A provisional checklist and gazetteer of the spiders of Chios, Greece (Arachnida: Araneae)

2011·0.25 MB·English

by  Russell-Smith, A

Bull.Br.arachnol.Soc.(2011)15(5),133–167 133
A provisional checklist and gazetteer of the spiders particularlyTurkeyandCyprus,isevenpoorerthanthat
of Chios, Greece (Arachnida: Araneae) of Greece and it is likely that many species currently
considered endemic to Greece will be found to have
A. Russell-Smith1, R. Allison2, M. Askins3, wider distributions once they have been better studied.
W. Blumsom4, R. Snazell5 and C. Spilling6 The Greek list is smaller than those for the other two
11BailiffsCottage,Doddington,Sittingbourne,Kent,ME90JU major Mediterranean areas in Europe, 1197 from the
2WalnutBarn,GussageSt.Michael,Dorset,BH215HX Iberian peninsula (Melic & Morano, 2004) and 1411
369SavillCrescent,Wroughton,Swindon,Wiltshire,SN49JG from Italy (Stoch, 2003). Given the geographic position
430WhiteStreet,WestLavington,Devizes,Wiltshire,SN104LP
of Greece at the crossroads between Europe, Asia and
5FourWinds,10BonAccordRoad,Swanage,Dorset,BH192DS
themiddleEast,thecurrenttotalalmostcertainlyrepre-
6Culls,GarfieldLane,LangtonMatravers,Swanage,Dorset,
BH193HJ sentsafractionofthetruenumberofspeciestobefound.
Most studies of Greek spiders have focused on par-
ticulargeneraorfamiliesandhaveoftenformedpartof
Summary
wider studies of the Mediterranean spider fauna. There
A provisional checklist of the spiders of the eastern appear to have been few published attempts to enumer-
AegeanislandofChios(Greece)isprovided.Thechecklist atethefaunaofadefinedareawithinthecountrywhich
includesdetaileddataonlocationsanddatesforallspecies
can provide information on the relative abundance of
recorded, together with information on their habitats and
differentspeciesandtheirhabitatpreferences,otherthan
overall distribution range. In total, 39 spider families and
315specieshavebeenrecordedfromtheislandtodate.The a recent checklist for the island of Lesbos (Bosmans
most species-rich families were the Gnaphosidae (53 et.al.,2009).However,theonlinechecklistofthespider
species),Salticidae(40species),andTheridiidae(40species), fauna of Crete (Eikamp & Chatzaki, 2007) provides a
which together accounted for 42% of all spiders. The list
useful comparative list from that region of Greece. At
includes 21 species which were previously unknown from
thesuggestionofMikeTaylor,anentomologistwhohas
Greece,sixofwhicharealsonewtoEurope.Thechecklist
alsoincludesaminimumofafurther45taxawhichremain spent many years working on the insects of Chios, a
to be fully identified but are clearly distinct. The family small group from the British Arachnological Society
Gnaphosidaeinparticularincludes17taxawhichcurrently decided to carry out an arachnological survey of the
cannot be assigned to known species. The composition of
island.Thispaperpresentsresultsfromthesurveywork
thespiderfaunaiscomparedwiththatfromotherareasof
Greece and the reasons for similarities and differences are which it is hoped will continue in future. Inevitably,
discussed. given the patchy state of knowledge of spiders of this
region, many species could not be fully identified at
present. We have decided, however, to include these in
Introduction
our list with the aim of providing a more complete
Compared with much of north-west and central account of the fauna and in the hope that relevant
Europe, the spider fauna of Greece has been poorly specialists will study them in the future.
studied. Early work by Caporiacco, Drensky, Fage,
C. L. Koch, Kulczyn´ski, Pavesi and Simon contributed Material and methods
to a list of 373 species produced by Bristowe (1935).
Study area
Subsequently, work on the spiders of the province of
AttikibyHadjissarantos(1940)producedacatalogueof Chios is an island in the north-east Aegean Sea lying
316species,ofwhichabouthalfwerenewtothatregion. closetotheTurkishcoast,southofLesbosandnorthof
More recently, the important work of Metzner (1999) Samos. The island is the fifth largest in Greece, with a
provided keys, descriptions and excellent figures for all total area of 904km2, a varied geology and a wide
the known species of Salticidae from Greece. Another topographic range, capped by Mt. Pelineon (1297m) in
landmark was the series of papers by Chatzaki and the north of the island. Its location close to the Turkish
colleagues which describe and figure all the known mainland, opposite the Izmir Peninsula, means that its
species of Gnaphosidae from the island of Crete fauna and flora consists of a mixture of European and
(Chatzaki et al., 2002a, b, 2003). While there have been Asian/Middle Eastern elements. Agricultural activity is
manymorerecentcontributionstoourknowledgeofthe non-intensiveandpatchyinnatureandthishascontrib-
spider fauna of Greece, these are widely scattered uted to the retention of much of the traditional land-
throughout the literature. They include significant con- scape and to the preservation and protection of native
tributions from Deeleman-Reinhold (Dysderidae), plantsandanimals.Furthermore,Chioshasnotsuffered
Knoflach(Theridiidae),Thaler(Lycosidae,Linyphiidae) from the excesses of environmentally damaging and
and Wunderlich (Linyphiidae and other families). Revi- inappropriate developments seen in many other Medi-
sionsofparticulargenerafromthewiderMediterranean terranean areas. A majority of the inhabitants are con-
region also include many species from Greece and these centratedintheprincipaltownontheeastcoastandthe
are listed in the catalogue mentioned below. humanpopulationdensityoftheremainderoftheisland
The most recent and complete listing of spiders from is exceptionally low.
Greece is the catalogue of Bosmans & Chatzaki (2005). Chioshasawiderangeofhabitats,includinglowland
This includes a total of 856 spider species, of which 213 phrygana (the Greek name for garrigue, Mediterranean
arecurrentlylistedasendemictothecountry.However, vegetation dominated by spiny under-shrubs), montane
our knowledge of the fauna of adjacent countries, phrygana, flower-rich grassland, pine forest, relict oak
134 ChecklistofspidersofChios
Locality Latitude Longitude Locality Latitude Longitude
AgiasmataBeach 38(35#25$N 25(36#08$E Kipouries 38(29#26$N 25(59#25$E
AghiosGalios 38(33#26$N 25(51#23$E Limenas 38(18#N 25(57#E
Anavatos 38(24#06$N 26(01#17$E Managros 38(28#N 25(57#E
Anavatosvalley 38(23#20$N 26(01#25$E ManagrosBeach 38(28#28$N 25(56#12$E
Valleynr.Armolia 38(14#57$N 26(21#13$E MarmaroMarsh 38(32#N 26(07#E
ElindaBay 38(23#03$N 25(59#23$E Mesta 38(15#23$N 25(55#13$E
Emborios 38(11#20$N 26(01#50$E Mt.Pelineon 38(34#N 26(01#E
Fytastream 38(31#41$N 26(00#27$E Nagos 38(33#31$N 26(04#55$E
Giosonas 38(33#34$N 24(04#28$E NeamoniMonastery 38(22#29$N 26(03#35$E
Gridia 38(12#56$N 26(06#08$E Ochrea 38(28#29$N 25(56#20$E
NearGridia 38(12#57$N 26(06#17$E Padoukios 38(27#43$N 26(08#12$E
Halandra 38(32#58$N 25(55#38$E Pagida 38(15#17$N 26(05#24$E
Issidoros 38(27#27$N 26(00#37$E PaleaPotamia 38(42#52$N 25(56#15$E
Kalimmasia 38(17#46$N 26(06#29$E Parparia 38(31#11$N 25(53#14$E
KampiaGorge 38(34#N 25(58#E PowerStation 38(19#N 26(09#34$E
KampiaBeach 38(35#37$N 25(58#48$E SalagonasBay 38(12#50$N 25(54#42$E
4kmN.ofKampia 38(35#10$N 26(00#40$E Salagonas 38(12#05$N 25(54#47$E
NearKataraktis 38(16#07$N 26(05#37$E Spartouna 38(33#39$N 25(58#57$E
HillE.ofKatavasi 38(28#00$N 25(59#00$E Viki 38(34#46$N 26(00#53$E
KatoFana 38(12#45$N 25(55#45$E Ochrea,nr.Volissos 38(28#18$N 25(56#24$E
KatoFanaBeach 38(12#23$N 25(55#43$E VoulamandisHouse 38(19#13$N 26(08#11$E
KeramiaClayPit 38(18#00$N 26(08#20$E Vrondados 38(24#57$N 27(07#53$E
Table1: Latitudeandlongitudecoordinatesforallsitesfromwhichspiderswererecorded.
forest, freshwater marsh, rocky stream valleys and many specialists who have given so freely of their time
gorges. Coastal habitats include both stony and sandy and expertise and who are listed in the acknowledge-
beaches, reed beds and rocky cliffs. There are a number mentsattheendofthispaper.Initially,eachparticipant
of caves on the island but these were not visited during in the survey sorted and identified their own specimens
the survey. as far as was possible using the available literature.
Specimens that were either unidentified or for which
Survey periods and methods somedoubtexistedwerethensubmittedtospecialistsfor
their scrutiny. Although this process helped to ensure
The main periods of survey were during two visits to that a large proportion of specimens were correctly
the island from 8–16 May 2006 and from 21 April–2 identifiedtospecies,therewerestillmanytaxawhichwe
May 2008. In addition, spiders collected in the period are currently unable to name. These have been included
1997–1999 by C. Felton, M. J. Taylor and G. R. in the list below, both for the sake of completeness and
Underwood from the collections of the Liverpool toindicatewherefurthersystematicresearchisrequired.
Museumwerealsoincluded.Themajorityofthesewere All material collected by G. R. Underwood, C. Felton
collectedbyhandbutthereweresomefrompitfalltraps andM.J.TaylorisdepositedintheLiverpoolMuseum,
collected by M. J. Taylor in spring 1999, spring and whiletheremainderisheldinthepersonalcollectionsof
autumn 2008 and spring and autumn 2009. the following individuals: R.A.=R. Allison, M.A.=M.
During the survey the majority of sampling was by Askins, W.G.B.=W. Blumsom, A.R.-S.=A. Russell-
handsearchingbutsweep-nettingofgrasslandandbeat- Smith, R.S.=R. Snazell, C.S.=C. Spilling, M.J.T.=
ing of shrubs and the lower branches of trees as well as M. J. Taylor.
sifting leaf-litter were also used. Sampling of specific
micro-habitats included under stones in rock phrygana
andgrasslands,onstonewalls,onandunderthebarkof Localities
trees (particularly old olive trees) and in the heads of
Latitude and longitude coordinates for all sites men-
reeds (Phragmites australis). On a number of occasions,
tioned in the text are given in Table 1.
pitfall traps were used to sample ground-active species.
These included those mentioned above, operated in
May and June 1999 by M. J. Taylor at three sites, in Checklist
Marmaro Marsh in April 2008, a rocky valley near
Armolia by M. J. Taylor in autumn 2008 and traps The following checklist provides as full detail as
placedinfoursitesinbothspringandautumn2009,also possible of the locations from which all specimens have
by M. J. Taylor. All specimens were preserved in 70% been taken, and summary details of their habitat and
ethanol for subsequent examination and identification. micro-habitats. We have included material collected by
C. Felton and G. R. Underwood between 1997 and
1999, as well as that collected by Mike Taylor over the
Identification
past 10 years, all now in the Liverpool Museum. It also
Identification has relied on the publications listed in includes earlier published records, some of which have
Bosmans & Chatzaki (2005) and, particularly, on the not been rediscovered during our survey and for many
A.Russell-Smithetal. 135
of which no precise locality data are available. These Family Anyphaenidae
earlier records are placed in square brackets to dis-
tinguishthemfromourownandthoseofcollectorsfrom
the Liverpool Museum. Anyphaena sabina L. Koch, 1866
Records:Viki,28April1997,3\(C.Felton).Mt.Pelineon(600m),
9May2006,1\(A.R.-S.),2\(W.G.B.);14May2006,1\(M.A.),1\
Family Agelenidae (R.A.).KampiaGorge,30April2008,1\(A.R.-S.).
Distribution: The commonest species of this genus in
Greece.DistributedthroughoutsouthernEuropeandas
Maimuna vestita (C. L. Koch, 1841)
far east as Azerbaijan. Most frequently taken on Chios
Records: [Without locality or habitat data (Strand, 1917)]. Kato
Fanabeach,27April1997,2\(C.Felton).Rockyvalleynr.Armolia, on foliage of shrubs and trees.
8May2006,1\(A.R.-S.).MarmaroMarsh,9May2006,1\(R.S.),1\
(R.A.). Fyta stream, 10 May 2006, 2\ (A.R.-S.). Kampia Gorge, 10
May2006,3\(A.R.-S.),3\(R.A.).ElindaBay,3\(W.G.B.).Ochrea, Family Araneidae
near Volissos, 12 May 2006, 1\ (M.A.), 1\ (R.S.), 5\ (R.A.). Mt.
Pelineon,14May2006,2\(R.A.).ManagrosBeach,22April2008,1\
(R.S.), 1\ (R.A.). Salagonas Bay, 23 April 2008, 2\ (A.R.-S.), 2\
Agalenatea redii (Scopoli, 1763)
(M.A.), 2\ (R.A.). Mt. Pelineon, 24 April 2008, 2\ (R.A.). Ochrea,
nearVolissos,25April2008,2\(M.A.),5\(R.A.).MarmaroMarsh, Records:4kmN.ofKampia,28April1997,1\(C.Felton).Near
25April2008,2\(A.R.-S.),3\(R.S.),1\(R.A.).KeramiaClayPit,26 Gridia,1\(G.R.Underwood).KatoFana,8May2006,1\(M.A.).
April2008,3\(R.A.).VoulamandisHouse,Kampos,27April2008, Marmaro Marsh, 9 May 2006, 1\ (R.A.). Ochrea, near Volissos, 1\
1\(A.R.-S.),1\(R.S.),1\(R.A.).Onhill1.5kmE.ofKatavasi,29 (A.R.-S.).KatoFana,28April2008,1\(R.A.).
April 2008, 1\ (A.R.-S.), 2\ (R.S.), 1\ (M.A.), 2\ (R.A.). Kampia Distribution: A widespread species in Greece but
Gorge,30April2008,1\(R.S.),1\(M.A.).Mt.Pelineon,1May2008,
apparentlynotparticularlycommononChios.Itoccurs
1\(R.S.),1_1\(A.R.-S.),1\(M.A.).Agiasmata,22April2008,3\
(C.S.). Mt. Pelineon, pitfall traps, 26 July–7 September 2009, 1\ in the Palaearctic region from western Europe to NW
(M.J.T.). China. All specimens were swept from the field layer of
herb-rich grasslands.
Distribution: A widespread and common species
throughout the island, as it is throughout the eastern
Mediterranean. Females are nearly always collected in
Araneus angulatus Clerck, 1757
small webs beneath stones, but it has also been found
Records:KatoFana,nearbeach,8May2006,1_(M.J.T,reared
under the bark of a pine tree on Chios. byW.G.B.)
Distribution: A Palaearctic species widespread in
Tegenaria angustipalpis Levy, 1996 Greece but it may be autumn-maturing, so possibly
Records: Kipouries, from pitfall trap in stream valley, 9–24 May under-recorded on Chios.
1999, 1_ (M.J.T.). Marmaro Marsh, 9 May 2006, 1\ (R.A.).
VoulamandisHouse,Kampos,inbathroom,22April2008,1_(C.S.).
ManagrosBeach,22April2008,1_1\(R.S.).
Araneus circe (Audouin, 1826)
Distribution:OriginallydescribedbyLevy(1996)from Records:[Withoutlocalityorhabitatdata,Werner,1935].Kampia
Israel, these are the first published records from Europe. Gorge,30April2008,1_(R.S.).
Specimens were collected in shady habitats and in a Distribution:ThisspeciesiswidelyrecordedinGreece
house in Kampos. and is distributed throughout the southern Palaearctic
region. On Chios it has been taken from shrubs in a
shady gorge.
Tegenaria parietina (Fourcroy, 1785)
Records:NagosSpring,Nagos,19September2006,1_(M.J.T.).
Distribution:AwidespreadspeciesinGreecebutsofar Araniella inconspicua (Simon, 1874)
only a single record from Chios. Found throughout Records: Mt. Pelineon (600m), 9 May 2006, 1\ (A.R.-S.). Beaten
Europe and as far east as central Asia. fromshrub,KampiaGorge,1\(M.A.).
Distribution: There are only two previous records
of this species from Greece, from Sterea Elada and
Family Amaurobiidae
Macedonia respectively. It is found throughout the
Palaearctic region. The specimens from Chios were
Amaurobius deelemanae Thaler & Knoflach, 1995 beaten from trees.
Records:Fytastream,10May2006,1_(R.S.).MarmaroMarsh,25
April 2008, 1_ 1\ (R.S.). Mt. Pelineon (600m), pitfall traps, 12–29
April2009,1_.Samesite,10–21October2009,1_(M.J.T.). Argiope lobata (Pallas, 1772)
Distribution:ThisspeciesisendemictoGreeceandhas Records:ElindaBay,20June2006,1_(M.J.T.).SalagonasBay,23
April2008,1imm.(R.A.).
previously been recorded from Crete, Rhodes and
Naxos. On Chios males were collected in montane Distribution: A common species in Greece, probably
phrygana and moist grassland, but females that cannot under-recorded on Chios because it is usually autumn-
becertainlyassignedtoaspeciesinisolationfrommales maturing. Distributed from southern Europe to central
were found in a wider range of habitats. Asia.
136 ChecklistofspidersofChios
Cyclosa sierrae Simon, 1870 3_ 3\ (A.R.-S.), 2_ (R.A.). Elinda Bay, beating, 11 May 2006, 1\
Records: Rocky valley nr. Armolia, 8 May 2006, 1\ (A.R.-S.). (M.A.). Managros Beach, 11 May 2006, 1\ (M.A.). Ochrea, near
Mt. Pelineon, 9 May 2006, 1\ (W.G.B.). Kampia Gorge, 10 May Volissos, 12 May 2006, 1\ (M.A.). Anavatos valley, 2km S. of
2006, 3_ 2\ (A.R.-S.), 1\ (R.A.). Elinda Bay, 11 May 2006, 2_ 2\ Anavatos,12May2006,1\(W.G.B.).Mt.Pelineon(600m),14May
(R.S.). Ochrea, near Volissos, 12 May 2006, 1_ (A.R.-S.), 1_ 4\ 2006,3_(M.A.).Gridia,16May2006,1\(M.A.).SalagonasBay,23
(R.A.). Anavatos valley, 2km S. of Anavatos, 12 May 2006, 4_ April2008,1\(C.S.),1_(R.S.),1\(R.A.).Ochrea,nearVolissos,25
(A.R.-S.), 1\ (R.A.). Salagonas Bay, 23 April 2008, 1_ (M.A.). April2008,1\(C.S.).Onhill,1.5kmE.ofKatavasi,2May2008,1_
Marmaro Marsh, 25 April 2008, 1\ (R.S.). Voulamandis House, 2\ (C.S.). Agiasmata, near derelict water mill, 22 April 2008, 1\
Kampos, 27 April 2008, 1_ (R.S.). Kato Fana, 27 April 2008, 1\ (A.R.-S.).Rockyvalleynr.Armolia,28April2008,1_(A.R.-S.),1\
(A.R.-S.).KampiaGorge,30April2008,1_2\(C.S.),3_3\(R.S.), (R.A.). Kampia Gorge, 30 April 2008, 1\ (M.A.), 1_ 4\ (R.A.).
2_1\(M.A.),1\(R.A.).Mt.Pelineon,nr.rockybluff,1May2008,1_ Pagida,pitfalltraps,18May2009,1\(M.J.T.).
(R.S.).
Distribution: Widespread and abundant throughout
Distribution: A widespread species throughout Chios, Chios,GreeceandacrossthePalaearcticregionasfaras
as in Greece as a whole. It is distributed from southern NWChina.Nearlyalwayssweptfromflower-richgrass-
EuropeeastwardstoGeorgia.Allspecimenswereswept landonChiosbutalsooccasionallyonshrubsandtrees.
from the field layer of herb-rich grasslands.
Neoscona adianta (Walckenaer, 1802)
Gibbaranea omoeda (Thorell, 1870)
Records:NearGridia,3April1998,1\(G.R.Underwood).Near Records:KatoFana,8May2006,1_(R.A.).MarmaroMarsh,in
Kataraktis,4April1998,1_1\(G.R.Underwood).Kalimmasia,29 long grass and reeds, 9 May 2006, 1_ (M.A.). Managros Beach, 11
April 1997, 1_ (C. Felton). Mt. Pelineon (600m), 9 May 2006, 1\ May2006,1_(R.S.).Ochrea,nearVolissos,12May2006,1_(R.S.),
(A.R.-S.). Fyta stream, 10 May 2006, 1\ (A.R.-S.). Kampia Gorge, 4_1\(R.A.).Anavatosvalley,2kmS.ofAnavatos,12May2006,1\
paths in gorge, 10 May 2006, 3_ 3\ (A.R.-S.). Elinda Bay, behind (W.G.B.). Salagonas Bay. 15 May 2006, 1_ (W.G.B.), 3\ (R.A.).
beach,beating,11May2006,1\(R.S.).Ochrea,nearVolissos,12May Gridia,16May2006,3_(W.G.B.).SalagonasBay,23April2008,1_
2006, 1\ (A.R.-S.). Anavatos valley, 2km S. of Anavatos, 12 May 1imm.(A.R.-S.).MarmaroMarsh,25April2008,1_(A.R.-S.).Kato
2006, 2\ (A.R.-S.). Kampia Gorge, 30 April 2008, 1_ 2\ (C.S.), 1_ Fana,27April2008,1_1imm.(C.S.).KampiaGorge,30April2008,
(M.A.),1\(R.A.). 1\ (R.A.). Gridia, track S. of village, 15 May 2009, 1_ (M.J.T.).
Pagida,pitfalltraps,18May2009,1_1\(M.J.T.).
Distribution: A widespread species on Chios and re-
Distribution: Widespread and abundant throughout
corded from several other areas of Greece. Recorded
Chios and across the Palaearctic region. On Chios,
fromthroughoutthePalaearcticregion.OnChiosfound
usually swept from grassland and particularly common
in the field layer and on foliage of bushes and trees.
in moist places.
Glyptogona sextuberculata (Keyserling, 1863)
Records:Agiasmata,nearderelictwatermill,22April2008,1imm.
Neoscona subfusca (C. L. Koch, 1837)
(A.R.-S.).Ochrea,nearVolissos,25April2008,2imm.(R.S.).
Records:MarmaroMarsh,9May2006,1_(R.A.).SalagonasBay,
Distribution: A very distinctive species, recognisable 15 May 2006, 1\ (W.G.B.), 1\ (R.A.). Keramia Clay Pit, 26 April
even as juveniles, which has been widely recorded in 2008, 1\ (M.A.). Gridia, track S. of village, pitfall traps, 23 April–5
May2009,1_(M.J.T.).
Greece but is apparently not common. It occurs
throughout the eastern Mediterranean, from Italy to Distribution: A species distributed from southern
Israel. EuropeandAfricatocentralAsia;widespreadinGreece
but apparently not common on Chios. Specimens were
Hypsosinga albovittata (Westring, 1851) collected in flower-rich grassland in an olive orchard, in
Records:NearKataraktis,4April1998,1_1\(G.R.Underwood). tall moist grassland in a clay pit and in dry phrygana.
Kalimmasia,29April1997,1_(C.Felton).ManagrosBeach,11May
2006, 2\ (R.S.). Elinda Bay, beating, 11 May 2006, 1_ (R.S.).
Anavatosvalley,2kmS.ofAnavatos,12May2006,1_2\(A.R.-S.),
Zilla diodia (Walckenaer, 1802)
1\ (R.A.). Ochrea, near Volissos, 12 May 2006, 1\ (R.S.). Rocky
Records: Mt. Pelineon (600m), in pine forest, 9 May 2006, 1_ 2\
valleynr.Armolia,28April2008,1_(C.S.).KampiaGorge,onpath
(A.R.-S.). Fyta stream, 10 May 2006, 2\ (A.R.-S.), 1\ (M.A.), 1\
down into gorge, 30 April 2008, 1\ (A.R.-S.). Rocky valley nr.
(R.A.).KampiaGorge,10May2006,2_1\(A.R.-S.).Mt.Pelineon,
Armolia,pitfalltraps,4–25April2009,1_(M.J.T.).
near rocky bluff, 24 April 2008, 1\ (M.A.), 1\ (R.A.). Same site, 30
Distribution: Widespread on Chios, as in Greece as a April 2008, 1\ (M.A.). Rocky valley nr. Armolia, 28 April 2008, 1\
whole, it is distributed throughout much of the Palae- (C.S.).KampiaGorge,30April2008,1\(R.A.).
arctic region. Generally swept from the field layer but Distribution: Fairly common on Chios and Greece
occasional individuals are found on the ground. generally, it occurs from W. Europe to Central Asia.
Normally swept from the field layer but one specimen
Larinioides cornutus (Clerck, 1757) taken on the ground.
Records:MarmaroMarsh,inlongmoistgrassland,25April2008,
1\(W.G.B.).
Zygiella atrica (C. L. Koch, 1845)
Distribution:ThisspecieshasaHolarcticdistribution.
Records:KampiaGorge,30April2008,1\(M.A.).
Distribution: Found throughout Europe and Euro-
Mangora acalypha (Walckenaer, 1802)
Records:MarmaroMarsh,9May2006,1\(W.G.B.),2_2\(R.A.). pean Russia, and widely distributed in Greece. It is
Fytastream,10May2006,1\(A.R.-S.).KampiaGorge,10May2006, apparently uncommon on Chios.
A.Russell-Smithetal. 137
Family Cithaeronidae Family Dictynidae
Cithaeron sp. Altella lucida (Simon, 1874)
Records:Rockyvalleynr.Armolia,28April2008,1imm.(R.S.). Records:Mt.Pelineon,openbluffbytracktorefuge,9May2006,1_
Distribution: This immature specimen may belong to (R.S.).
C. praedonius O. Pickard-Cambridge, 1872, previously Distribution: Recently recorded for the first time in
recorded from a single locality in Greece. On Chios, it Greece from Lesbos (Bosmans et al., 2009). Specimen
was found on rocky ground on a scrubby hillside. collected in rocky grassland on Chios. Distributed
throughout Europe except Scandinavia.
Family Clubionidae
Dictyna civica (Lucas, 1850)
Clubiona vegeta Simon, 1918
Records: Kato Fana, in flower-rich meadow, 8 May 2006, 1\
Records: Nr. Armolia, 25 April 1997, 1\ (C. Felton). Gridia, 29 (M.A.),1\(R.S.),1\(A.R.-S.).Rockyvalleynr.Armolia,8May2006,
April1997,3\(C.Felton).Kalimmasia,29April1997,2\(C.Felton). 1\(R.A.).MarmaroMarsh,9May2006,1_(A.R.-S.).Ochrea,near
Kato Fana, 8 May 2006, 1\ (A.R.-S.). Rocky valley nr. Armolia, 8 Volissos, 12 May 2006, 1_ (R.A.). Anavatos valley, 2km S. of
May2006,1\(M.A.).Mt.Pelineon(600m),9May2006,1\(A.R.-S.). Anavatos,12May2006,1_2\(A.R.-S.),2_(R.A.).Rockyvalleynr.
Ochrea,nearVolissos,12May2006,1\(M.A.).Anavatosvalley,2km Armolia,13May2006,1_(M.A.),1_(R.A.).Gridia,13May2006,1\
S. of Anavatos, 12 May 2006, 1\ (M.A.). Gridia, 21 April 2008, 1\ (M.A.). Same site, 16 May 2006, 3\ (M.A.), 4_ 2\ (R.A.). Voula-
(C.S.). Agiasmata, 22 April 2008, 1\ (A.R.-S.). Managros Beach, 22 mandisHouse,Kampos,16May2006,1_(R.A.).Gridia,trackN.of
April2008,1\(R.S.).SalagonasBay,23April2008,1\(A.R.-S.),1\ village,21April2009,1_(M.A.).Agiasmata,nearderelictwatermill,
(R.S.).Rockyvalleynr.Armolia,25April2008,1\(R.S.).Hill1.5km 22 April 2008, 1_ (A.R.-S.). Salagonas Bay, 23 April 2008, 2_ 1\
E. of Katavasi, 29 April 2008, 1\ (A.R.-S.). Elinda Beach, 29 April (M.A.),1_ (A.R.-S.), 1\ (C.S.). Kato Fana, 27 April 2008, 1_ 1\
2008, 1\ (A.R.-S.). Palea Potamia, 9 May 2008, 1\ (A.R.-S.). (A.R.-S.), 1_ (R.A.). Keramia Clay Pit, 26 April 2008, 1\ (C.S.).
MarmaroMarsh,25April2008,1\(A.R.-S.).NeamoniMonastery,2 Rocky valley nr. Armolia, 28 April 2008, 1_ 1\ (A.R.-S.). On hill
May2008,1\(R.S.).Mt.Pelineon,nr.rockybluff,pitfalltraps,7–14 1.5kmE.ofKatavasi,29April2008,1_(M.A.).Mt.Pelineon,near
May2009,1_(M.J.T.). rockybluff,1May2009,1_(M.A.).Gridia,trackS.ofvillage,15May
Distribution: Widespread and common throughout 2009,2_1\(M.J.T.).Pagida,18May2009,4_2\(M.J.T.).
the eastern Mediterranean and on Chios, it occurs from Distribution:AwidespreadspeciesinGreece,foundin
W. Europe to central Asia. Found both under stones southern Europe and North Africa, as well as N.
and in the field layer, mainly in grasslands. America. All specimens from Chios were swept in herb-
rich grassland.
Family Corinnidae
Lathys arabs Simon, 1910
Arabelia pheidoleicomes Bosselaers, 2009
Records:KeramiaClayPit,26April2008,1\(R.S.),1\(R.A.).
Records: Managros Beach, under stone in dry meadow, 11 May
2006, 1\ (R.S.). Managros Beach, 22 April 2008, 1\ (C.S.). On hill, Distribution: Described from Algeria and earlier con-
1.5kmE.ofKatavasi,29April2008,1\(M.A.). sidered a synonym of L. stigmatisata Menge, 1869, this
Distribution: This species has recently been described specieswasrecentlycollectedonLesbos(Bosmansetal.,
from the neighbouring island of Lesbos (Bosselaers, 2009). It was found in long grass in a moist area on
2009). On Chios it was collected in dry grassland and Chios.
degraded phrygana.
Lathys humilis (Blackwall, 1855)
Phrurolithus sp. Records: Rocky valley nr. Armolia, 8 May 2006, 1\ (M.A.). Mt.
Records: Marmaro Marsh, 9 May 2006, 1\ (R.S.). Ochrea, near Pelineon(600m),9May2006,1\(A.R.-S.),1\(M.A.).Fytastream,
Volissos,12May2006,1_(R.A.).MarmaroMarsh,25April2008,1\ 10 May 2006, 1\ (M.A.). Elinda Bay, 11 May 2006, 1\ (M.A.).
(A.R.-S.). Anavatos valley, 2km S. of Anavatos, 12 May 2006, 7\ (M.A.), 1\
(R.A.).Mt.Pelineon(600m),14May2006,4\(R.A.).Ochrea,near
Distribution:Collectedamongtherootsoflong,moist Volissos,25April2008,1\(M.A.).KeramiaClayPit,26April2008,
grassland and in a shady meadow beneath large oak. 1\ (M.A.). Kampia Gorge, near mouth of gorge, 30 April 2008, 1\
(A.R.-S.).
Distribution:ThespeciesisknownfromtheW.Medi-
Family Ctenizidae
terranean but has only recently been reported from
Lesbos (Bosmans et al., 2009) in Greece; otherwise
Cyrtocarenum cunicularium (Olivier, 1811)
recorded throughout the Palaearctic. Specimens were
Records: [Without locality or habitat data (Strand, 1917)].
Vrondados, 22 September 1996, 1_ (M.J.T.). Rocky valley nr. beaten from the foliage of shrubs and trees.
Armolia, pitfall traps, 12–17 September 2008, 2_ (M.J.T.). Keramia
ClayPit,inburrow,26April2008,1imm.(A.R.-S.).Mt.Pelineon,nr.
rockybluff,inburrow,1May2008,1\(A.R.-S.). Marilynia bicolor (Simon, 1870)
Distribution: A common and widespread species in Records:Gridia,21April2008,1_(A.R.-S.).MarmaroMarsh,25
April2008,1_(M.A.).
Greece and also recorded from Turkey. It is probably
under-recorded on Chios because of the difficulty of Distribution:ThisspeciesisdistributedfromtheMedi-
locating its burrows. terranean to central Asia. In Greece it has only been
138 ChecklistofspidersofChios
recorded previously from the mainland in Attiki and Distribution: From North Africa, through the eastern
fromLesbos.ItwascollectedonChiosunderastonein Mediterranean to the Crimea and the Caucasus. On
dry phrygana and in tall moist grassland. Chios taken in the same habitats as D. crocata
Nigma flavescens (Walckenaer, 1825)
Records: Voulamandis House, Kampos, 16 May 2006, 1_ (R.S.). [Dysdera longirostris Doblika, 1853]
Voulamandis House, in garden, 27 April 2008, 1_ (C.S.). Kampia Records: [Parparia, litière de platane, 26 February 1982, 1 juv.,
Gorge,30April2008,3_(A.R.-S.). Deeleman-Reinhold&Deeleman,1988.]
Distribution: There is only one previous published Distribution: A species recorded from eastern Europe
record for this species from Greece, on the mainland in to the Ukraine.
Evvoia.OnChiosfoundinashadygorgeandgarden.It
is a West Palaearctic species.
[Dysdera punctata C. L. Koch, 1838]
Records:[Withoutlocalityorhabitatdata,Bristowe,1935.]
Nigma puella (Simon, 1870)
Records:MarmaroMarsh,swept,9May2006,1_(M.A.).Kampia Distribution: Recorded from southeastern Europe to
Gorge,10May2006,1\(M.A.).Ochrea,nearVolissos,12May2006, Georgia.
1\ (M.A.). Same site, 25 April 2008, 2\ (M.A.). Kampia Gorge, 30
April2008,1_1\(C.S.).KampiaBeach,30April2008,1_(M.A.).
Distribution: There are three previous records for N.
Dysdera romantica Deeleman-Reinhold, 1988
puellafromGreece,includingfromRhodesandLesbos.
Records: [Parparia, litière de platane au bord d’un ruisseau, 26
It is distributed throughout Europe and its offshore February1982,Deeleman-Reinhold&Deeleman,1988.]Mt.Pelineon
islands. Most specimens on Chios were swept in herb- (600m),14May2006,1\(R.S.).Ochrea,nearVolissos,12May2006,
rich grassland but one male was taken from under a 1\. Same site, 25 April 2008, 1\ (A.R.-S.). Salagonas Bay, 23 April
2008, 1\ (R.S.). Marmaro Marsh, 25 April 2008, 1\ (R.S.). Kampia
stone.
Gorge, 30 April 2008, 1\ (A.R.-S.). Elinda Beach, 6 May 2008, 1\
(A.R.-S.).Mt.Pelineon(600m),pitfalltraps,25March–12April2009,
1\;12–29April2009,1\(M.J.T.).
Nigma walckenaeri (Roewer, 1951)
Records:KampiaGorge,10May2006,2_(A.R.-S.).Ochrea,near Distribution: The species is endemic to Greece and
Volissos,12May2006,1_(A.R.-S.). reported from several of the eastern Aegean islands. On
Distribution: This is the commonest species of the Chiositwasfoundunderstonesinvarioushabitatsand
genusinGreeceandisrecordedthroughoutEuropeand in a winter-wet area of tall grassland.
as far east as Azerbaijan. On Chios, specimens were
sweptfromtheherblayerandcollectedinlitterbeneath
large oak trees. Harpactea cf. abantia (Simon, 1884)
Records: Marmaro Marsh, 25 April 2008, 1_ (R.S.). Ochrea, near
Volissos,25April2008,1_(A.R.-S.).
Scotolathys simplex (Simon, 1884)
Records: Salagonas, under stones, 15 May 2006, 1\ (M.A.). Distribution: A species endemic to Greece, previously
ManagrosBeach,11May2006,2\(M.A.).Agiasmata,22April2008, only recorded from the mainland. On Chios recorded
1\(R.S.). fromunderstonesbeneathlargeoaktreesandinanarea
of tall, moist grassland.
Distribution:WidespreadintheMediterraneanregion
and recently recorded from Lesbos (Bosmans et al.,
2009).CollectedonChiosunderstonesinadrymeadow
and near abandoned building. Harpactea sp. A
Records:Rockyvalleynr.Armolia,pitfalltraps,24March–4April
2009, 1_ (M.J.T.). Same site, 4–25 April 2009, 2_ (M.J.T.). Gridia,
Family Dysderidae trackS.ofvillage,pitfalltraps,24March–4April2009,2_.Samesite,
23April–5May2009,1_(M.J.T.).Mt.Pelineon(650m).pitfalltraps,
25March–12April2009,2_(M.J.T.).
Dysdera crocata C. L. Koch, 1838
Remarks:Thisspeciescannotbematchedwithanyof
Records: Gridia, track S. of village, 21 April 2008, 1_ (A.R.-S.).
Voulamandis House, Kampos, 27 April 2008, 1_ (A.R.-S.). the Harpactea males hitherto described from Greece or
Padoukios,pitfalltraps,6–22May2009,2_(M.J.T.).Samesite,22–31 Turkeyandiscurrentlybeingdescribed.OnChiositwas
May2009,1_(M.J.T.). only captured in pitfall traps in spring.
Distribution: A cosmopolitan species that is wide-
spreadinGreece.OnChiosoftenfoundaroundornear
houses. Harpactea sp. B
Records: Mt. Pelineon (650m), pitfall traps, 25 March–12 April
2009,1_;12–29April2009,2_;29April–7May2009,1_(M.J.T.).
Dysdera lata Wider, 1834
Remarks: Another species that cannot be matched
Records: Marmaro Marsh, 25 April 2008, 1_ (R.S.). Voulamandis
House,Kampos,27April2008,3\(A.R.-S.),2\(R.A.).Gridia,track withanyoftheHarpacteamaleshithertodescribedfrom
S.ofvillage,pitfalltraps,23April–5May2009,1\(M.J.T.). Greece or Turkey and a description is in preparation.
A.Russell-Smithetal. 139
Harpactea sp. rubicunda group A Distribution: Ubiquitous throughout Greece and the
Records:Ochrea,nearVolissos,25April2008,1_(R.S.).Marmaro Mediterranean region and distributed eastwards as far
Marsh,pitfalltraps,25April–1May2008,5_(A.R.-S.). asTurkmenistan.Femalesspinuntidysheetwebs,often
under stones but also under the loose bark of trees.
Harpactea sp. rubicunda group B
Records: Rocky valley nr. Armolia, 8 May 2006, 1\ (R.S.). Mt.
Pelineon(600m),9May2006,1\(R.S.).Gridia,trackS.ofvillage,21 Pritha nana (Simon, 1868)
April2008,1\(A.R.-S.). Records: Gridia, by track N. of village, 13 May 2006, 1\ (R.S.).
Kampia Gorge, near mouth of gorge, 30 April 2008, 1_ 1 imm.
Remarks:Itispossiblethatthefemalesofoneorboth (A.R.-S.).KampiaBeach,30April2008,1_(M.A.).
of these two different species may belong to the males
Distribution: This tiny circum-Mediterranean species
listed above but since they were taken at different
is apparently much less abundant in Greece than
localities they cannot yet be positively associated.
Filistata. It is nearly always found under stones in very
dry habitats.
Stalagtia christoi Van Keer & Bosmans, 2009
Records: Mt. Pelineon (650m), pitfall traps, 25 March–12 April
2009,5_;12–29April2009,2_(M.J.T.). Family Gnaphosidae
Distribution: This species has recently been described
from Lesbos by Bosmans and colleagues. On Chios it Anagraphis pallens Simon, 1893
wasrecordedfrompitfalltrapssetinalimestonebluffin Records:ManagrosBeach,22April2008,1\(A.R.-S.).
montane phrygana. Distribution: In Greece, this species has only pre-
viously been recorded from Crete and is otherwise
known from N. Africa and Israel. The specimen from
Family Eresidae
Chios was taken from beneath a stone in an olive
orchard.
[Eresus cinnaberinus (Olivier, 1789)]?
Records:[Withoutlocalityorhabitatdata(Werner,1935)]
Aphantaulax trifasciata (O. P.-Cambridge, 1872)
Remarks:Aspeciesnowregardedasanomendubium.
Records:KatoFana,8May2006,1\(R.A.).ManagrosBeach,11
It is possible that this was an erroneous record for E.
May2006,1_(M.A.).Anavatosvalley,2kmS.ofAnavatos,12May
walckenaeri. 2006, 1_ (M.A.). Ochrea, near Volissos, 12 May 2006, 1_ (R.S.).
Gridia, 16 May 2006, 1\ (R.A.). Gridia, 21 April 2008, 1_ (R.S.).
Samesite,25April2008,1_(A.R.-S.).Ochrea,nearVolissos,25April
Eresus walckenaeri Brullé, 1832?
2008,2_(M.A.).MarmaroMarsh,25April2008,1_(C.S.).Keramia
Records:Mesta,April2006,1_(M.J.T.).ManagrosBeach,11May ClayPit,26April2008,1_(C.S.),1_(R.S.),1_(M.A.).ElindaBay,29
2006,1_1\(M.A.).Samesite,1\(W.G.B.).Gridia,16May2006,1_ April2008,1_(R.S.).Rockyvalleynr.Armolia,pitfalltraps,15–29
(W.G.B.). Mt. Pelineon (650m), pitfall traps, 14–31 May 2009, 1_ May2009,1_(M.J.T.).
(M.J.T.).
Distribution: A widespread species in the Mediterra-
Distribution: Although likely to be this species, the
nean and distributed throughout the Palaearctic region.
commonest of the genus in the eastern Mediterranean,
Unlike most other gnaphosids, it is commonly swept
thereissomedoubtastherearenogoodrecentfiguresof
fromthefieldlayerofgrasslandsbutisalsooccasionally
thegenitaliaavailable.Thisspeciesisrecordedthrough-
found on the ground.
out the Mediterranean region. On Chios, specimens
collected under stones and males running on open
ground. Aphantaulax sp.?
Records:Agiasmata,ingrassnearderelictwatermill,22April2008,
1\(A.R.-S.).
Stegodyphus lineatus (Latreille, 1817)
Remarks:Theepigyneofthisspecimenresemblesthat
Records:SalagonasBay,27April1997,1imm.(C.Felton).Onhill
1.5km E. of Katavasi, 29 April 2008, 1 imm. (R.A.). Kato Fana, 4 of this genus but the abdomen lacks white spots.
May2008,1\(A.R.-S.).
Distribution: A widespread species throughout the Berinda ensiger (O. P.-Cambridge, 1874)
Mediterranean and extending eastwards to Tajikistan. Records: Rocky valley nr. Armolia, 8 May 2006, 1\ (A.R.-S.), 1_
On Chios it spins untidy cribellate webs in the upper (R.A.). Kampia Gorge, 10 May 2006, 1_ (R.S.). Giosonas, 10 June
branches of phrygana. 2006, 1\ (M.J.T.). Managros Beach, 11 May 2006, 1\ 1 imm.
(A.R.-S.),1_(R.A.).Anavatosvalley,12May2006,1_(R.S.).Gridia,
bytrackS.ofvillage,13May2006,3_2\(R.S.),1_(A.R.-S.).Kato
Family Filistatidae Fana, 15 May 2006, 1_ (M.J.T.). Salagonas Bay, behind beach, 15
May 2006, 1_ 1\ (R.S.); 23 April 2008, 1_ (A.R.-S.), 1_ (M.A.).
Giosonas,24May2006,1_(M.J.T.).SalagonasBay,6June2006,1\
Filistata insidiatrix (Forskål, 1775) (M.J.T.). Salagonas Bay, 23 April 2008, 1_ 1\ (R.A.). Ochrea, near
Records:KatoFanaBeach,27April1997,2\2imm.(C.Felton). Volissos,25April2008,1_(A.R.-S.).KatoFana,28April2008,1_
KatoFana,8May2006,1\1imm.(A.R.-S.).SalagonasBay,5June (A.R.-S.), 1_ (R.A.). Elinda Beach, 29 April 2008, 1_ (A.R.-S.).
2006,1\(M.J.T.).SalagonasBay,23April2008,1_1imm.(A.R.-S.). KampiaGorge,30April2008,1_(R.A.).Mt.Pelineon(650m),pitfall
Ochrea,nearVolissos,25April2008,1_(R.A.).KatoFana,27April traps,14–31May2009,1_(M.J.T.).SalagonasBay,12May2009,1_
2008,1\(A.R.-S.).Samesite,3May2008,1_2\(A.R.-S.). (M.J.T.).
140 ChecklistofspidersofChios
Distribution: A widespread species in Greece, also shallow cave, 27 April 1997, 1_ (C. Felton). Salagonas Bay, above
found in Turkey. Nearly always taken under stones in rocky headland, 27 April 1997, 1\ (C. Felton). Issidoros, 28 April
1997, 1_ (C. Felton). Managros, 30 April 1997, 1_ (C. Felton).
phrygana and other dry habitats.
Kipouries,pitfalltraps,9–24May1999,1_1\(M.J.T.).Mt.Pelineon
(600m),nearrockybluff,9May2006,1_3\(A.R.-S.),1_2\(M.A.),
1\(W.G.B.),1\(R.A.).Samesite,14May2006,2\(M.A.).Kampia
Berinda sp.
Gorge,10May2006,1\(A.R.-S.).ManagrosBeach,11May2006,2\
Records: Managros Beach, 22 April 2008, 1_ (A.R.-S.). On hill (A.R.-S.),2\(R.A.).Anavatosvalley,2kmS.ofAnavatos,12May
1.5kmE.ofKatavasi,2May2008,1\(A.R.-S.). 2006,1_(A.R.-S.).Gridia,13May2006,2\(A.R.-S.),1\(R.S.).Same
Remarks: These specimens do not correspond to any site,16May2006,1\(W.G.B.),1\(R.A.).Rockyvalleynr.Armolia,
13May2006,1\(R.S.).Mt.Pelineon,14May2006,1\(R.A.).Gridia,
of the three previously described species of Berinda.
21April2008,2\(R.S.),1\(M.A.).ManagrosBeach,23April2008,
Specimens were found under stones in degraded 2\ (M.A.), 1\ (R.S.). Salagonas Bay, 22 April 2008, 1\ (C.S.), 1\
phrygana. (A.R.-S.), 1\ (R.S.), 1\ (R.A.). Mt. Pelineon, 24 April 2008, 1_
(A.R.-S.).Rockyvalleynr.Armolia,28April2008,1_3\(R.S.).On
hill1.5kmE.ofKatavasi,29April2008,1\(A.R.-S.),1_(C.S.),1_
Camillina metellus (Roewer, 1928) (R.S.), 1\ (M.A.), 1\ (R.A.). Kampia Gorge, 30 April 2008, 1\
Records:Mt.Pelineon,nearrockybluff,14May2006,1\(M.A.). (A.R.-S.), 2\ (R.S.), 2\ (M.A.). Kampia Beach, 30 April 2008, 2\
Managros Beach, 22 April 2008, 1_ (R.S.). Salagonas Bay, 23 April (R.S.),2\(M.A.).Mt.Pelineon,2May2008,4_2\(M.A.).Neamoni
2008, 1_ (R.S.). On hill 1.5km E. of Katavasi, 29 April 2008, 1_ Monastery,2May2008,1\(R.S.).
(C.S.). Same site, 2 May 2008, 1_ (R.S.). Mt. Pelineon, near rocky Distribution:Thisisbyfarthecommonestmemberof
bluff,8May2008,1_(A.R.-S.).
the genus on Chios and is found from the eastern
Distribution: A species endemic to Greece, quite Mediterranean to Pakistan. It occurs in similar habitats
widely recorded in Crete but otherwise only previously to the last species.
known from Karpathos and Lesbos (Chatzaki, 2009).
Specimens from Chios were collected beneath stones in
Drassodes pubescens (Thorell, 1856)
phrygana.
Records: Marmaro Marsh, 9 May 2006, 1\ (R.S.). Same site, 25
April2008,2\(M.A.).
Cesonia sp. Distribution: D. pubescens is recorded throughout the
Palaearctic but is apparently an uncommon species in
Records:Rockyvalleynr.Armolia,understonesinmaquis,8May
2006,1\(M.A.). Greece,recordedpreviouslyfromCrete,Rhodes,Lesbos
Remarks:Theepigyneofthisfemaleappearsdifferent and Macedonia. On Chios collected in long grass and
reeds.
from that of Cesonia aspida, recently described from
Crete (Chatzaki et al., 2002b) but until further speci-
mens are found it would be unwise to describe it as a Drassodes serratichelis (Roewer, 1928)
new species. Records:SalagonasBay,15May2006,1_1\(M.A.).Onhill1.5km
E. of Katavasi, 29 April 2008, 1_ (C.S.). Kampia Gorge, 30 April
2008,3_(M.A.).
Cryptodrassus hungaricus Balogh, 1935? Distribution: Apparently, not a particularly common
Records: Gridia, 13 May 2006, 1_ (R.A.). Gridia, N. of village, speciesinGreecealthoughreportedfromseveralsitesin
pitfalltrap,29May–7June2009,1_(M.J.T.).
Crete by Chatzaki et al. (2002b). Elsewhere it has been
Remarks: These specimens show small differences recorded in Israel and the USA. Found on Chios under
fromthemaleofC.hungaricusbutcouldbeconspecific. stones both in dry areas and a shady gorge.
Drassodes lapidosus (Walckenaer, 1802) Drassodes sp.
Records: Mt. Pelineon (600m), near rocky bluff, 9 May 2006, 1_ Records:Ochrea,nearVolissos,25April2008,2_(R.S.).
(A.R.-S.),1_(M.A.),1_(R.S.),1\(W.G.B.),2_3\(R.A.).Kampia Remarks:Thesespecimenscouldnotbematchedwith
Gorge,trackintogorgebelowchurch,10May2006,1\(R.S.).Fyta any described species. Collected in litter beneath large
stream,10May2006,1\(M.A.).Mt.Pelineon,200mabovebluff,14
oak trees.
May2006,1_1\(R.S.),Mt.Pelineon,nearabandonedrefuge,14May
2006,1_(R.S.),2_1\(R.A.).Mt.Pelineon(600m),nearrockybluff,
24April2008,1_2\(C.S.),1_(A.R.-S.),4_(R.S.),4_3\(M.A.),2_
Drassyllus crimeaensis Kovblyuk, 2003
1\(R.A.).MarmaroMarsh,25April2006,1_(R.S.).KeramiaClay
Records: Rocky valley nr. Armolia, 8 May 2006, 1_ (M.A.).
Pit,26April2008,2_1\(R.S.).Mt.Pelineon,nearabandonedrefuge,
Anavatosvalley,12May2006,1\(R.S.).Mt.Pelineon,24April2008,
1May2008,2_(A.R.-S.),2_(R.S.).
1_(M.A.).
Distribution: This large robust spider, recorded Distribution: Originally described from the Crimea in
throughout Europe and as far east as the Urals, is less the Ukraine, this species has recently been recorded
widespread on Chios than the congeneric D. lutescens. from several sites in southern Turkey (Kovblyuk et al.,
All specimens were collected under stones in phrygana 2009). These records are the first for Greece. On Chios
and similar dry habitats. found under stones in phrygana.
Drassodes lutescens (C. L. Koch, 1839) Drassyllus praeficus (L. Koch, 1866)
Records: [Without locality or habitat data (Strand, 1917)]. Nr. Records: Kipouries, rocky stream, 28 April 1997, 1\ (C. Felton).
Armolia, 25 April 1997, 1_ 1\ (C. Felton). Kato Fana Beach, in MarmaroMarsh,9May2006,1_6\(A.R.-S.).ManagrosBeach,11
A.Russell-Smithetal. 141
May2006,1_1\(A.R.-S.).Ochrea,nearVolissos,12May2006,1\ Remarks: These specimens do not appear to be L.
(A.R.-S.). Managros Beach, 21 April 2008, 1\ (R.A.). Marmaro femineus or any of the other six species recorded from
Marsh, 25 April 2008, 1_ (R.A.), 1\ (A.R.-S.). Marmaro Marsh,
Greece. Collected under stones in phrygana.
pitfalltraps,25April–1May2008,9_2\(M.J.T.).Padoukios,pitfall
traps, 20 April–6 May 2009, 5_; 6–22 May 2009, 3_ (M.J.T.). Mt.
Pelineon(650m),pitfalltraps,1_,14–31May2009(M.J.T.).
Micaria cyrnea Brignoli, 1983?
Distribution: The commonest species of the genus in Records:Ochrea,nearVolissos,infieldinrelictoakwood,12May
Greece. Its distribution extends from Europe to at least 2006,1\(M.A.).
LakeBaikalinRussia.OnChios,particularlyabundant Distribution: Previously known only from Italy and
in the moist grassland of Marmaro Marsh but has also Corsica. There is some doubt about this record and
been found in leaf litter and under stones. males are needed to confirm it.
Drassyllus jubatopalpis Levy, 1998
Nodocion sp.
Records:Gridia,bytrackN.ofvillage,16May2006,1_(R.S.). Records:SalagonasBay,23April2008,1\(R.S.).
Distribution: This species was originally described
Remarks: This specimen could not be matched with
from Israel by Levy (1998) and has recently been re-
any described species known to J. Murphy but might
corded from Lesbos by Bosmans et al. (2009). It was
have been described under Poecilochroa. Collected
collected beneath chalky shale rock in phrygana on
under a stone in olive orchard.
Chios.
Nomisia aussereri (L. Koch, 1872)
Gnaphosa dolosa Herman, 1879
Records:ManagrosBeach,22April2008,1\(A.R.-S.).Hill1.5km
Records: Gridia, by track N. of village, 21 April 2008, 1\ (R.S.). E.ofKatavasi,2May2008,1_(A.R.-S.).Rockyvalleynr.Armolia,
ElindaBeach,29April2008,1\(R.A.).KampiaBeach,30April2008, pitfalltraps,26August–7September2009,1_(M.J.T.).Rockyvalley
1\(M.A.). nr.Armolia,pitfalltraps,7–18September2009,1_(M.J.T.).Gridia,
near shore N. of village, pitfall traps, 18–29 September 2009, 1_
Distribution: This species is distributed from southern
(M.J.T.).
Europe to Central Asia, but in Greece has only been
recorded from Crete and Lesbos. Collected beneath Distribution: A species distributed from the Mediter-
stones in phrygana near the coast on Chios. ranean to Mongolia and widespread in Greece. Col-
lected on Chios under stones in dry gravelly area and
degraded phrygana.
Haplodrassus dalmatensis (L. Koch, 1866)
Records:SalagonasBay,aboverockyheadland,27April1997,1\
(C. Felton). Managros, pitfall traps, 6 April 1999, 1_ (M.J.T.). Nomisia excerpta (O. P.-Cambridge, 1872)
Managros Beach, 11 May 2006, 1\ (A.R.-S.). Managros Beach, 22 Records:KampiaGorge,30April2008,1_(A.R.-S.).Onhill1.5km
April2008,1\(A.R.-S.). E.ofKatavasi,2May2008,1_(A.R.-S.).PaleaPotamia,9May2008,
Distribution: A reasonably common species in Greece 1_ (A.R.-S.). Voulamandis House, 27 April 2008, 1_ (R.S.). Mt.
Pelineon(600m),1May2008,1\(R.S.).
but known from only two sites on Chios. Found in low
herbsandgrassandunderastoneinadrygravellyarea. Distribution: A widespread and very common species
Its distribution extends from western Europe to central in Greece which is unaccountably scarce on Chios.
Asia. Otherwise recorded from the Canary Isles, Tunisia and
Israel. Collected on Chios under stones near stream
courses.
Haplodrassus silvestris (Blackwall, 1833)
Records:Managros,pitfalltraps,6April1999,1_(M.J.T.).
Nomisia sp.
Distribution: There is only a single previous record of
Records: Kampia Gorge, under stones by edge of stream, 10 May
this European species from Greece, from the island of
2006,1\(W.G.B.).
Lefkada in the Ionian island group. On Chios it was
collected in a pitfall trap in a dry stony area. Remarks:Theepigyneofthisspecimenisclosetothat
of N. excerpta but differs somewhat from those of
typical representatives of that species. Males will be
Leptodrassus femineus (Simon, 1873)
needed to correctly place the species.
Records:KatoFana,8May2006,1\(R.S.).Gridia,13May2006,
1_(M.A.).Samesite,16May2006,1_(M.A.).1_(R.S.).
Nomisia exornata (C. L. Koch, 1839)
Distribution: This southern European species is only
Records:Kipouries,pitfalltraps,9–24May1999,2_(M.J.T.).Mt.
knownelsewhereinGreecefromCretewhereithasbeen
Pelineon, 9 May 2006, 1_ 1\ (R.S.). Same site, 14 May 2006, 1_
recorded from several sites. Collected on Chios under
(M.A.), 1_ (W.G.B.). Rocky valley nr. Armolia, 8 May 2006, 2_
stones in dry phrygana. (R.A.);13May2006,1_(M.A.).Fytastream,10May2006,1\(R.A.).
Kampia Gorge, 10 May 2008, 1_ (R.A.). Salagonas Bay, behind
beach,15May2006,1\(M.A.).Gridia,bytrackN.ofvillage,16May
Leptodrassus sp. 2006,1_1\(R.S.).KatoFana,8May2006,1\(R.S.).Gridia,21April
Records: Kato Fana, under stones in phrygana, 8 May 2006, 1\ 2008, 2_ 1\ (A.R.-S.), 2_ (M.A.). Salagonas Bay, 23 April 2008, 1\
(R.S.).Gridia,16May2006,1_(R.A.). (M.A.).Mt.Pelineon,24April2008,1_(M.A.).KeramiaClayPit,25
142 ChecklistofspidersofChios
April2008,1_(M.A.).Rockyvalleynr.Armolia,28April2008,1_ Distribution: A species as widespread on Chios as
(R.S.),1\(A.R.-S.).Onhill1.5kmE.ofKatavasi,29April2008,1\ elsewhere in the eastern Mediterranean. Always found
(A.R.-S.), 1_ (M.A.), 1_ (R.A.). Kampia Gorge, 30 April 2008, 1_
running on the ground and under stones in phrygana
(R.S.), 1_ 1\ (M.A.), 1\ (R.A.). Rocky valley nr. Armolia, pitfall
traps, 15–29 May 2009, 2_ (M.J.T.). Padoukios, pitfall traps, 6–22 and other hot, dry habitats.
May2009,1_(M.J.T.).Mt.Pelineon,rockybluff,pitfalltraps,14–21
May2009,4_(M.J.T.).Rockyvalleynr.Armolia,pitfalltraps,5–15
May2009,3_(M.J.T.). Scotophaeus insularis Berland, 1936
Records:ManagrosBeach,11May2006,1\(R.A.).
Distribution: An abundant species throughout the
MediterraneanandextendingeastwardstocentralAsia. Distribution: Originally recorded from the Cape Verde
It is nearly always found running on the ground and islands. A single specimen was recorded from under a
under stones in dry habitats. stone on an eroded beach area.
Scotophaeus peninsularis Roewer, 1928
Nomisia ripariensis (O. P.-Cambridge, 1872)
Records:MarmaroMarsh,25May2008,1\(A.R.-S.).
Records:Limenas,pitfalltraps,25May–1June1999,2_(M.J.T.).
Rockyvalleynr.Armolia,8May2006,1_(A.R.-S.).Mt.Pelineon,8 Distribution: This species is recorded from Greece
May 2008, 1\ (A.R.-S.). Kato Fana, 8 May 2006, 1_ (A.R.-S.), 2\ (Arkadia and Crete) and Israel. Collected in long moist
(R.A.).ManagrosBeach,11May2006,1_(W.G.B.).ElindaBay,11
grassland on Chios.
May2006,2_(R.A.).Rockyvalleynr.Armolia,13May2006,1_1\
(A.R.-S.). Gridia, by track N. of village, 13 May 2006, 1\ (M.A.).
Samesite,16May2006,1\(M.A.).SalagonasBay,15May2006,3\
Scotophaeus scutulatus (L. Koch, 1866)
(M.A.), 1\ (R.A.). Gridia, 21 April 2008, 1\ (R.A.). Agiasmata, 22
April 2008, 1\ (R.A.). Salagonas Bay, 23 April 2008, 1_ (R.A.). Records: Rocky valley nr. Armolia, 8 May 2006, 1\ (A.R.-S.).
KeramiaClayPit,26April2008,1_(R.S.).KatoFana,28April2008, Gridia,bytrackN.ofvillage,13May2006,1\(R.S.).Samesite,16
1_ (C.S.), 3_ (A.R.-S.), 1_ (R.A.). Kampia Gorge, near mouth of May2006,1\(W.G.B.).ElindaBay,11May2006,1\(M.A.).
gorge,30April2008,1\(A.R.-S.).Padoukios,pitfalltraps,6–22May Distribution: A widespread species in Greece and
2009,2_(M.J.T.).
throughout the Mediterranean, also recorded in the
Distribution: Widespread and abundant throughout Ukraine,butnotparticularlycommononChios.Found
Greece, where it has similar habitat preferences to the under stones in phrygana and other dry habitats.
previous species. Its general distribution extends from
Bulgaria and Greece to Azerbaijan.
Scotophaeus sp.?
Records:Rockyvalleynr.Armolia,8May2006,1\(W.G.B.).Same
site,28April2008,1\(R.S.).Mt.Pelineon(650m),14May2006,1\
Poecilochroa furcata Simon, 1914
(M.A.).
Records:Rockyvalleynr.Armolia,28May2008,1_(R.S.).
Distribution: These female specimens do not corre-
Distribution:PreviouslyrecordedonlyfromFranceand
spond to any of the four species previously recorded
Italy. Collected under stones in degraded phrygana.
fromGreece.Foundunderstonesinphryganaandother
dry habitats.
Poecilochroa senilis (O. P.-Cambridge, 1872)
Records:Mt.Pelineon,9May2006,1_(M.A.).ElindaBay,11May Setaphis carmeli (O. P.-Cambridge, 1872)
2006,1\(W.G.B.).Samesite,13May2006,1\(A.R.-S.).Salagonas Records:Rockyvalleynr.Armolia,pitfalltrap,4–25April2009,1_;
Bay,2April2008,1_(M.A.). pitfalltrap,25April–5May2009,1_(M.J.T.).
Distribution: This species has been recorded from the
Distribution: This circum-Mediterranean species has
eastern Mediterranean to Turkmenistan but in Greece
been quite widely recorded in Crete. It was taken in
previously recorded only from a number of sites in
pitfall traps in phrygana on Chios.
Crete. Found under stones and in long, moist grassland
on Chios.
Setaphis sp.
Records:Agiasmata,nearbeach,22April2008,1\(A.R.-S.).
Pterotricha lentiginosa (L. Koch, 1837)
Remarks: The epigyne of this species does not appear
Records: Kipouries, pitfall traps, 9–24 May 1999, 3_ 1\ (M.J.T.).
to correspond to any of the seven species previously
Mt.Pelineon(600m),9May2006,2\(M.A.).KampiaGorge,10May
2006,1\(M.A.).Anavatosvalley,2kmS.ofAnavatos,12May2006, recorded from the Mediterranean region.
1\ (A.R.-S.). Gridia, by track N. of village, 13 May 2006, 1_ 2\
(A.R.-S.).Samesite,16May2006,2_(M.A.),2_(R.S.).Rockyvalley
nr.Armolia,13May2006,1\(R.S.).SalagonasBay,behindbeach.15 Synaphosus trichopus (Roewer, 1928)
May2006,1\(M.A.),1_(R.A.).Gridia,21April2008,1_(A.R.-S.), Records:Rockyvalleynr.Armolia,8May2006,1_(A.R.-S.).Same
1_(M.A.).SalagonasBay,23April2008,1_(R.S.).Armolia,28April site,13May2006,1_(M.A.),1\(R.S.).Nr.rivercourse,KatoFana,
2008,2\(R.S.),1_(R.A.).Mt.Pelineon,24April2008,1_(C.S.),2_ 8May2006,1_2\(A.R.-S.).SalagonasBay,behindbeach,15May
(M.A.).KeramiaClayPit,26April2008,1_(M.A.).KampiaGorge, 2006, 1\ (R.S.). Gridia, by track N. of village, 16 May 2006, 1\
30April2008,1\(R.S.),1\(R.A.).Onhill1.5kmE.ofKatavasi,2 (M.A.). Kato Fana, 27 April 2008, 2_ (A.R.-S.). Rocky valley nr.
May2008,1\(A.R.-S.).Gridia,N.ofvillage,pitfalltraps,29May–7 Armolia, 28 April 2008, 1\ (R.S.). Elinda Beach, 29 April 2008, 1_
June 2009, 1_ (M.J.T.). Mt. Pelineon (600m), 7–14 May 2009, 1_ (A.R.-S.).Rockyvalleynr.Armolia,pitfalltrap,29May–7June2009,
(M.J.T.). 1_(M.J.T.).