A New Tachycarabus (Coleoptera, Carabidae) from the Southernmost Part of the Min Shan Mountains in Northern Sichuan, Southwest China

2006·3.6 MB·

by  Imura, Y

i:/、・t1-l1.T1)k、-().34(l):7 12,May20.2006
ANewTa hycarabus(Coleoptera,Carabidae) fromthe
Southernmost Part oftheMinShan Mountainsin
NorthernSichuan,SouthwestChina
Ytiki IMURA
Shinohara-cho l24)-S.Kuhoku-ku、Yokohama,222-0026Japan
and
Jaroslav KAL,、B
Jinacovicc119.66434Kuiim.(zcchRepublic
Abstract Ano、vcarabidbeetleofthegenusT(lc/1、・cal-cit)ll、.,isdescribedfromthe
southernmostpartofthe Min Shan Mountains in northern Sichuan、Southwest China.
underthenameCpc/)e、/-.
In thesummer of 2005, alongseries ofcarabidspecimens belonging to the
Rhlgocalabuscomplexwas foundfromthetwolocalitiesat thesouthernmostpartof
theMinShanMountainsstretchingfromnorthtosouthalongtheeasternbankofthe
MinJiangRiver.Afteracarefulcomparativestudy,wehavecometotheconclusion
that thespeciesinquestionmustbenewtoscience. Inthepresentpaper,wearegoing
todescribeitasanewspeciesbelongingto thegenusT11c/1、,calabus(sensuIMURA,
2002)inthesubtribeCarabina.IncasethesubtribeCarabinaisregardedasequivalent
tothegenusCa・labus(s lat)asisadoptedbymostEuropeanauthors,itwillbeplaced
inthesubgenusRhigoc(」1-abusof thegrandgenusCal・abus.Theabbreviationsused
hereinarethesameasthoseexplainedinpreviouspapersofthefirstauthor(cf.IMuRA,
1990,p.139;2002,p.130).
Beforegoinginfodescription,thefirstauthorwishestoexpresshissinceregrati-
tude Io Messrs. Igor BELoUsov(St.Petersburg,Russia)and IlyaKABAK(Almaty,
Kazakhstan)fortheirkindcooperation.ThanksarealsoduetoDr.Shun-lobiUENo
(Naljona1scienceMuseum,Tokyo)forreviewingthemanuscriptofthisPaPe「.
T‘acllycarabuspopekIMURAetKALAB,SP nov.
(Figs.l-3)
Length (jncluding mandibles): , 16.5-19.0 (arithmetic mean 17.7)mm;
,
、fzukyixI、wlUvltuAtasnrdqJpaoronslmavlkKj、ih_lAgllfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
17.0-19.5 (arithmetic mean t8.5)mm (in thepopulation from Zhenjiangguan);
,
14.6-17.3(arithmeticmeant6.0)mm;9,15.5-18.2(arithmeticmeant7.1)mm(inthe
population fromJiaochangba).
Uppersurfaceofbody notstrongly polished,withthecolorationdark reddish
copperytodarkbrownoftenwithafaintgreenishtinge,ornearlyentirelyblackishor
greenishinsomeindividuals;venterandappendagesbrownishblack,thoughtibiaeand
basalpartsofmandibles,paIpiandantennaearealittlemorestronglyreddish.
Headnothypertrophicinbothsexes,withtheeyesalmostnormal insizeandde-
greeofprotrusion; fronsweaklyconvexabove,oftenroughlyrugu1oseandsometimes
scatteredwithsmall punctures; frontal furrowsnotsowideandnotdeeplyconcave,
andirregularlyrugu1oseonthesurface;vertextoneck irregularlyandroughlywrin-
kled and often sporadically scattered with vaguepunctures; retinaculum of right
mandiblenarrowerandsmallerthanthatoftheleftone,withtheanteriortoothusually
alittleshorterthantheposterioronbothsides;terminalsegmentsofpalpinotremark-
ablydilatedinbothsexes;penultimatesegmentoflabialpalpusbisetose;mediantooth
of mentum shorter than lateral lobes, slightly protruded ventrad, and triangularly
shapedthoughnotsharply pointedat thetip;submentumbisetose;antennaenotso
long,barelyreachingbasalthirdofelytrainmaleandreachingbasalquarterinfemale;
hairlessventraldepression,orthiridium,ofmaleantennaerecognizable fromsegment
5 to9or10.
Pronotumrathersmall,subcordate,muchwiderthanlongandwidestneartheapi-
calquartertothird;apicalmarginweaklyemarginate, frontanglesobtuselyrounded
andhardly protrudedanteriad; lateral sidesdistinctlymarginedthroughout, gently
roundedinfrontandeithersinuatelyornearlystraightlynarrowedtowardshindangles
which aresubtriangularlyproducedposteriadwithblunt tips; discweakly
convex
above,withthesurfaceweaklywrinkledandsometimesvaguelypunctateinmedian
portion,oftenbecomingratherscabrousinperipheralportion;basal foveaesmall,not
So deeply concavethoughdistinct, andmedian longitudinal line clearly impressed
th「oughout;onetothreemarginalsetaeinsertedoneithersideofpronotum,oneortwo
inmedianportionandzerooronebeforehindangle.
Elyt「aelongatedoval,weaklyconvexabove,widestalittleorobvjouslybehjnd
themiddle,moregraduallynarrowedtowardsbasesthantowardsapjces,wjtheffaced
ShOulde「S; late「alSidesgentlyarcuatethroughoutthoughoftennearlyslrajghtbefore
theWidestPartinmale,withthemarginsnarrowlyreflexedabove;sculpturetr_j or
hePtaP1oidhete「odyname-prmi areisthestrongest,ratherirregularlysegmentedby
Shallow, notSo la「gep「imary foveolesto form rows of narrow costae; secondaries
muchWeake「andnarrowerthanprmi aries,indicatedbylongitudinallyarrangedrows
ofSmall9「anuIeS・Pa「flybecomingcontiguoustoformirregularlyjnterruptedcostae,
to「tia「ieSalsoindicatedbyrowsofsmallgranulesasinsecondarjes,oftenseparated
intotwo「oWStofo「mquarternariesandsometimesadhesivetoacjjacentjnterva1sto
fo「m「etCiula「Pattern;umbilicateseriesindicatedbyirregularlyandsporadicallyset
「oWSoffine9「anuleS;eachelytronwithashallowbutratherremarkabledepressiona
NewTaclnca,・abtlsfromNorthernSichuan
Fjgs.1_2. Tahyca,abuspopekfromSEofZhenjiangguan(4,100-4,185m)innorthernSichuan. - 1,
holotype,2, ,paratype.
,
littlebeforeapex.
Epjsternaalmostsmooth,sidesofsternitesweaklywrinkle sternal sulci un「eC-
ognjzed;metacoxatrisetose;basalfoursegmentsofmael foretarsusdilatedWithhai「
padsonventralsurface.
MalegenjtaljaasshownjnFig 3;aedeagusslender,gentlyarcuateth「eu9hOut
andweaklybentventradapicadinlateralview;apical loberathe「She「t,about1・5
tmi esaslongaswjde,fjnger_tjplikeinshapewiththeapexobtusely「oundedinlate「al
viewandsubtrjangularlyshapedindorsalview;0Lun1iobedand「athe「Small;1i9ulum
Indicatedbylongitudinallysetrowsofgranuelstoformanarrow「id9e;neithe「BLno「
MLrecognizedthoughbasalpartofendophallusnearaedeaga1apexaPPa「entlyin-
ated;PREIndicatedbyapajrofhemispherica11yprotrudedhai「lyinflations;PAR
ratherstronglyandsymmetricallyprotrudeddorsadonbothSides; PP ta「9e,hemi-
atedandnotremarkablypigmented;ALweaklyinflated;PLnotSola「9
spher1ca11yIn
butwellrecognizable;AGGweaklysclerotizedandpigmente indicatedbyaPal「o
smallterminalplateweaklyprojectedtowardsin exedSidesofendOPha11uS・
YukiIMURAandJaroslavKALAB
10
Malegenjta1organofTachycalabusppek. - a,Aedeaguswithfullyeve「tedendOPhaliuSin
Fig 3
rightlateralvjew;b,apicalpartofaedeagusinrightlateralview;c,dittoindo「SalView・Scale:1mm
fora;0.5mmforb&c.
Holotype: (S,SEofZhenjiangguan [領江美] (32° l7'54-57"N/
Type series.
103°47'56″_47'56″E), 4,100-4,185m in altitude, in southeastern SOngPan Xian
],ofnorthernSichuan,SouthwestChina,17-VII-2005,tobedepositedinthe
[松潘
Departmentof Zoology,NationalScienceMuseum(Nat. Hist), Tokyo.ParatypeS:
13 14 ,samedataas for theholotype; Ie 12 ,samearea (32°17'57″一
, ,
18'17″Nile3°47'56″一48'07〃E, 4,185-4,421m in altitude), 20-VII-2005; 42
,
45 ,8kmESEofJiaochangba[較i ](=Jiaochang[較%i ])(32°01'N/103°46'E),
4,300m inaltitude, innorthernMao Xian [茂 ], ofnorthern Sichuan,Southwest
China, l5~l7-VII-2005,separately preservedinthecollectionsoftheauthors, the
ZoologicalInstituteofAcademyofSciences(St.Petersburg)andB.BREzlNA(Prague).
Notes. Thepresent new specieswascollected fromtwodifferentsites in the
alpinezoneover 4,100monthesamemountainrange. Thepopulation fromZhen-
jiangguanissomewhatdiffferent fromthatofJiaochangbainhavinglargersizeonav-
erage,relativelylargepronotum,longerandslendererelytra,narrowerprimarycostae,
etc.Alltheseare,however,consideredtobeamerevariationwithinthesamespecies
mainlyduetothedifferenceofaltitudeofeachlocality,andwehesitatetoseparate
themintotwodifferentsubspecies.
The new species is characterized by less hypertrophic head, irregularly and
roughly rugose-punctatedorsalsurfaceof theheadandpronotum, relativelysmall
pronotum,onlyweaklyconvexpronotumandelytra, tri-orheptaploidheterodynamjc
elytra1sculpturewiththestrongestprimarycostae,andlongandslenderappendages,
NcwTlac/n'(,t1;・alii、.,fromNorthernSichuan
etc.
ItismostcloselyalliedtoTpus1oSEMENow(1898,p362)described from“Sun_
pan” (=Songpan)ofnorthernSichuan,but isreadilydiscriminated fromthatspecies
mainlybythefollowingrespects:1)eyeslessstronglyprotrudedlaterad;2)pronotum
relativelysmall,withmuchmoreunevendiscaIsurface,deeperbasal foveaeandless
stronglyprojectedhindangles;3)elytrausuallyalittlelongerandslenderer,withmore
effacedshoulders;4)aedeagusslenderer, lessstronglyarcuate,withthemedianper_
tionlessstronglyinflatedrightlaterad,apicallobealittlelongerandslenderer,andless
remarkablybent right lateradindorsal view;endophalIuswith 0LandPREsmaller,
PARlargerandmorestronglyprojecteddorsad.
FromTpseudopus1oDEuvE(1996,p 22)described from“va11eea10kmausud-
est deSanggarpar, env 4200metres” ofnorthernSichuan, T popeknov. is distin_
guishedbymuchslendererbody,lesshypertrophicboa lesstransversepronotumwith
shorterhindangles,andshorterapicallobeofaedeaguswhichislessremarkablybent
ventradin lateralview.
ItisnecessarytocomparethenewspecieswithTko1・sake、、,1SEMENo、v(1898,p.
361)described from“Ta-tz'ao-pinsupraangustiasCho-dgi-gou,baudprecutaburbe
Lun-ngan-fu,att ic.-13.000'sm” Regrettably,however, itisimpossibletomakea
fullcomparativestudybetweenthetwospeciesbasedonbothsexes,sinceSEMENow's
specieshasbeenknownsofaronly fromasinglefemalespecimen.So faraswehave
examinedtheholotypeof T、/,orsako、、,1nowpreservedintheZoological Instituteof
Academy ofSciences(St.Petersburg), thenew speciesseemstobedistinguishable
from SEMENows' raceby lightercolorationofappendages, narrower pronotum with
more remarkably protruded front angles and less strongly uneven discal surface,
shorterandrobusterelytrawithnarrowerprimarycostaeandlessstronglydeveloped
otherintervals.
Thenewspeciessomewhatresembles (?)/ll7fa17g1ctl.''DEUvE(2001,p 90)de-
scribedfrom“environsdoJintang,4200metres''ofSichuan,buttheformerisreadily
discriminatedfromthelatterasfollows:1)appendageslongerandslenderer;2)dorsal
surfaceofheadandpronotumlessroughlyrugose-punctate;3)pronotumdifferent in
shape,wjththelateralsjdesmoreroundlyarcuateinfront,moreremarkablysinuate
behjnd,thedjscmorestronglyconvexaboveandthehindan9lesnarrowe「andme「e
sharplypojntedattheapices;4)elytraalittlelessstronglyconvexabove.Withthelat-
eralsideslessroundlyarcuate;5)elytra1discdifferentlysculpture with theP「ima「y
costaealilliewjderandsegmentedbydeeperprimaryfoveoleStheSeCOnda「ieSand
elr1lar1esindicatedbylessregularlysetrowsofgranulesandthequa「te「na「ieSme「e「e-
duced;6)apicalportionofaedeaguslessstronglybentvent「adinlaet「alView,Withthe
tipnotSubquad「atebutobtusely「ounded
resemblethoseof‘‘ ca,.a加s” eli(0g1-川SE_
Aedeaga1featuresoftheneWSPeCieS
MENov(lg03,p 349)describedflem Ju9・Amnenkol'alt 13 _14000,s m,, the
mounatinrangelocatedbetweenthesourcesofthe「VIe「SHuan9Heand h 19J1an
HowevertheformerIseasilydiscrmi inatedfromthelatte「inta「9e「Size・Sonclef.Pr
l2 Yuki IMURAandJaroslav KALAB
portionwithlesshypertrophichead,muchmoreunevendorsalsurface,etc.ItSi most
, probable that SEMENovs' speciesbelongs, together with . 'c o''ows ｽll t ''aeo-
calabusjntheRhigocarabuscomplexaccordingtothemolecularphy1ogenetiCa1Study
(IMURA,2002,p i t).
Etymology. Thenewspecificnamecomesfromthatofacolleagueofthesecond
author,Mr.Popek(=Josef)SKoTAK.PopekisanotherdenominationofJosefinCzech
language.
要 約
井村有希・J.KALAB:中国四川省北部から発見されたプーシオードゥガネオサムシ属の1新
種. _ 2005年の夏, 中国四川省北部松活 から茂 にかけての山民江東岸を南北に走る山民山
山脈最南端部から, プーシオードゥガネオサムシ属に属する未記載種が発見されたので,
Tach、ca,abuspepekという名を与え,本論文において記繊した.
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