 9, same yellow with basal joint pale brown; jugum
data as the male—holotype (BPBM). yellow with punctures reddish brown; pron-
Distribution. Only known from the otum dark to pale yellow with punctures
type material. chestnut orange and intercallar space black;
—
Discussion. This species is diagnosed scutellum pale yellow with punctures chest-
mostly on the basis ofthe shape ofthe pos- nut orange, and basal angle black; clavus
teroventral edge of male genital capsule and corium pale yellow brown with punc-
(Figs. 11-12), the intercallar space narrow tures reddish brown; claval and corial veins,
with prominent carina, the humeral angles and costal and apical margin pale yellow;
subacute and slightly laminate, and the head dorsal abdominal segments dark orange.
mostly yellow with punctures chestnut or- Ventral coloration: Head black with two
ange. In Acanthotyla distinguenda and A. short longitudinal yellow stripes lateral to
fasciata the intercallar space is broad and middle line and close to eyes; buccula yel-
flat, the humeral angles are rounded and not low with punctures chestnut orange; rostral
laminate, the head is mostly black, and the segments pale chestnut orange; propleura,
VOLUME 107, NUMBER 2 287
Fig. 36. Dorsal view oiAcanthotyla distinguenda, male.
—
PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON
288
mesopleura, and metapleura yellow with III pale chestnut orange. 3, Antennal seg-
punctures chestnut orange; coxae dark red- ment IV chestnut orange with basal joint
dish brown with apex yellow; trochanters dark brown. 4, Hemelytral membrane dark
yellow; fore and middle femora yellow, brown to black with veins pale yellow. 5,
speckled with pale brown discoidal spots, Outer margin of calli black. 6, Trochanter
and with subdistal pale brown ring; hind fe- yellow with tiny—pale brown stripe.
mur yellow, speckled with pale brown dis- Distribution. This species is known
coidal spots, and with two pale brown rings, only from New Guinea. The only previous-
one near middle third and other subdistal; ly known record came from the original de-
tibiae pale brown with two yellow rings, scription in which the holotype, allotype
one subbasal, other near middle third; tarsi and paratypes were collected: Indonesia:
pale chestnut yellow; middle third of ab- Irian Jaya (Dutch New Guinea): Manok-
dominal sterna dark reddish brown, and lat- wari, Andai, Hattam, Assiki on Digul, Kop-
erally yellow with punctures reddish brown stein, and Sekroe (Blote 1936). The new re-
to chestnut orange; genital capsule reddish cords listed below show it occurs exten-
brown. sively in New Guinea.
Head: Rostrum reaching posterior mar- Material examined.—1 c?, INDONESIA:
gin of abdominal sternite III. Irian Jaya (Dutch New Guinea), Manok-
Thorax: Humeral angles rounded, slight- wari, J. W. van Nouhuys (holotype RNHL).
ly prominent, elevated, and higherthan pos- PAPUA NEW GUINEA: 2 S, Bisianumu,
terior pronotal disk; intercallar space broad E. of Port Moresby, 500 m, 23 September
and flat. 1955, J. L. Gressitt (BPBM); 1 cJ, 1 9,
Genital capsule: Posteroventral edge Middle Fly River, 250-300 m, July 1928,
transversely straight or slightly concave; Pemberton (BPBM); 1 S, Morobe, D. Bul-
lateral angles with short blunt processes; log Rd., S. Slopes Yaningya, 600-1300 m,
middle third with stout spinous conical pro- 19-20 March 1968, Reni (BPBM); Sogeri,
jection, obliquely directed upward (Figs. 27 October 1968, Tawi and Mena (BPBM).
15-16). Parameres in Figs. 7-8. Indonesia: Irian Jaya: 1 S,l 9, Jayawijaya
Female: Color similar to male. Connex- Prov., Brazza River, Dekai, ca. 100 m, 21-
ival segments VIII and IX yellow with pos- 22 June 1994, A. Riedel (ZSMC); 1 9, Jay-
terior third reddish brown; dorsal abdomi- awijaya Prov., Samboka, upper KoiffRiver,
nal segments VIII and IX dark reddish ca. 200 m, 10-14 October 1996, A. Riedel
brown; genital plates yellow with punc- (ZSMC); 1 S, Jayawijaya Prov., Kec Wai-
tures, posterior third ofparatergite VIII and geo Sel., Gamang Isl., 0-100 m, 26 No-
IX, and middle third of gonocoxae I dark vember 1996, A. Riedel (ZSMC); Merauke
reddish brown. Prov., Asmat-Patipi, Brazza River, 100 m,
Genital plates: Gonocoxae I reniform, A. Riedel (USNM); 1 S, \ 9, Manokwari
enlarged dorsoventrally, with deep depres- Prov., Wasior, 0-500 m, 8-12 January
sion close to middle third; mesial margin 2001, A. Riedel (UNAM); 5 9, Manok-
broadly contiguous and not emarginate; wari, Ransiki, Mayuby, 300 m, 26-30 Sep-
paratergite VIII and IX totally exposed; tember 1990, A. Riedel (ZSMC); 1 d, Na-
paratergite VIII small, triangular, with spi- bire, Pusppensaat km. 60, 200 m, 15 Au-
racle visible; paratergite IX not overlapping gust 1991, A. Riedel (ZSMC).
at middle third, larger than paratergite VIII, Discussion. Acanthotyla distingiienda
strongly carinated, exposed, and reflexed, is easily distinguished by the shape of the
with lower t—hird bifurcate (Figs. 29, 33). posteroventral edge ofthe male genital cap-
Variation. 1, Antennal segment I pale sule which shows a stout conical obliquely
chestnut orange, speckled with pale brown erected projection, with short blunt pro-
irregular spots. 2, Antennal segments II and cesses at lateral angles (Figs. 15-16). In A.