A New Variety of Schoenoplectus mucronatus (L.) Palla (Cyperaceae) from Japan

2004·0.78 MB·English

by  Yu Iokawa

植物研究雑誌
J. Jpn. Bot.
Originals 79: 1-3 (2004)
A New Variety of Schoenoplectus mucronatus (L.)
Palla (Cyperaceae) from Japan
Yu IOKAWAa，K azuhiro KOHNOb and Sigeru DAIGOBOc
aDepartment of Biology，Jo etsu University of Education，Jo etsu，Ni igata，9 43-8512 JAPAN;
bpukui Prefectural Pujishima High School，B unkyo 2-8-30，P ukui，9 100017JAPAN;
・
CDepartment of Comprehensive Studies，Jo etsu University of Education，Jo etsu，Ni igata 943-8512 JAPAN
(Received on May 8，2 003)
A new variety of Schoenoplectus mucronatus (L.) Palla，va r. antrorsispinulosus，is
described from Okayama Prefecture，s outhwestem Japan. This varlety is characteristic in
having antrorsely spinulose hypogynous bristles while other infraspecific taxa of the spe-
cies have retrorsely spinulose ones.
Key words: Cyperaceae，h ypogynous bristles，n ew variety，S choenoplectus mucronatus
rsispinulosus.
W紅.α~ntro
Schoenoplectus mucronatus (L.) Palla is plant as ad ifferent variety of S. mucronatus，
distributed widely in Eurasian continent，b ut and name it var. antrorsispinulosus after the
rarely in Japan (Kadono 1994). It is found on feature of hypogynous bristles. This variety
the margins of Showa-ike pond in Okayama， grows on the wet sandy place along the
southwestem Japan (Kobatake et al. 1999). stream at the m ginof the Showa-ike pond
征
We examined specimens from the habitat with Sanguisorbαofficinalis L.，Il ex serrata
and found that the hypogynous bristles of the Thunb.， Eupatorium lindleyanum DC.，
plant eantrorsely spinulose (Fig. 1a，b ). Miscanthus sinensis Andersson，I schaemum
訂
The feature is stable in the population of the aristatum L. subsp. glaucum (Honda)
habitat and entirely different from that of the T.Koyama，C yperus pilosus Vahl，R hyncho-
rest of the species which is retrorsely spora rugosa (Vahl) Gale and Scirpus
spinulose (DeFilipps 1980，E gorova 1999， wichurae Boeck. Although we examined
Koyama 1978: Fig. 1c，d) . Hypogynous bris- herbarium specimens of the genus in KPM，
tles esmooth，s cabrid or plumose in the KYO，M AK，T I，T NS and TUS，n o other 10-
征
genus Schoenoplectus (Goetghebeur 1998). cality of the variety has been found so far.
Spinules on the scabrid hypogynous bristles Further field surveys especially around the
are mostly retrorse in the genus except for a type locality are needed to clarify the
few species， in which the bristles are biogeographical extent of the variety.
retrorsely to antrorsely spinulose (Smith and
Hayasaka 2001). In a recent taxonomical Schoenoplectus mucronatus (L.) Palla in
work on the genus (Smith and Hayasaka Bot. Jahrb. 10: 299 (1888).
2002)，h ypogynous bristle differences，s uch V紅.antrorsispinulosus Iokawa，K .Kohno
as density and length of spinules 'econsid- & Daigobo，va r. nov. [Fig. 1e，f]
，紅
ered to have an enough value for dividing A typo setis hypogynis an仕orsespinulosis
taxa as varieties. We，t herefore，r egard this differt.
2 植物研究雑誌第79巻 第1号 平成16年2月
……一一
..官d.に(.，.場..四-t， _.同町". V...一"..~_ .~. 山哨払ω11・.a1“柚a・・.悼... ..-...・租---T叫
晶、加。 u..，‘".山・.伺包.岨・.内、匂..・..隠.(遁.'.陛、I同'‘6，_1"....“.. 帆..~..~.臥.)，'均(".r0、・.・......_亀
、羽."'"I
iLO.j:{川...._伽軸，MI\rJf"lfu..... 蜘陥"".
Fig. 1. Fruits with hypogynous bristles (a，c :do rsal view) and spinules on hypogynous bristles (b，
d) of two varieties of Schoenoplectus mucronatus，v ar. antrorsispinulosus (a，b ) and var.
mucronatus (c，d) ，ha bit (e: 23 Sept. 2002，Sh owa-ike pond，Ok ayama-shi) and type specimen
(f: Y. Iokawa & al. 6032，T US) of S.m ucronatus var. antrorsispinulosus.Sc ale bars = 1m m(a ，
c) or 0.1 mm( b，d ). Vouchers: var.αntrorsispinulosus (Y. Iokawa & al. 6032 TUS)，v ar.
mucronatus (Z. Tashiro，7 X 1917，K YO).
Type: Japan; Okayama Pre ，.f Okayama- Dr. Hiroyoshi Ohashi for his valuable advice
shi，O saki，S howa-ike. Y. Iokawa，K . Kohno on the manuscript. Thanks are due to Ms.
& S. Daigobo 6032，23 Sep. 2002 (TUS). Hiroko Kobatake for giving us information
Japanese name:I nu-hime-kang紅白 (nov.). on the locality. Wea re grateful to the cura-
tors of KPM，K YO，M AK，T I，T NS and TUS
Wew ish to express our sincere thanks to for allowing us to examine their specimens.
Febn，Iary 2004 Joumal of Japanese Botany Vol. 79 No. 1 3
References Kobatake H.，K ariyama S.，K atayama H.，Ki noshita N.
DeFilipps R. A. 1980. Scirpus. In: Tutin T. G.， and Enomoto T. 1999. Okayama-shi-san-syoku-
Heywood V. H.，B urges N. A.，M oore D. M.， butu-mokuroku. 153 pp. Okayama-shi-hoken-fuku-
Valentine D. H.，W alters S. M. & Webb D. A. shi-kyoku，O kayama. (in Japanese).
(eds.)，F lora Europaea 5: 277-280. Cambridge Koyama T. 1978. Cyperaceae. In: Li H. L.， Huang T.
，.c ，
University Press，C ambridge. Koyama T. & DeVol C. E. (eds.)Flora of
Egorova T. V. 1999. Cyperaceae. In: Fedorov A. A. Taiwan 5: 191-372. Epoch Publishing，T aipei.
(ed.)，Fl ora of Russia 2:102-304. A. A. Balkema， Smith S. G. and Hayasaka E. 2001. Delineation of
Rotterdam (Translation of Flora Everopeiskoi Schoenoplectus sect. Malacogeton (Cyperaceae)，
Chasti SSSR，t om 11. Na uka Publishers，Le ningrad， new combination，a nd distinctions of species. J.
1976，in Russian). Jpn. Bot. 76: 339-343.
Goetghebeur P. 1998. Cyperaceae. In: Kubitzki K. 一一-and一一一2002.New combinations within North
(edよThefamilies and genera of vascular plants 4: American Schoenoplectus smithii and S.
141-190. Springer，B erlin. purshianus (sect. Actaeogeton，C yperaceae) and
Kadono Y. 1994. Aquatic Plants of Japan. 179 pp. comparison with eastem Asian allies. Novon 12:
Bun-ichi Sogo Shuppan，T okyo (In Japanese). 106-111.
五百川裕，河野和博 ，大悟法滋 日本産ヒメカ
a b C.
ンガレイの1新変種
カヤツリグサ科フトイ属のヒメカンガレイ イ属の標本を調査したが これまでのところタイ
(Schoenoplectus mucronatus (L.) Palla)の新変種イ プ産地(岡山市大崎正和池)以外からは見つかっ
ヌヒメカンガレイ (v antrorsispinulosus lokawa， ていない.
訂.
K.Kohno & Daigobo) を記載した.この変種は刺 (a上越教育大学自然系生物学教室，
針が上向きであり，下向きの母種から明瞭に区別 福井県立藤島高等学校，
b
できる.国内の主要なハーバリウムにおいてフト 上越教育大学学習臨床総合学習教室)
c