A New Species of Tiberioides (Coleoptera, Passalidae) from Myanmar, with a Key to the Species of Tiberioides

2007·6 MB·English

by  Kon, M

Elyt・,a,Tokyo,35(1):313-318,May30,2007
A NewSpeciesofTiberioldes (Coleoptera,Passalidae) from
Myanmar, witha Key to theSpeciesofTiberioides
Masahiro KoN
SchoolofEnvironmentalScience,TheUniversityofShigaPrefecture
Hassaka-cho2500, Hikone,Shiga,522-8533Japan
and
Kuni0 ARAYA
GraduateSchoolofSocialandCulturalStudies,KyushuUniversity,
Ropponmatsu4-2-1,Chuo-ku,Fukuoka,810-8560Japan
Abstract AnewspeciesofTiberioldesisdescribedfromMyanmarunderthe
nameof 「. er/ey1sp nov. 「fber1oldes erfeyl'sp nov resembles「. kMwert1(ARROW)
inhavingtheelytronmatinthelateralgrooves,butcanbedistinguishedfromthelatter
bytheanteriorlowertoothsimpleattheapex. AkeytothespeciesofTiber1oldesisalso
provided.
Thegenus Tiberioldeswasestablished byGRAVELY (1912) for Tiberius kuwert1
ARROW, l907. Up to thepresent, threespeciesof Tiberioides, T kuwerti (ARROW,
1907),T borealis(ARROW,1907),andT austen1GRAVELY, 1914, havebeen known
fromtheeasternHimalayasandMyanmar(GRAVELY,1914;ARROW,1950;KoNetal.,
1999a,b).
WhenweexaminedaseriesofTiberioldesspecimenspreservedinthecollectionof
the Natural History Museum, London, we found aseriesofspecimensof aspecies
distinct fromall theknownmembersof thisgenus. After a close examination and
comparisons,weconcludedthat this formwasnewtoscience. Thereafter, weobtained
someadditionalspecimensof this formcollected from Kachin, Myanmar. Thus, we
describeanewspeciesofTiberioidesfromMyanmarbasedontheabovespecimens. In
addition, weprovideakeyto thespeciesofthisgenus.
Tiberioideskerleyi KoNetARAYA,sp nov.
(Figs.1-5)
Descriptionofholotype. Male. Bodylengthfromanteriormarginofhead totip
ofelytron: 35.0mm.
314 Masahiro KoNand Kunio ARAYA
Fig.1. HabitusofTiberioldeskerleyisp nov.,male,scale,10mm
Anteriorangleofheadrounded;outer tuberclessymmetrical,triangular,vertically
bifidatdistalends;lowertipofouter tuberclealittlemoreproducedforwardsthanthe
upperone; distancebetweeninner tuberclesalittlelonger than that between innerand
outer tubercles; tip of inner tuberclemuch larger than that of outer tubercle; ridge
between inner and outer tubercles distinct; ridgebetween inner tubercles distinct;
anteriorareabetweenouter tuberclessmooth, hollowedalittle;uppersurfaceof head
impunctate and hairless, rugose in areabehind outer tubercle and depressed area,
smooth around parietal ridge; parietal ridge blunt; central tubercle low and blunt;
canthusnot broadened distally, with aridge on proximal portion of upper surface;
frontalridgevanishingpriortoinner tubercle.
Mandiblessymmetrical; upper toothroundedat apex; upper marginofmandible
concavebehinduppertooth,swollennearbase;loweroutermarginofmandibleangulate
nearbase; lowest terminal tooth largerthananterior lower tooth;anterior lower tooth
triangular,simplypointedatapex;upperinnersurfaceofmandiblepuncturedandhairy
in proximal portion. Anterior margin of labrum concave, left angle a littlemore
produced forwardsthan theright one; ligulaweaklypointedat themiddleofanterior
NewTibe・tiotdesfromMyanmar 315
Figs 2-3. Tiberioldeskerley1sp nov.,male,scale,1mm;2,anteriorpartofhead,indorsalview;3,
mentumandligula
margin,withdistinctmediankeel;mentumimpunctateandwithoutdistinct transverse
carinaincentralportion,puncturedandhairyinlateralportion;submentumpunctured
asawhole;hypostoma1processimpunctateandhairless. Antennaewithsixpubescent
lamellae,thefourthtosixthlamellaelong,thethirdone0.5timesaslongasthefourth
one,thefirst andsecondveryshort.
Pronotumwithveryslightmediansulcus,withshallowlateralscar,withafewfine
puncturesinlateralscar;posteriorplateofprosternumimpunctateandhairless;scutel-
lumimpunctateandhairless;mesothoracicepisternunimpunctateandhairlessinlower
portion,puncturedinupperportion;mesosternumimpunctateandhairless,withshallow
slightscar; posterior intermediateareaofmetasternum impunctateandhairless, with
irregular dents along posterior margin of central area; anterior intermediate area
puncturedand hairy in inner portion; ridgebetween lateral and intermediate areas
distinct;lateralareadenselypuncturedandhairy;centralareaimpunctateandhairless.
Lateralgroovesofelytronmat,wider thanadjacent ribs,withoblongpunctures.
Firstvisibleabdominalsterniteimpunctateandhairless;secondvisibleabdominal
316 Masahiro KoNand Kunio ARAYA
Figs 4-5. MalegenitaliaofTiberioldeskerleyisp nov.,male,scale,1.0mm;4,ventralview;5,right
lateral view.
sternite punctured and hairy behind transverse ridge; 3rd to 6th visible abdominal
sternitesimpunctateandhairless.
Penisofmalegenitalialarge,withlongitudinalmembranousareaalongmidlineon
ventralside;parameresunitedonventralside,shorter thanbasalpieceinventralview,
latero-proxima1endpointedproximallylikeahook;distalmarginofbasalpieceobtusely
V-shapedinventralview, lateralmarginsdivergentdistally.
Variation. Nosexualdimorphismisevident. Bodylengthofparatypes,37.9mm
±1.65,34.6-40.6mm(mean±SD,range,N=18).
Typeseries. Holotype: Sin Lum,Bhamo,Upper Burma, 6000ft, l908. Para-
,
types:3 2早早,thesamedataasfortheholotype;1(,'l,2早早,Bhamo,UpperBurma,
,
1910; l 4Q9,ditto,1911;3 2早早,Dahtingzen,2000m,N.E.Kachin,Myanmar,
, ,
VI-2000. Theholotypeisdepositedinthecollectionof theNaturalHistoryMuseum,
London.
Etymology. ThisspeciesisdedicatedtoMr.M.KERLEY,whogaveuskindhelp
duringourstayintheNaturalHistoryMuseum,London.
Notes. Tiberioideskerley1sp nov. isclosely related toT kuwerti (ARROW),but
canbedistinguidhed fromthelatterbyhavingthefollowingcharacters: theanterior
lower toothsimply pointed; theupper tooth rounded; thescar ofmesosternumvery
slight.
New llberio1clesfromMyanmar 317
KeytotheSpeciesofTiberioides
1. Lateralgroovesofelytronwide,mat,withoblongpunctures 2
- Lateralgroovesofelytronnarrow,notmat,simplypunctured
2. Anterior lower toothsimplypointedindorsalview_____ T kerleyisp nov
- Anterior lower toothbidentatedista1o-proximaliy T kuwerti
3. Mentumwithstrongtransversecarinaincentralportion T borealis
- Mentumwithout transversecarinaincentralportion _. 「. austem
.
Specimenscompared. Tiberioidesausteni GRAVELY:cotype,sexunknown,Dikran
V.,ANDREwEsBequest(inthecollectionoftheNaturalHistoryMuseum,London);T
borealis (ARROW): holotype, sex unknown, Assam Nagas (in the collection of the
Natural History Museum, London); T kuwerti (ARROW): holotype, sex unknown,
Nepal (inthecollectionoftheHessischerLandesmuseum,Darmstadt).
Acknowledgments
WewishtoexpressourcordialthankstoW.SCHNEIDER, theHessischer Landes-
museum, Darmstadt, M. KERLEY, the Natural HistoryMuseum, London, andM.
MATsUI,KyotoUniversity, forgivingustheopportunitiestoexaminethetypespeci-
mens. ThisstudywassupportedinpartbyaGrant-in-AidfromtheJapanSocietyfor
thePromotionofScience(No. 18405012).
要 約
近 雅博・荒谷邦雄: ミャンマーからのnberlo1des属クロツヤムシの1新種とnber1o1des属の
種への検索表. - ミャンマーからTiberioides属の1新極を記載し, Tkerleyispnov.と名付け
た. T kerleyisp nov.は, 側面の満が 消しの華肖 をもつ点でT kuwert1(ARROW)に似ている
が, anteriorlowertoothの先端が単純に尖っていることによって区別できる. さらに, Tiberioides
属の種への検索表を作成した.
References
ARROW,J.A.,1907.AcontributiontotheclassificationoftheColeopterafamilyPassalidae.Transcfit.Soc.
London,1906:441-469.
1950. Coleoptera, Lame11icornia, Lucanidae and Passalidae, vol. IV. In the: Fauna of India
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IncludingPakistan, Cey1ott,Burmaa,tdMalaya.Taylor&Francis,London.
GRAVELY,F.H.,1912.PreliminaryaccountofarevisedclassificationofIndo-AustralianPassalidae.J.Aslat.
Soc.Bengal,8:403-407.
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Me,n. Ind. Mus.,3:177-353.
KoN,M.,M.TANAKA&K.ARAYA,1999a.Passalidbeetles(Coleoptera,Passalidae)COllectedf「omBhutan,
withdescriptionofthemalegenitaliaofTiberioidesaustenlGRAVELY.Elytra,Tokyo,27:471-474.
318 Masahiro KoNand Kunio ARAYA
KON,M.,M.TANAKA,K. ARAYA&Y.NOSE,1999b.Passalidbeetles(Coleoptera,Passalidae)collected
fromnorthernMyanmar,withnewrecordsofCeracupeschingkin1andTiberioidesborealis.Elytra,Tlokyo,
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Elytra,Tokyo,35(1):318,May30,2007
ANewRecordof Heliocopristyrannus
(Coleoptera,Scarabaeidae) fromBorneo
TeruoOcHIi)andMasahiroKoN2)
1)Kofudai5-21-6,Toyono-cho,Toyono-gun,Osaka,563-0104Japan
2)SchoolofEnvironmentalScience,TheUniversityofShigaPrefecture
Hassaka-cho2500,Hikone,Shiga,522-8533Japan
HeliocopristyrannusTHOMSONhasbeenrecordedfromMyanmar(Tenasserim),theMalay
Peninsula,Sumat「aandJava(BALTHASAR,1963). ThroughthecourtesyofDr.Y.CAMBEFoRT,
wehaveexaminedaspecimenofthisspeciesfromBorneopreservedinthecollectionofMNHN,
Paris. ThisisthefirstrecordofthegenusHeliocoprisfromBorneoaswellasthatofH tyrannus.
The collection dataareasfollows: 1ex.,MittenBorneo,21-X-1925, H.F.SIEDERsleg., R.
PAULIAN dot.
Reference
BALTHASAR,V.,1963.MonographicderScarabaeidaeunciAphodiidaederpalaearktischenundorientalischen
Region,Band1392pp.Tschechos1owakischeAkademiederWissenschaften,Prag.