A revision of Meigen's Chloropidae collection in the Museum National d'Histoire Naturelle, Paris (Diptera)

2006·0.79 MB·

by  Nartshuk, E P

© Zoological Institute, St.Petersburg, 2006
A revision of Meigen's Chloropidae collection
in the Museum National d'Histoire Naturelle, Paris (Diptera)
E.P. Nartshuk
Nartshuk, E.P. 2006. A revision ofMeigen's Chloropidae collection in the Museum Na
tional d'Histoire Naturelle, Paris (Diptera). Zoosystematica Rossica, 15(1 ): 173-184.
Chloropidae ofMeigen 's collection in Paris ( 54 species) are examined, including 35 spe
cies described by Meigen (1830, 1838). New synonymy is established: Oscinella nigerri
ma (Macquart, 1835) = Ch/orops nigritus Meigen, 1838; Lasiambia palposa (Fallen,
1820) = Chlorops ater Meigen, 1838; Rhopalopterum anthracinum (Meigen, 1830) =
Chlorops glaberrimus Mei gen, 1838; Oscinella vindicata (Meigen, 1830) = 0. hortensis
Collin, 1946; Meromyza variegata Meigen, 1830 = M lidiae Nartshuk, 1992; M. femo
rata Macquart, 1835 = M. laeta Meigen, 1838. Ch/orops lucida Meigen, 1838 and C.
vindicata (Meigen, 1830) are restored from synonymy and considered to be good species
Thaumatomyia lucida (Meigen) and Oscinella vindicata (Meigen), respectively. Lecto
types are designated for following Meigen's species: Eurina lurida, Chlorops calceatus,
C. hypostigma, C. geminatus, C. cingulatus, C. pygmaeus, C. simplex, C. lucidus, C.
femoralis, C. sulcicollis, C. glaberrimus, C. fascipes, C. nitidissimus, Meromyza prato
rum, M. variegata, and M. /aeta. The following currently accepted synonymy is con
firmed by examination of Meigen's types: Chlorops strigulus (Fabricius, 1794) = C. cin
gulata Meigen, 1830; Thaumatomyia notata (Meigen, 1830) = C. circumdata Meigen,
1830; T. rufa (Macquart, 1835) = C. simplex Meigen, 1838; Siphonella oscinina (Fallen,
1820) = C. angustifrons Meigen, 1830; Elachiptera comuta (Fallen, 1820) =C. femora
lis Meigen, 1838; Rhopalopterum anthracinum (Meigen, 1830) =C. glaberrimus Mei
gen, 1838; Rhopalopterumfasciolum (Meigen, 1830) = C. fascipes Meigen, 1838. Types
of 9 species described by Meigen are found neither in MNHP, nor in NHMW. A new
species Meromyza meigeni sp. n. is described for M. laeta sensu Ismay (1981) and Nartshuk
(1992). Meromyza variegata sensu Ismay (1981 ), Beschovski (1985) and Nartshuk (1992)
is considered to be M. athletica Fedoseeva, 1974, and a redescription of this species is
given.
E.P. Nartshuk, Zoological Institute, Russian Academy ofS ciences, Universitetskaya nab.
1. St.Petersburg 199034. Russia. E-mail: ch/[email protected]
This paper is the next attempt to clarify the iden has seen Meigen's type of Chlorops pygmaea
tity of Chloropidae described by earlier authors. (Andersson, 1966).
Preceding ones dealt with species described by Meigen's Chloropidae collection in MHNP
Th. Becker and 0. Duda, C. von Roser, J.W. is kept in drawers 55, 59, and 60 separately
Meigen and other authors (Nartshuk, 1970, 1994, from other collections of Chloropidae. Most
1997a, 1997b). Becker (1902) revised the types specimens are pinned, a few are glued on pa
of species described by J.W. Meigen in the Mu per 2 x 5 mm. Some glued specimens are dam
seum National d'Histoire Naturelle in Paris aged, and it is believed that they were pinned
(MHNP in the following text) and Naturhistor previously. Each specimen described in 1830
isches Museum in Vienna (NHMW in the fol has a square label on brownish paper with the
lowing text). Later, I (Nartshuk, 1997) revised names of the genus and species in Meigen's
Meigen 's types in NHMW with new synonyms handwriting; species described or listed by
and designation of lectotypes. In 2002, I had an Meigen in 1838 have on the label only the spe
opportunity to examine Meigen 's collection in cific epithet without the generic name. In ad
MHNP in course of preparing the Chloropidae dition, every specimen has a round label with
part for the project "Fauna Europaea". After Th. the word "Meigen" and a printed number, and
Becker, Meigen's collection in MHNP was ex each specimen in drawers 59 and 60 has an
amined by E. Seguy; his paper on Chloropidae other number on the back of this label. The lat
(Seguy, 1934) contained a statement "sec. typ." ter numbers are accompanied by figures 40
for 10 Meigen's species. H. Andersson has also showing registration in 1840, they are listed
seen the collection (pers. comm.), but did not by me in parentheses. There are also collec
publish the results. He mentioned only that he tion labels with the names of the genus and
174 E.P Nartshuk: Meigen s Ch/oropidae in Paris • ZOOSYST. ROSSICA Vol. 15
species. Some specimens have additional la and hind legs; other parts in good conditions.
bels "Museum Paris coll." or "Museum Paris Meigen mentioned "mein Exemplar" from vicin
Meigen coll.". ity of Paris. Therefore the examined specimen is
There are 51 collection labels in Meigen's interpreted as holotype. - Current interpretation:
Chloropidae with 1-4 flies, pinned or glued, af a good species Chlorops graci/is Meigen.
ter each. All flies after the collection label are Chlorops calceata Meigen, 1830: 146. No.
lost only in two cases. The collection contains 2522 (2721 40). One female, corresponds to the
35 species described by Meigen in 1830 and 1838 description and recent treatment of the species.
and 16 species described by other authors: Lin It is designated here as lectotype. First flagel
naeus, Fallen, Fabricius and Macquart. lomere on inner side yellow in central part, not
Species are discussed in the following text entirely black. Specimens in NHMW do not cor
according to their arrangement in drawers. Ar respond to the description (Nartshuk, 1997a).
rangement of Chlorops species in Becker (1902) There is one more label "Museum Paris coll.
was different, probably the material had been Meigen." - Current interpretation: a good spe
moved into other drawers. Already Becker (1902) cies Ch/orops calceatus Meigen.
did not find types of Chlorops f asciatus Meigen, Chlorops ornata Meigen, 1830: 152. No.
1830, C. gentilis Meigen, 1830, C. confluens 2523. No specimen, only collection label and two
Meigen, 1830, C. ruficeps Meigen, 1830, C. holes from pins. Becker (1902) mentioned only
geniculatus Meigen, 1838 (described from "coll. one specimen in NHMW and did not mention
Baumhauer"; the type is probably in Berlin), C. any specimen in MHNP. - Current interpretation:
varipes Meigen, 1838, C. pictus Meigen, 1838, a junior synonym of Thaumatomyia notata (Mei
C. amoenus Meigen, 1838, C. albitarsis Meigen, gen, 1830: 144).
1838, and C. vagans Meigen, 1838. Now, there ChloropshypostigmaMeigen, 1830: 141. No.
are no collection labels for these species, except 2524 (2723 40). Four specimens, two of them
C. vagans. All these species, except C. vagans, (male and female) on one pin, both without first
are listed as nomina dubia in the "Catalogue of flagellomere. One male without head and one
Palaearctic Diptera" (Nartshuk, 1984). Lyneborg more specimen with abdomen in small tube on
(1965) listed C. geniculata as a synonym of Thau the same pin, both with additional label "Muse
matomyia obscurella (now it is T. hallandica um Paris coll." All specimens correspond to the
Andersson), but without special explanation. description. The male on the pin with the female
is here designated as lectotype; the female is a
Drawer55 paralectotype. Becker ( 1902) mentioned only two
specimens on a pin. - Current interpretation: a
Eurina lurida Meigen, 1830: 5. No. 2363. One good species Chlorops hypostigma Meigen.
male, in good condition, only right first flagel Chlorops geminata Meigen, 1830: 141. No.
lomere absent. The species was described from 2525 (2724 40). One female in good condition,
"Baumhauerischen Sammlung", type locality designated here as /ectotype. There are males in
unknown. The specimen corresponds to the de Winthem's collection in NHMW, but they are
scription. It is designated here as lectotype; there labelled "Halle" and do not belong to the type
is no material in NHMW (Nartshuk, I 997a). Pont series (Nartshuk, 1997a). - Current interpreta
(1986: 202) wrote "I can see no reason for not tion: a good species Chlorops geminatus Mei
regarding specimens in his [Meigen's] collection gen.
in Paris of the species described from Baumhau Chlorops nasuta. No. 2526 (2724 40 and 2725
er 's collection as being syntypes, if they agree 40). One female in good condition and the sec
with the original description". - Current inter ond specimen (only thorax) are C. meigenii Loew
pretation: Eurina lurida Meigen. (darkened wings and yellow spot on katepister
Camarota flavitarsis Meigen, 1830: 7. No. num). Meigen 's specimen is not a type of Ch.
2365. One female (Becker, 1902 recorded one nasuta, and there is no reason to change the cur
male), corresponds to the description. This spec rent interpretation. - Current interpretation: Mus
imen is a paralectotype, the lectotype is desig ca nasuta Schrank, 1781 is a junior synonym of
nated from Winthem's collection in NHMW Chlorops pumilionis (Bjerkander, 1778). Loew
(Nartshuk, 1997a). - Current interpretation: a (1866) correctly named Chlorops nasuta sensu
junior synonym of Camarata curvipennis (La Meigen as C. meigenii Loew.
treille, 1805). Chlorops scalaris Meigen, 1830: 145. No.
2527 (2726 40). One female without antennae
Drawer59 and right wing, corresponds to the description:
enlarged palpi, black shining spot on katepister
Chlorops gracilis Meigen, 1830: 140. No. num, ocellar triangle black with narrow yellow
2521 (2720 40). One specimen without abdomen sides and yellow hind corners. This female is a
ZOOSYST. ROSSICA Vol. 15 • E.P. Nartshuk: Meigen :S Chloropidae in Paris 175
paralectotype, the lectotype is designated from Cblorops lineata. No. 2534 (2733 40). Two
Winthem's collection in NHMW (Nartshuk, females, one of them with label "Ch/oropisca
1997a). - Current interpretation: a good species glabra Meig. ( det. Th. Becker, 1901 )", specimen
Chlorops scalaris Meigen. in good condition but without right first flagel
Cblorops speciosa Meigen, 1830: 146. No. lomere. Another female without head, has addi
2528 (2727 40). One female, ofrather large size, tional text''\'. Fall." on the label. Both are Thau
corresponds to the description, paralectotype. The matomyia glabra (Meigen). The specimens are
lectotype is designated from Winthem's collec not types. Becker (1902) mentioned only a fe
tion in NHMW (Nartshuk, 1997a). - Current in male in Paris as Ch/oropisca glabra. - Current
terpretation: a good species Chlorops speciosus interpretation: Musca lineata Fabricius, 1781 is
Mei gen. considered now a junior synon)'m of Chlorops
Cblorops cingulata Meigen, 1830: 146. No. pumilionis (Bj erkander, 1787). According to Zim
2529 (2728 40). One female without antennae, sen (1964), Fabricius's type is lost: "Kiel only
with other parts in good conditions, corresponds the namelabel".
to the description. It is designated here as lecto Cblorops circumdataMeigen 1830: 147. No.
type. - Current interpretation: junior synonym of 2535 (2734 40). One female without head, dam
Chlorops strigulus (Fabricius, 1794); synonymy aged by insects. Scutellum flat, apical setae ap
confirmed here. proximated, stripes on scutum as in Thaumato
Chlorops strigula. No. 2530 (2729 40). Two myia notata Meigen. The specimen is designat
females, both are C. pumilionis Bjerkander, 1778, ed here as /ectotype, it corresponds to the descrip
one oft hem has an additional label "Museum Paris tion. Meigen described the species from Win
coll." Meigen's specimens are not types, there is them's collection, but specimens in Winthem's
no reason for nomenclature change. - Current collection in NHMW do not belong to the type
interpretation: a good species Chlorops strigula series (see Nartshuk, 1997a, p. 393). - Current
(Fabricius, 1794). Note: According to Zimsen interpretation: junior synonym of Thaumatomyia
(1964), the type is in MHNP, but I have not seen notata (Mei gen, 183 O); synonymy confirmed
it. Seguy (1934) listed "Ch/orops strigula Mei here.
gen (sec. typ. )"as a synonym of Chlorops pumil Chlorops limbata Meigen, 1830: 150. No.
ionis Bjerkander, and "Chlorops cingulata Mei 2536 (2735 40). On the label, only "limbata ",
gen (sec. typ. )"as a synonym of Chlorops strigu without generic name. The specimen, a female,
la Fabricius, but he did not mention Fabricius type. glued, is Lasiosina brevisurstylata Dely-Drask
Oscinis cereris. No. 2531 (2730 40). Three ovits, 1977. Ocellar triangle with shining rhom
specimens, one of them is glued. Two females boid spot on the tip; legs entirely yellow; only
with dark arista in good condition, one of them one hind notopleural seta; first flagellomere
has additional words "v. Fall." on the label, both black; palpi not seen. The species was described
are Cetema spp. One male is Cetema myopinum from Winthem's collection, and the lectotype of
Loew, specimen in good condidion. All speci C. limbatus Meigen was designated from collec
mens are not types. - Current interpretation: Ce tion in NHMW (Nartshuk, 1997a). - Current in
tema cereris (Fallen, 1820). The lectotype is in terpretation: a good species Ch/orops limbatus
Zoological Museum of Lund University (Anders Meigen, 1830, a senior synonym of C. brevi
son, 1963). manus Loew, 1866 (Nartshuk, 1997a).
Chlorops messoria. No. 2532 (2731 40). One Cblorops tarsata. No. 2537 (2736 40). One
female in good condition, is Diplotoxa messoria female in good condition, is Cryptonevra jlavi
(Fallen, 1820). The specimen is not a type. - tarsis (Fallen, 1820). The specimen is not a
Current interpretation: Diplotoxa messoria (Fal type. - Current interpretation: a good species
len, 1820). The lectotype is in Zoological Muse Neohaplegis tarsata (Fallen, 1820). The lecto
um of Lund University (Andersson, 1963). type is in Zoological Museum of Lund Universi
Chlorops taeniopa Meigen, 1830: 144. No. ty (Andersson, 1963).
2533 (2732 40). One female, without left flagel Chlorops pygmaea Meigen, 1838: 385. No.
lomere and part oflegs and with damaged right 2538 (2737 40). One male, glued, but was prob
wing. It is a light coloured specimen of C. pumi ably pinned previously; right side of thorax dam
lionis (Bjerkander, 1787); ocellar triangle most aged. Cheeks produced, stripes on scutum brown
ly yellow, except for black ocellar tubercle and ish. The specimen is here designated as lecto
middle line, other characters correspond to C. type. - Current interpretation: a good species
pumilionis. It is a paralectotype, the lectotype is Trachysiphonella pygmaea (Meigen, 1838). An
designated from Winthem's collection in NHMW dersson {1966), who has seen Meigen's type,
(Nartshuk, 1997a). - Current interpretation: a considered T. pygmaea to be a good species.
junior synonym of Ch/orops pumilionis (Bjer Chlorops simplex Meigen, 1838: 385. No.
kander, 1787). 2539 (2738 40). One specimen, glued, tip of ab-
176 E.P Nartshuk: Meigen s Chloropidae in Paris • ZOOSYST. ROSSICA Vol. 15
1 3
4
Figs 1-4. Thaumatomyia. 1, 2, T simplex (Meigen), !ectotype (1, antenna; 2, scutum); 3, 4, T /ucida (Meigen), lecto
type (3, antenna; 4, frons).
domen not seen (female, according to Becker, belongs to Lasiambia palposa (Fallen, 1820). It
1902), additional label "Museum Paris Mei gen". is not a type. - Current interpretation: a good
The specimen has strongly damaged right side species Lasiambia palposa (Fallen, 1820). The
of head and thorax; left wing and scutellum ab lectotype is in Riksmuseum in Stockholm (An
sent. Median stripe on scutum does not extend dersson, 1963 ).
to scutellum and red in the middle (Fig. 2). Ocel Chlorops phaeoptera Meigen, 1830: 153. No.
lar triangie yellow, extending to front of frons, 2542 (2741 40). One specimen without tip of
with one row of setulae along sides. First flagel abdomen (Becker, 1902 listed the specimen as
lomere longer than deep, yellow, darkened above female) and right flagellomere, corresponds to
(Fig. 1) . Meigen stated: "Schildchen gelb, flach". the description. The specimen is a paralectotype.
The ~En :is des:ignatB'.:l. here as lectotype. - The iectotype is designated from the NHMW
Current interpretation: junior synonym of Thau collection (Nartshuk, l 997a). -Current interpre
matomyia rufa (Macquart, 1835); synonymy con tation: junior synonym of Capnoptera scutata
firmed here. (Rossi, 1790).
Chlorops lucida Meigen, 1838: 384. No. 2540 Chlorops rufipes Meigen, 1830: 154. No.
(2739 40). One female, with label "lucida" (with 2543 (2742 40). On the back side of the Mei
out generic name), and additional label "Muse gen 's label is a striked out inscription "Chlorops
um Paris Meigen". The specimen corresponds to frontelia". It is a female with entirely yeliow legs,
the description. Meigen stated: "Schildchen gelb, belonging to Conioscine!la mimula Collin. Tnis
flach." Ocellar triangle shining, extends nearly specimen does not correspond to the description.
to front of frons, yellow, but ocellar tubercle, The species was described from Winthem's col
sides and middle line black. There are a small lection and the lectotype was designated from
ridge on apical part of triangle and two rows of NHMW (Nartshuk. l997a). - Current interpre
setulae along sides (Fig. 4). Cheeks narrow, as tation: junior synonym of Oscinisoma cognatum
fore tibia. First flagellomere round, black (Fig. (Meigen, 1830).
3). Stripes on scutum black; median stripe does Chlorops laevigata. No. 2544 (2743 40).
not extend to scutellum. Scutellum flat; apical Specimen without head and abdomen, only tho
setae approximated. The specimen is designated rax remained on the thick pin. Thorax black, shin
here as lectotype. It is a species of Thaumato ing. Legs dark; halteres yellow. Becker (1902)
myia, very similar to T. notata (Meigen, 1830), mentioned a strongly damaged specimen. - Cur
but small ridge on apical part of ocellar triangle rent interpretation: Madiza laevigata Fallen, 1820
distinguish it from T notata. Therefore, I prefer is a junior synonym of Neohaplegis tarsata (Fal
to consider it a good species Thaumatomyia luc len, 1820).
ida (Meigen, 1838); the species name is restored Chlorops pusilla Meigen, 1830: 157. No. 2545
from synonymy with T notata. Becker (1902) (2744 40). One female, without head, right wing
wrote that Chlorops lucida is not identical with and part of legs. Only a middle leg, a hind leg
Chloropisca ornata (a synonym of Thaumato and another hind femur are preserved. Middle
myia notata). tibia entirely yellow; hind leg black, except for
Chlorops palposa. No. 2541 (2740 40). One tarsus. Scutum dusted; notopleural setae l +2 .
female, with additional inscription "von Fallen", Becker (1902) mentioned one damaged speci-
ZOOSYST. ROSSICA Vol. 15 • E.P. Nartshuk: Meigen:S Chloropidae in Paris 177
men. It is a specimen of Oscinella; it is strongly Current interpretation: Chlorops ater Meigen,
damaged and cannot help to clarify the case with 1838, syn. n. is a junior synonym of Lasiambia
Oscinella pusilla. See Nartshuk, l 997a: 401. palposa (Fallen, 1820).
Chlorops lepida Meigen, 1830: 157. No. 2546 Chlorops brevipennis Meigen, 1830: 152. No.
(2745 40). One specimen without abdomen, right 2552 (2751 40). Two specimens, one female (or
wing and hind legs. Fore and middle femora black male?) pinned, and another female glued. Both
with yellow apex, tibiae and tarsi yellow, but the specimens with short wings, in good condition;
last segment slightly darkened. - Current inter they are paralectotypes. The lectotype is desig
pretation: a junior synonym of Rhodesiella plu nated from Winthem's collection in NHMW
miger (Meigen, 1830). The lectotype is designat (Nartshuk, l 997a). - Current interpretation: a
ed from the NHMW collection (Nartshuk, good species Elachiptera brevipennis (Meigen).
1997a); it has the last segment of tarsi darkened Chlorops albiseta Meigen, 1830: 153. No.
as well. 2553 (2752 40). Two specimens. One female in
Chlorops angustifrons Meigen, 1830: 157. good condition, except absence of right wing; the
No. 2547 (2746 40). One female, designated here white arista is well seen. The other female with
as lectotype. It corresponds to the description. - broken both first flagellomeres, but wings dark
Current interpretation: junior synonym of Si ened, legs black as in Oscinella maura (Fallen,
phonel/a oscinina (Fallen, 1820); synonymy con 1820). I consider both females to be paralecto
firmed here. types, although Becker (1902) mentioned no
Chlorops cornuta. No. 2548 (2747 40). One specimens in MHNP. The lectotype is designat
male of Elachiptera cornuta (Fallen, 1820) with ed from Winthem's collection in NHMW
yellow legs. Another male is E. diastema Collin, (Nartshuk, 1997a). - Current interpretation: a
1946; apical setae widely spaced; dusting before junior synonym of Oscinella maura (Fallen,
scutellum is not seen, as the scutum is covered 1820).
with artificial dust. Both specimens are not Chlorops maura. No. 2554 (2753 40). Only
types. - Current interpretation: Elachiptera cor "maura" on the label, without generic name. One
nuta (Fallen, 1820). The lectotype in Zoological female, pinned, is a species of Agromyzidae.
MuseumofLund University(Andersson, 1963). Another female, glued, a specimen in good con
Chlorops femoralis Meigen, 1838: 390. No. dition, but without fore legs; middle tibiae yel
2549 (2748 40). One male, glued, in good con low; hind tibiae black. It is Oscinella frit (Lin
dition, only right apical seta of scutellum bro naeus, 1758), s. I. Both specimens are not types.
ken. It is Elachiptera cornuta (Fallen, 1820) with Becker (1902) listed one female in Paris. - Cur
black femora. The specimen is here designated rent interpretation: Oscinella maura (Fallen,
as lectotype (Meigen listed two specimens). - 1820).
Current interpretation: junior synonym of Ela Chlorops vindicata Meigen, 1830: 160. No.
chiptera cornuta (Fallen, 1820); synonymy con 2555 (2754 40). Three specimens. One female
firmed here. E. cornuta is a widespread species pinned, in good condition, with hind tibia black,
rather variable in the colour of legs and thick fore and middle tibia black with both ends yel
ness of the arista. Additional studies are needed low corresponds to the description and is Oscinel
of quite many described related species and forms la hortensis Collin, 1946. Another female, glued,
to clarify their taxonomic status. in good condition, with darker legs, is 0. frit (Lin
Chlorops minuta. No. 2550 (2749 40). One naeus, 1758), s. 1. The third specimen is a male
female, glued, in good condition, is Oscine/la frit of 0. angustipennis Duda, it has an additional
(Linnaeus, 1758), s. I. Meigen described no spe label "Museum Paris coll." with glued red square
cies with the name minuta in Chloropidae and 3 x 3 mm. Becker (1902) mentioned "in Pariser
describedEphydra minuta Meigen, 1830: 124 in Museum 1 ~ ". The lectotype was designated from
Ephydridae. There is no species minuta in Mei Winthem's collection in NHMW (Nartshuk,
gen 's Ephydridae collection in MHNP. Becker 1997a). Only one pinned specimen (female) is
(1902: 312) mentioned no Chlorops minuta, only similar to the lectotype and here considered a
Ephydra minuta specimen in NHMW. paralectotype. C. vindicata Meigen, 1830 was
Chlorops atra Meigen, 1838: 388. No. 2551 listed as a synonym of Oscinella pusilla (Mei
(2750 40). One female, glued, with additional gen, 1830) by Nartshuk (1984). Now, the spe
label "Paris Museum, coll. Meigen". The speci cies is restored from synonymy and considered a
men corresponds to the description and belongs good species. - Current interpretation: Oscinel
to Lasiambia palposa (Fallen, 1820). npl 1+1, la vindicata (Meigen, 1830) is a good species
wing cell ha not widened. This specimen is des and 0. hortensis Collin, 1946, syn. n. is its jun
ignated here as /ectotype. Synonymy with Poly ior synonym. I did not synonymize these species
odaspis sulcicol/is (Meigen, 1830) given by Duda earlier (Nartshuk, 1997a), before examination of
(1932-33) and Nartshuk (1984) is incorrect. - specimens in MHNP.
178 E.P. Nartshuk: Meigen s Chloropidae in Paris • ZOOSYST. ROSSICA Vol. 15
Chlorops frit. No. 2556 (2755 40). Label only rent interpretation: junior synonym of Rhopal
".frif', without generic name. Two specimens, opterum fasciolum (Meigen, 1830); synonymy
both glued. A male with very narrow cheek is confirmed here.
Oscinella cariciphila Collin, 1946; in female Chlorops nitidissima Meigen, 1838: 388. No.
cheeks are not seen, fore and middle legs black. 2562 (2761 40). Only "nitidissima", without ge
There is a third pin with a piece ofp aper but with neric name, on the label. There is an additional
out fly. The specimens are not types. Becker label "Museum Paris coll. Meigen". One female,
( 1902) listed only one specimen. - Current inter glued, in rather good condition, only left flagel
pretation: Oscinellafrit (Linnaeus, 1758). lomere and part of right arista broken. Scutum
Chlorops lineella. No. 2557 (2756 40). There shining, cheek nearly as first flagellomere, all
is a mark of female ( 9) on the label. One female femora black, fore tibia slightly darkened in the
in good condition, but apical setae of scutellum middle, middle tibia yellow, hind tibia black, tarsi
broken. Scutellum black with yellow tip, legs yellowish. The specimen corresponds to the de
entirelly yellow. It is Tricimba cincta Meigen, scription and is designated here as lectotype. -
1830. The specimen is not a type. - Current in Current interpretation: a good species Oscinella
terpretation: Tricimba lineella (Fallen, 1820). The nitidissima (Meigen, 1838).
lectotype is in Riksmuseum in Stockholm (An
dersson, 1963). Drawer60
Chlorops sulcicollis Meigen, 1830: 387. No.
2558 (2757 40). One female, glued, cheeks Chlorops longula Meigen, 1838: 389. No.
strongly produced, yellowish, antenna! fovea 2563 (2762 40). One specimen, glued, tip of ab
entirely separated by facial keel. Antennae dark. domen broken. Meigen 's description: "Kopf,
The specimen corresponds to the description and Fiihler und Stime rothgelb". The examined spec
is designated here as lectotype. - Current inter imen has antennae, including basal segments,
pretation: a good species Polyodaspis su/cicollis black, palpi with darkened tip, not enlarged. Ocel
(Meigen, 1830). lar triangle black with yellow sides. Stripes on
Chlorops vagans Meigen, 1838: 389. No. scutum black dusted, narrow; middle stripe short;
2559 (2758 40). Only "vagans", without generic outer lateral stripe very narrow, as stroke. Abdo
name, on the label. No specimen, only a piece of men brownish dorsally and ventrally, like in fe
paper on pin. Becker (1902) wrote "leere male of C. varsoviensis Becker. Pleura darkened
Nadel". - Current interpretation: a good species and with shining blackish marks. Anepisternum
Dicraeus vagans (Meigen, 1838). The only Eu with whitish setulae. Basitarsus of fore leg long.
ropean species with partly yellow thorax. Collin The specimen does not correspond to the descrip
(l 946) published a figure of the male genitalia, tion, it is a female of C. varsoviensis Becker,
and the species is easily distinguished. 1910. Becker (1902) only mentioned that the
Chlorops glaberrima Meigen, 1838: 389. No. specimen in NHMP is a female. - Current inter
2560 (2759 40). Only "glaberrima", without ge pretation: Chlorops longula Meigen, 1838, no
neric name, on the label. There is also a label men dubium.
"Museum Paris coll. Meigen". One female, glued, Chlorops hyalipennis Meigen, 1838: 389. No.
without head and part of legs (left fore leg and 2564 (2763 40). No specimen, only a piece of
both hind legs absent, except coxae). Body with paper on pin. Becker ( 1902) mentioned one male,
greenish lustre, preserved legs entirely yellow. Oscinis sp. - Current interpretation: Chlorops
The specimen corresponds to the description. It hyalipennis Meigen, 1838, nomen dubium.
is a female ofR hopalopterum anthracinum (Mei Chlorops nigrita Meigen, 1838: 391. No.
gen, 1830). Becker (1902) named the female in 2565 (2764 40). One female, in rather good con
Paris as Oscinis s. str. The specimen is here de dition, only right flagellomere broken. Legs en
signated as lectotype. - Current interpretation: tirely black, only tarsi brownish. Cheek as wide
junior synonym of Rhopa/opterum anthracinum as first flagellomere. It is Oscinella nige"ima
(Meigen, 1830); synonymy confirmed here. (Macquart, 1835). The species was considered a
Chlorops fascipes Meigen, 1838: 388. No. junior synonym of 0. frit Linnaeus (Duda, 1933)
2561 (2760 40). Only "fascipes", without gener or a good species (Collin, 1946; Nartshuk,
ic name, on the label. There is an additional la 1984). - Current interpretation: Chlorops nigri
bel "Paris Museum coll. Meigen". One male, tus Meigen, 1838, syn. n. is a junior synonym of
glued, in rather good condition, only arista and Oscinella nigerrima (Macquart, 1835).
left flagellomere broken. Legs mostly yellow with Chlorops polita. No. 2566 (2765 40). Label:
middle tibia dark. Becker (1902) mentioned one ''polita Mg. Bayern". Meigen (1838: 394) listed
female and identified it as Oscinis s. str. The spec Chlorops polita Macquart. Two specimens, glued,
imen is Rhopalopterum fasciolum (Meigen, and one piece of paper without fly. Both speci
1830) and is designated here as lectotype. -Cur- mens arePsilopa sp. (Ephydridae). Cogan (1984:
ZOOSYST. ROSSICA Vol. 15 • E.P. Nartshuk: Meigen s Chloropidae in Paris 179
141) listedPsilopapolita (Macquart, 1835: 524, Notes on M variegata sensu Fedoseeva (1960,
as Hydrellia), but Macquart described also Os 2003), Ismay (1981} and Nartshuk (1992) are
cinis polita Macquart, 1835: 603. The last spe given at the end of the paper.
cies is considered nomen dubium (Nartshuk, Meromyza saltatrix. No. 2569 (2769 40). Two
1984). specimens. There are marks males on the labels,
Chlorops exornata, no number on collection but the specimens are females of Meromyza tri
label (2766 40). No description or mention of angulina Fedoseeva with narrow hind femora,
species with this name is found by me in Meigen black setulae on cheeks and black sides of ocel
(1830, 1838) and it is not listed within Acalyp lar triangle. Both specimens are not types. - Cur
tratae in both catalogues of Palaearctic Diptera rent interpretation: a good species Meromyza sal
(Becker et al., 1905; Soos & Papp, 1984). One tatrix (Linnaeus, 1761).
male in good condition, is Chlorops speciosus Meromyza laeta Meigen, 1838: 395. No. 2570
Meigen. The name Chlorops exornata is a no (2770 40). One female, rather large, with black
men nudum (name in collection). apices of palpi, red stripes on scutum and strongly
Meromyza pratorum Meigen, 1830: 165. No. swollen hind femora. The specimen corresponds
2567 (2767 40). Two females, one of them with to Meromyza femorata Macquart, 1835 sensu
genitalia in a tube on the same pin and the label Fedoseeva(l960, 2003) andlsmay(1981). Beck
"Meromyza pratorum"; another female with the er (1902) mentioned that the specimen has black
label ''pratorum", without generic name. Both apices of palpi and considered the species to be
females in good condition and correspond to the identical with M. variegata Meigen. Meigen
description. The female with the genitalia in a mentioned red scutal stripes and gave no note on
tube is designated here as lectotype, the other the thickness of hind femora and colour ofpalpi.
female is a paralectotype. - Current interpreta There is no illustration in Meigen's "Dipteren
tion: a good species Meromyza pratorum Mei Farbtafeln" (Morge, 1976). Based on the red
gen, 1830. scutal stripes, I consider this specimen as corre
Meromyza variegata Meigen, 1830: 165. No. sponding to the description and designate it here
2568 (2768 40). One male and two females. Male as lectotype. Black apices ofpalpi, red stripes on
with genitalia in a tube on the same pin, exam scutum, and especially thick hind femora are char
ined by Ismay(1981). Labels of pins with a male acteristic of M. femorata Macquart. - Current
and a female only with specific name without interpretation: Meromyza laeta Meigen, 1838,
generic name, another female without label at all. syn. n. is a junior synonym of Meromyza femo
Both females are M femorata Macquart sensu rata Macquart, 1835.
Fedoseeva (1960, 2003) and Ismay(1981): palpi Meromyza laeta sensu Fedoseeva (1960, 2003 ),
black in apical half, scutal stripes red and no mark Ismay (1981) and Nartshuk (1992) is described
on scutellurn (possible to see only in a female), below as Meromyza meigeni sp. n. (description
hind femora strongly thickened. The male is an at the end of the paper).
other species. In the male, palpi narrowly black Meromyza nigriventris. No. 2571 (2771 40).
ish at apex, scutal stripes reddish, mark on scutel One female with wide black stripes on scutum,
lum not seen (the pin goes through the scutel black abdomen, black palpi, corresponds to Mero
lum), femora moderately swollen, 2.5 times as myza nigriventris Macquart, 1835. -Current in
thick as tibia. Surstyli rather long and directed terpretation: a good species Meromyza nigriven
laterally. Fore process of postgonites narrower tris Macquart, 1835.
than in M. femorata and aedeagus with bifurcat
ed base (Figs 5-9). In M femorata, the aedeagus Notes on Meromyza with description of
is with arrow-shaped base. I know only one spe a new species
cies of Meromyza with bifurcate base of aedea
gus, M. lidiae Nartshuk described from Finland, Meigen and other early authors did not include
Russia (Leningrad and Moscow provinces) and drawings or characteristics of the male genitalia
Bulgaria (Nartshuk, 1992). Other characters of in descriptions of new species of Meromyza.
Meigen's specimen also correspond to M lidiae Now, correct identification of most species is
(Figs 11-16). M variegata on Meigen 's drawing impossible without examination oft he male gen
(Morge, 1976: Tafel 162, 13) has reddish scutal italia. Fedoseeva (1960) published drawings of
stripes and no mark on scutellum; unfortunately, the male genitalia of many European species be
the palpi are not figured and their colour not fore the type specimens were examined, based
mentioned in Meigen's description. The male is only on other characters, mainly on colour. There
designated here as lectotype. - Current interpre fore, controversial opinions exist on identity of
tation: Meromyza lidiae Nartshuk, 1992, syn. n. some species and food-plants of their larvae, es
is a junior synonym of M variegata Meigen, pecially for M. femorata and M variegata. Fe
1830. doseeva (1960, 2003) named the grass Daclylis
180 E.P. Nartshuk: Meigen s Chloropidae in Paris • ZOOSYST. ROSSICA Vol. 15
I ,~
~ ~
I
f8 19 I 110 14
I I
I
(
15
16
Figs S-16. Meromyza variegata Meigen, male genitalia (5-10, lectotype, orig.; 11-16, from Nartshuk, 1992, as M.
lidiae Nartshuk). S, 11, epandrium; 6, sursylus, lateral view; 7, 13, postgonites, lateral view; 8-10, 14-16, aedeagus in
different positions; 12, epandrium, lateral view. Scale line: 0.1 mm.
ZOOSYST. ROSSICA Vol. 15 • E.P Nartshuk: Meigen s Chloropidae in Paris 181
glomerata as the food-plant of M. femorata and our of scutum, colour of abdomen, thickness of
Phleum pratense, as that of M. variegata. How hind femora. However, there are some differenc
ever, the European authors listed Daclylis glom es: larger size of body (3.0-3.5 mm), mark on
erata as the food-plant of M. variegata. Exami scutellum in the species described by Fedosee
nation of the types of two Meigen's species re va. In the description by Fedoseeva (1960), the
sults in description of a new European species palpi are described as narrow, in keys of Fedo
and change of interpretation of another species. seeva (2003) and Beschovski (1985) palpi are
characterised as swollen. I have seen only a fe
Meromyza meigeni sp. n. male determined by Fedoseeva as M. laeta Mei
(Figs 17-25) gen. Only postgonites of the male genitalia are
illustrated in Fedoseeva's paper, they correspond
= Meromyza laeta sensu Ismay {1981) and Nartshuk well to drawings by Ismay ( 1981) and Nartshuk
(1992). (1992), but other structures of the male genitalia
of her specimens are unknown. M. laeta sensu
Holotype: o', Slovenia, Postoina, 13.VII-l.Vill.1958, Fedoseeva occurs in Russia north to Moscow and
edge of mixed forest, R.L. Coe. east to Kemerovo, in Ukraine, and Kazakhstan
Paratypes. Slovenia: 2 o', same data as in holotype;
(Fedoseeva, 1960). Fedoseeva (1960) noted that
Italy: a series, Zuel, near Cortina, 13. Vffi.1969, V.F. Eas
top (determined as Meromyza laeta by J. Hubicka). this species occurs in dry sandy places in Mos
The holotype and both paratypes from Slove cow Province, host-plants are Agrostis vulgaris
nia are deposited at Zoological Institute, Russian With. and Festuca rubra L. Beschovski (1985)
Academy of Sciences, St.Petersburg; they were recorded the species from Bulgaria, Albania,
determined by E.P. Nartshuk as Meromyza laeta Macedonia and Bosnia.
and so listed by Coe (1968). The paratypes from
Italy are kept at the Natural History Museum, Meromyza athletica Fedoseeva, 1974
London; they are included in the type series on (Figs 27-40)
the basis of drawings of the male genitalia by
Ismay (1981 ), who listed these specimens as M. = Meromyza variegata sensu Ismay (J 981), Beschovski
(1985) and Nartshuk (1992).
/aeta Meigen.
Description. Body yellow. Frons yellow, slight Description. Body yellow. Frons yellow, slight
ly longer than wide, covered with short black ly longer than wide, covered with short black and
setulae; head setae black. Setae vte the longest, 3 white setulae; setae black. Setae vte the longest,
times as long as vti and twice as long as poc. vti and poc very small. Ocellar triangle yellow,
Ocellar triangle yellow, finely rugose, extending rugose in apical part, extending nearly to 2/3-3/4
to nearly 2/3 of frons. Parafacialia half as wide of frons. Parafacialia narrower than genae. Ge
as first flagellomere. Genae subequal in width to nae equal in width to first flagellomere. First flag
first flagellomere. First flagellomere slightly long ellomere slightly longer than wide, yellow, slight
er than deep, yellow, slightly darkened above. ly darkened above. Arista dark. Palpi narrow,
Arista dark. Palpi narrow, yellow, narrowly black yellow. Scutum longer than wide. Central scutal
at apex. Scutum longer than wide. Scutal stripes stripes red, short, sharply bordered, not extend
mainly red; outer lateral stripes brownish. Bor ing to scutellum by distance equal to scutellum.
ders of scutal stripes diffuse, not sharp, especial Inner part of inner lateral stripes reddish, remain
ly hind end of central stripe. Pleura yellow with der part and outer lateral stripes black dusted, or
reddish mark on katepistemum and katepimeron. both lateral stripes black. Pleura yellow with red
Hind femora moderately swollen, no more than dish mark on katepistemum and katepimeron.
2.5 times as thick as tibia. Abdomen with central Hind femora strongly swollen, 3.5-4.0 times as
line of black spots, and tergites brownish in fore thick as tibia. Abdomen with three black spots
2/3 or sometimes without spots. Epandrium on tergites. Epandrium rather large, with short
small, with short setae; surstyli short. Postgonites setae. Surstyli short, squared or slightly rounded
black. Anterior process of postgonites rather nar apically. Postgonites black. Anterior process of
row and rounded. Aedeagus narrow, with arrow postgonites moderately massive, with short acute
shaped base. Body length 2.0-2.5 mm. tip and convex upper margin near tip. Aedeagus
Distribution. A species of South European dis with median constriction and two projections in
tribution: Slovenia, Italy. Ismay (1981) did not lateral view. Body length 3.5-4.0 mm.
found the species in England. Distribution. Southern part of European Rus
Note. Meromyza laeta sensu Fedoseeva ( 1960, sia north to Kursk, Ukraine, Georgia, Bulgaria,
2003) and Beschovski (1985) is probably the Great Britain. Host-plant: Phleum pratense (Fe
same species, it has the same structure of post doseeva, 2003).
gonites (Fig. 26) and some other characters, as Note. Fedoseeva (197 4) described this species
the shape of the first flagellomere, size and col- from the south of the European part of the former
182 E.P. Nartshuk: Meigen '.!-Chloropidae in Paris • ZOOSYST. ROSSICA Vol. 15
19
I
1
20
~~!'0
I
22 24 . /
\'':: 1 I
0
I I
1 \
..._....._)
' ' .. --~~ .... --) 25
I
23
Figs 17-26. Meromyza meigeni sp. n. (17-20, from Nartshuk, 1992, asM. laeta, paratype; 21-23, from Ismay, 1981, as
M. laeta; 26, M. laeta sensu Fedoseeva, 2003). 17, 21, epandrium, ventral view; 18, 22, epandrium, lateral view; 19,
23, 26, postgonites, lateral view; 20, 24, 25, aedeagus in different positions. Scale line: 0.1 mm.
USSR, she compared it with M variegata sensu and Nartshuk (1992) to be M. athletica Fedosee
Fedoseeva (1960), and the recorded differences va (1974). The difference in the shape ofpostgo
were insignificant. The main difference is in the nites in the drawing of M athletica by Fedosee
shape ofpostgonites (Figs 38, 39). I have seen a va (1974, 2003) is explained, in my opinion, by
paratype of M athletica and examined the male not strongly lateral position of this structure.
genitalia of another specimen determined as M.
athletica by Fedoseeva. The shape of postgonites Acknowledgements
is the same as figured by Fedoseeva ( 1960, 2003 ),
Ismay (1981), Beschovski (1985) and Nartshuk I am greatly thankful to Dr. T. Pape (Zoological Muse
(1992) for M variegata, but not as figured by um in Copenhagen, Denmark) for the opportunity to visit
MHNP in course of the Programme "Fauna Europaea'',
Fedoseeva (1974, 2003) for M. athletica. Fedo
Dr. D. Matile-Ferrero, Dr. M. Bailak and J. Charbonel
seeva never figured other parts of the male geni for kind help during my visit to MHNP in Paris in 2002.
talia of her M. variegata, but gave a drawing of The study was supported by the Russian Foundation for
the aedeagus of M. athletica (Fig. 40), in which Basic Research (grant 05-04-48258), grant of the Presi
is well visible the projected base of aedeagus, as dent of the Russian Federation for support of the leading
scientific schools (NSh 4686. 2006.4), Programmes of
figured byismay(l981) and Nartshuk(1992) for
the Ptesidium of the Russian Academy of Sciences "Sci
M. variegata. On these grounds, I consider M entific bases of conservation of biodiversity in Russia"
variegata sensu Ismay ( 1981 ), Beschovski ( 1985)