A new species of Stenelmis (Coleoptera: Elmidae) found west of the Mississippi River

11 Pages·1991·5.2 MB·

by  K L Schmude

PROC. ENTOMOL. SOC. WASH.
93(1), 1991, pp. 51-61
A NEW SPECIES OF STENELMIS (COLEOPTERA: ELMIDAE)
FOUND WEST OF THE MISSISSIPPI RIVER
Kurt L. Schmude and Harley P. Brown
(KLS) Department ofEntomology, University ofWisconsin, Madison, Wisconsin 53706;
(HPB) Department ofZoology, The UniversityofOklahoma, Norman, Oklahoma 73019.
—
Abstract. \d\x\is ofa new Nearctic riffle beetle, Stenelmis occidentalis, are described.
Its range is restricted to areas west ofthe Mississippi River where it is one ofonly a few
species in this genus to occur in Mexico, and the only Stenelmis known to occur in the
northwestern U.S., Montana, Oregon, and Utah. Diagnostically useful characters include
size, pronotal sculpturing, elytral color pattern, comparative lengths of the tarsomeres,
metatibial spinous ridge on males, and the distinctive male genitalia. Adults are most
similar to adults of Stenelmis decorata, and differences between the two, and adults of
other species, are discussed. Adults occur in several types oflotic situations and they are
found on a variety ofsubstrates; they are also readily attracted to lights.
Key Words: Elmidae, Stenelmis occidentalis, new species, riffle beetle, western United
States, Mexico
StenelmisDufouris the mostdiverse and from Oregon have also been mentioned
widespreadNearcticgenusofElmidae(riffle (Brown 1972). But structuraldifferencesbe-
beetles),with30describedspeciesoccurring tween northern and southern individuals,
throughout most of North America north and their similarity to adults ofother spe-
ofMexico(Brown 1983, 1987,White 1982). cies, some of which are yet undescribed,
The senior author is revising the genus in preventedthejuniorauthorfromdescribing
North America, and this paper is part of thespeciesseparatefromarevisionarystudy.
that study. Adults of the following species
Materials and Methods
are beingdescribed at thistime to make the
nameavailable foraconcurrent revisionary Extensive collectingwas accomplished in
study of the riffle beetles of mid-America the southern range ofthe species by theju-
andtheWestIndies(SpanglerandSantiago, niorauthor, andadditional materialwasex-
in prep). amined from the following collections:
This "new" species ofStenelmis actually CASC—California Academy of Sciences,
has been known for some time. Texas spec- San Francisco, D. H. Kavanaugh & R.
imenshavebeenmentionedintheliterature Brett
as Stenelmis sp. (near decorata) as early as CNCI—CanadianNationalCollections, Ot-
1963 by Burke, and by Stewart et al. (1973) tawa, Ontario, L. LeSage
and Poole and Stewart (1976) as Stenelmis INHS—Illinois Natural History Survey,
mexicana(us), one of Brown's manuscript Urbana, K. C. McGiffen
names. In studies on a Montana stream, MCZC—MuseumofComparativeZoology,
Gore (1975, 1977, 1978) referred to this Harvard University, Cambridge, Mas-
species as Stenelmis sp. a, and specimens sachusetts, S. R. Shaw, D. Furth, S. Pratt
52 PROCEEDINGS OFTHE ENTOMOLOGICALSOCIETY OFWASHINGTON
LSUC—Louisiana State University, Baton PW/PL—ratio ofpronotal width vs length;
Rouge, C. B. Barr, J. B. Chapin EL—maximum length of elytra measured
NMNH—National Museum of Natural fromanteriormargintoapexalongelytral
History, Smithsonian Institution, Wash- suture with beetle's venter situated in
ington, D.C., P. J. Spangler horizontal plane;
ODAC—Oregon Department of Agricul- EW—maximum width ofelytra;
ture, Salem, R. L. Westcott EL/EW—ratio ofelytral length vs width;
PERC—Purdue University, West Lafay- PE length—length ofpronotum and elytra,
ette, Indiana, A. Provonsha measured separately, and summed.
TAMU—Texas A&M University, College
Station, H. R. Burke, E. G. Riley Stenelmis occidentalis
UGAM—UniversityofGeorgiaMuseumof Schmude and Brown, New Species
Natural History, Athens, C. L. Smith
UMRM—University of Missouri, Colum- Stenelmis sp. (near decorata) Burke, 1963:
113.
bia, R. L. Blinn, K. B. Simpson
Stenelmiss^.^GovQ, 1975:215, 1977: 115,
USUC-Utah State University, Logan, W.
1978: 147.
Hanson
WSJ.UC—Washington State University, Stenelmis mexicana Stewart et al., 1973:
960 [nomen nudum].
Pullman, R. S. Zack Stenelmis mexicanus Poole and Stewart,
CBB—
Cheryl B. Barr, Sacramento, Califor-
1976: 154 [nomen nudum].
nia
HPB—Harley P. Brown, Oklahoma Muse- Holotype m2At.-Head: lOW: 0.38 mm.
um ofNatural History, Norman Median dark band between eyes narrowed
JRD—Jack R. Davis, Texas Water Com- posteriorly; each light band equal in width
mission, Austin to dark band at point between eyes. Anten-
DLG—Daniel L. Gustafson, MontanaState nae and palpi testaceous. Antennal and
University, Bozeman pronotal lengths equal.
EGR-Edward G. Riley, Texas A&M Uni- Pronotum (Figs. 1, 2): PL: 1.00 mm, PW:
versity, College Station 0.91 mm. Widestjust behind middle; sides
KLS—KurtL. Schmude, UniversityofWis- narrowed toward base with a slight sinua-
consin, Madison tion before base; abruptly narrowed before
WDS-WilliamD. Shepard,CaliforniaState middle, then subparallel to apex. Median
University, Sacramento sulcus deep, widest and deepest anteriorly,
A Hitachi S-570 scanning electron mi- gradually narrowed toward base. Median
costae prominent, progressively narrowed
cthreosScEopMewphaostuosgerdapahts5.kDVraowri1ngskVoftothoebtgaeinn- andelevatedtowardbase. Sulcusandcostae
obsolete in anterior 0.25 and before basal
italia were made using a Prado Universal
margin. Lateraltuberculiprominent,clearly
(Leitz Wetzlar) microprojector. All mea-
surementswere madeat 72x magnification separated by shallow, oblique lateral de-
pression. Anteriortubercleround; posterior
withan ocularmicrometeron a Leitz Wetz-
tubercle more elongate, weakly costate, and
lar dissecting microscope. Abbreviations
obsoletewell before basal margin. Granules
and explanations ofmeasurements used in
denselybutuniformlydistributedoverdor-
the text and Table 1 are listed below: sal surface, except less dense in median sul-
lOW—minimum interocular width; cus; granules oftwo or more sizes, smallest
PL—maximum length of pronotum mea- granules less than one-halfthe size oflarg-
sured along mesal plane; est. Color grey; costae, median sulcus, and
PW—maximum width ofpronotum; basal halflighter than apical half.
VOLUME NUMBER
93, 1 53
-a
3
O
CO
2 T3
O 3
^
o
Si
54 PROCEEDINGS OFTHE ENTOMOLOGICALSOCIETY OFWASHINGTON
Figs. 1. 2. Slenelmisoccidentalispronota; scalelines0.5 mm. 1,alt—anteriorlateraltubercle;pit—posterior
lateral tubercle; mc—median costae; ms—median sulcus; old—oblique lateral depression; Texas specimen. 2,
Montana specimen.
Scutellum: Sixteen granules that are merous granules ofdifferent sizes scattered
slightlylargerthan those on pronotum scat- over surface. Femur and most ofeach tibia
tered over surface; smallergranules present same color as pronotum; apex oftibia and
also. each tarsus testaceous. Indorsalview, basal
Elytra (Fig. 3): EL: 2.30 mm, EW: 1.24 halfofeach 5thtarsomerenarrowwith sides
mm. Base ofinterval 3 moderately costate subparallel, nextquarterlengthdilated, api-
for 0.20 elytral length; more costate ante- cal quarter length subparallel. Fifth tarso-
riorly but obsolete before basal margin. mere of pro- and mesotarsi slightly longer
Backgroundcolorbrown;eachelytronfaint- (0.03 mm) than their preceding four tarso-
ly vittate. Medial portion of vitta diffuses meres combined; 5th metatarsomere equal
with background color; anterior and pos- in length to its four preceding tarsomeres.
teriorportionsbrighter. Vittadoesnotover- Apicoventral margin of 5th tarsomere
lap humeral angle; confined anteriorly be- slightly convex. Conspicuous spinous ridge
tween striae 4-5; medially expanded to on distal halfofinner margin on mesotibia.
include portion ofinterval 4; includes stria Distalhalfofmetatibiawithaweakspinous
3 posteriorly; endsbeyondlateralcarinabut ridge on inner margin.
before apical margin. Interval 1, and much Genitalia(Figs. 6, 7): Peniswidestatbase,
of posterior portion on outside of lateral constrictedtowardmiddlewhereitbecomes
carina, also lighter than background color. swollen,thensubparalleltoitsroundedapex.
Venter: Apical emargination oflast ster- Each paramere with basodorsal portion
num wide and deep, but narrowerthan api- wide, elongate, nearly truncate distally, and
cal width of5th metatarsomere. longer than apicodorsal portion; inner dor-
Legs (Figs. 4, 5): Each femur with nu- sal margin slightly divergent distally, and
VOLUME NUMBER
93, 55
Fig. 3. Stenelmis occidentaliselytra; scale line 0.5
mm; c—costaeon thirdintervals; Montanaspecimen.
abruptlyangledtowardoutermarginatapex;
outer margin subparallel basally and con-
vergent apically with a shallow sinuation
before apex; inner ventral margin slightly
sinuate apically, then broadly concave to
base.
Allotype.-IOW:0.39mm,PL: 1.05 mm,
PW: 0.91 mm, EL: 2.40mm, EW: 1.25 mm.
Essentially similar, but larger. Sides ofpro-
notum more sinuate at base, more conver-
gent at apex. Color of interval 1 on elytra
not noticeably lighter than background col-
or. Fifth tarsomere of pro- and mesotarsi
slightlylonger(0.04mm)thanprecedingfour
tarsomerescombined. Without sexualchar- Figs. 4, 5. Sienelnus occidentalis meso- and me-
acters (spinous ridge) on meso- and meta- tatibiae ofmale; scale lines 0.1 mm, insert scale lines
0.5 mm; Montana specimen. 4, mesotibia with con-
tibiae.
spicuous spinous ridge on inner margin. 5, metatibia
Variations.—Variation within the type with weak spinous ridge on inner margin.
series is minimal. The antennae can be
slightly shorter than the pronotum. Some
specimens are essentially and faintly bi-
56 PROCEEDINGS OFTHE ENTOMOLOGICALSOCIETY OFWASHINGTON
swellingofthe penis is reduced and not dis-
tinctly evident in some.
Ingeneral, northern specimensare larger.
We found a male and female from South
Dakota and a female from Utah with PE
lengths more than four millimeters; these
are the largest Stenelmis recorded from
North America. In northern specimens the
antennae are distinctly shorter than the
comparatively longer pronotum. The an-
terolateral pronotal margin can be sinuate.
The pronotal and elytral sculpturing are
more pronounced. Up to 27 large granules
occuronthescutellarsurface. Theelytraare
maculatetovittateandthetestaceousmark-
ings are usually wider and cover as much
as a part ofinterval 3 to stria 5 anteriorly,
andstriae 2-5 posteriorly. Whenvittate,the
Figs. 6, 7. Stenelmis occidentalis male genitalia; medial portion variesconsiderablyin width,
dorsal;parameresspreadwiderthannatural; scaleline from narrow (striae 4-5) to wide (interval
0.25 mm. 6, Mexico specimen. 7, Montanaspecimen. 3 to stria 5), and it is usually fainter than
other portions, while it nearly diffuses with
maculate, with narrow anterior and poste- the background color in some beetles. Pos-
rior maculae confined to interval 5, and terior area on the outside ofthe lateral ca-
interval 3 to stria 5, respectively. Themeso- rinacanalso be lighterthanthebackground
and metatibial spinous ridges on males can color. Lengths ofthe 5th tarsomere on the
be slightly less or more conspicuous. pro-and mesotarsi offemalesare0.03-0.06
mm
Variation of specimens from the south- longer than the preceding four tarso-
mm
easternportionoftherangeissimilartothat meres combined, but are equal to 0.05
found within the type series. Pronotal and longer in males. The 5th metatarsomere is
mm mm
elytral sculpturing can be less pronounced. 0.01 shorter to 0.01 longer in fe-
mm
As few as nine large granules occur on the males, and 0.04 shorter to equal in
scutellarsurface. Elytralcolorpatternvaries males. The metatibial spinous ridge on
from nearly immaculate, narrowly (interval males, although low, is distinct on most in-
5) to broadly (interval 4 to stria 5) maculate dividuals. Besides being larger, the penis of
orvittate. Backgroundcolorofelytraislight most individuals is even more swollen me-
to dark brown to black; diffusion ofthe tes- dially, and the outer dorsal margin of the
taceousmarkingswiththelightbrownback- parameres can be more deeply sinuate be-
ground color is common. In males, the 5th fore its apex.
tarsomere ofthe pro- and mesotarsi can be Diagnosis.—This species cannot be in-
even longer (0.04-0.06 mm) than the pre- serted into the adult keys of Sanderson
ceding four tarsomeres combined. In both (1938) or Brown (1972) because the distin-
sexes, the 5th metatarsomere can vary from guishing characters are not in accordance
mm mm
being 0.03 shorter to 0.03 longer with the key's structure. Rearranging the
than the preceding four tarsomeres com- keys to accommodate adults of5. occiden-
bined. The weak metatibial spinous ridge talis would not be prudent at this time be-
on males is inconspicuous to distinct. Male causeseveralnew, undescribedspeciesmust
genitahavary slightlyin sizeandthe medial alsobeincluded,andareorganizationofthe
VOLUME NUMBER
93, 1 57
keybased, in part, on different characters is seum, University ofKansas, 2 Instituto de
required. Insteadofprovidingakey, wedis- Biologia, Universidad Nacional Autonoma
tinguishadultsofS. occidentalisfromaduhs de Mexico, Mexico, D.F., 2 HPB.
ofsimilar species in the discussion below. Additionalspecimensexamined.—Anad-
Although individualsin itsnorthern range ditional 2626 specimens were examined.
make this species the largest Stenelmis in Following each collection record, the num-
North America, some specimens from bers of individuals are in parentheses and
southern Texas are relatively small and immediately precede depository institu-
could easily be confused with specimens of tions or private collections, which are ab-
other species. Adults ofS. occidentalis are breviated as in Methods section. Arkansas.
most similar to those of S. decorata San- Scott Co. Mill Cr., 5 mi E Y City, 9 Jul
derson and S. n. sp. (identified as S. bicari- 1965, J. L. Donahue (1 KLS). Washington
nata in Barr and Chapin 1988), but can be Co. 24 Jul 1940 (1 INHS). Arizona. Yuma,
separated by comparing measurements and HornColl H3951 (1 MCZC). Yuma,
ratios ofthe head, pronotum, and elytra as HomColl H (1 MCZC). Colorado.Mesa Co.
listed in Table 1. In addition, the more pro- CameoPowerPlant, 8Jul 1977, A. A. Bork-
UMRM;
nounced pronotal sculpturing, the elongate si(2 1 KLS).Louisiana. GrantPar.
but not distinctly costate basolateral tuber- Stuart Lk. Rec. Area, 22-29 Jul 1982, light
cle, and the dense but uniform pronotal trap, EGR (2 LSUC). Montana. Tongue R.,
granulation, with different-sized granules, 21 Sep 1978, R. Oswald (26 HPB). Bighorn
aredistinctivecharacters. Theelytral mark- Co. Busby, RosebudCr., 29Aug 1989, DLG
ings of most specimens do not contrast as (34 DLG; 16 KLS). Carter Co. Alzada, Lit-
sharply with the background color as they tleMissouriR., 30Aug 1989, DLG(1 KLS).
do in individualsofmost species. Thecom- Chouteau Co. Ft. Benton, Missouri R., R.
bination of the 5th tarsomere on the pro- Oswald, summer 1978 (2 HPB). Golden
andmesotarsi mostcommonlybeinglonger Valley Co. Ryegate, Musselshell R., 1 Sep
than their preceding four tarsomeres, while 1989,' DLG (18 DLG; 15 KLS). Musselshell
it is shorter or subequal on the metatarsi, Co. Melstone, Musselshell R., 1 Sep 1989,
alongwith thelarge mesotibial spinousridge DLG (2 KLS). Rosebud Co. Ashland,
and smaller metatibial ridge in males, is di- Tongue R., 30 Aug 1989, DLG (19 DLG;
agnostic. The male genitalia, without a lat- 13 KLS). Bimey, Tongue R., 1975, J. A.
eral flange on the penis, is very similar to Gore (7 KLS). Mouth of Rosebud Cr., 31
the genitalia ofmost species in Sanderson's Aug 1989, DLG (43 DLG; 28 KLS). Rose-
(1938) crenata group, but especially to S. bud Cr., 10 Jun 1976, S. Baril (51 HPB).
decoratainhissinuata/humerosagroup. The Nebraska. Dawes Co. White R., 8 mi NE
constrictedandthenswollen medialareaon Chadron, 17 Jun 1984, W. P. McCafferty,
the penis is unique, and when combined A. V. Provonsha(6 PERC; 6 KLS). Nevada.
with its rounded apex and the characteris- ClarkCo. Moapa, Muddy R., 12 Dec 1986,
tics ofthe parameres, males are easily iden- WDS (2 KLS; 4 WDS), Muddy R. & Warm
NW
tified. Springsoutflow stream, 6.5 mi Moapa,
Type data.—Holotype, allotype, 33 para- 12 Dec 1986, CBB (2 CBB). Lincoln Co.
types: TEXAS: Bastrop Co., Bastrop State Ash Springs outflow streams, U.S. Hwy 93,
Park, 24 June 1963, at light, S. G. Wellso. 12 Dec 1986, CBB (7 LSUC; 16 CBB; 9
Holotype and allotype are on permanent KLS; 10WDS). Oklahoma. CaddoCo. Ana-
loantotheNMNH from TAMU. Paratypes darko, Washita R., 20 Jul 1963, HPB (7
will be deposited in the following collec- HPB). Verden, Washita R., 31 May 1971,
tions: 15 TAMU, 4 NMNH, 4 CASC, 2 HPB (45 HPB). Cleveland Co. Norman, 25
CNCI, 2 INHS, 2 SnowEntomological Mu- Aug 1975, 3 Jun-13 Aug 1976, light trap.
58 PROCEEDINGSOFTHE ENTOMOLOGICALSOCIETY OF WASHINGTON
WDS (3 WDS). Comanche Co. East Cache 10 May 1964, H. R. Burke (1 TAMU), 5-
Cr., 1 Jul 1963, HPWB (36 HPB). Custer Co. 10 Aug 1968, 1 Jun 1970, V. V. Board (30
Washita R., 2 mi Clinton, 5 Nov 1977, TAMU), 1966, M. H. Sweet (4 TAMU), 21
WDS(5 WDS), 8 miECordell, 5 Nov 1977, Aug 1972, lighttrap, W. E. Clark(1 TAMU),
WDS (6 WDS). Garvin Co. Washita R. N 17Jun 1989, EGR(4TAMU). LittleBrazos
Wynnewood, 1 Sep 1967, HPB (21 HPB). R., Hwy 21, 3 Jul 1970, V. Board & R.
Grady Co. Washita R., 6 Oct 1963, HPB (1 Phelps(18 TAMU). CaldwellCo. Plum Cr.,
HPB). Johnston Co. Tishomingo, Penning- U.S. Hwy 183, 1 1 Sep 1987, CBB (3 LSUC;
tonCr., 2W5 Jul 1972, HPB(1 HPB). Latimer 4CBB). CokeCo. NE Bronte, 29 May 1969,
Co. 5 mi RedOak,2Jul 1977,K. Stephan HPB(25 HPB). ColoradoCo. Columbus(29
(1 HPB). Love Co. Mud Cr., 5 JuWl 1963, NMNH). Coleman Co. Coleman, 29 May
HPB (1 HPB). Major Co. creek Cleo 1969, HPB (1 HPB). ComalCo. Guadalupe
Springs, 1800 ft elev., 5 Sep 1971, HPB (10 R., 1.8 mi E Spring Branch, Rte 33, 16 Dec
HPB). McClain Co. S. Canadian R., New- 1989, R. S. Zack (15 WSUC). Guadalupe
castle, 18 Aug 1976, WDS (2 WDS). R. St. Pk., 1 1 Sep 1987, light trap, CBB (3
McCurtain Co. GloverR., 25 Jul 1972, HPB LSUC; 13 CBB; 1 KLS). Wimberly, Blanco
(1 HPB). Stephens Co. Wild Horse Cr., 26 R. 25 May 1969, HPB (1 HPB). Comanche
Jul 1962, HPB (34 HPB). Washita Co. Cor- Co. De Leon, Sabanna R., 29 May 1969,
dell, 4 Jul 1969, HPB (30 HPB). Oregon. HPB (8 HPB). Concho Co. Concho R. S
MalheurCo. Ontario, Exp. Sta., 2Aug 1960, Lowake, 23 Jul 1982, JRD (8 JRD; 4 KLS).
light trap, F. P. Larson (2 ODAC; 2 HPB). Cooke Co. Gainesville, Elm FTc. Trinity R.,
South Dakota. FallRiver Co. Cheyenne R., 21 Jun 1978, WDS (1 WDS). Coryell Co.
U.S. Hwy 38, 11 Jun 1975, W. P. Mc- Gatesville, 20 Aug 1970, HPB (8 HPB).
Cafferty, A. V. Provonsha, B. L. Heath (4 Crosby Co. Crosbyton, 22 Apr 1972, HPB
PERC; 3 KLS). Texas.Anderson Co. 10 mi (1 HPB). Denton Co. Argyle, Denton Cr.,
SW Elkhart, 23 Jul 1960, light trap, H. R. 28 Apr 1939, J. A. & H. H. Ross ( 8 INHS).
Burke (3 TAMU). Salmon, 27 Jun 1975, Denton Cr., 12 mi S Denton, 21 Jun 1978
light trap, H. R. Burke (4 TAMU). Trinity (3 WDS). 3 mi S Denton, U.S. Hwy 35, 22
R., U.S. Hwy 287 nr Cayuga, 6 Oct 1987, Jul 1975 (12 WDS). Elm Fk., 15 Nov 1970,
JRD (2 JRD). Bastrop Co. Bastrop St. Pk., D. McNeely (7 WDS). DeWitt Co. Coleto
24 Apr 1962, light trap, S. G. Wellso (10 Cr. nrWeser, 28 Mar 1974, HPB (29 HPB).
CASC; 10 TAMU; 6 INHS), 13 Jun 1969, NEWesthoff, 28 Mar 1974, HPB(77 HPB).
S. Peck (5 INHS). Bee Co. Aransas R. E Brook S Westhoff, 28 Mar 1974, HPB (9
Papalote, 29Apr 1982,JRD(3JRD;2KLS). HPB). Duval Co. Sepulveda Ranch, 4 Jul
Bexar Co. San Antonio, 2 Jun 1958, J. F. 1970, C. W. Griffen (1 NMNH). Ellis Co.
Lawrence(1 HPB; 2 MCZC), 4Jun 1975, M. Waxahachie, 31 Aug 1967, HPB (7 HPB).
Druckenbrod(3 NMNH). BlancoCo. John- FortBendCo. Missouri City, 31 May 1977,
son City, 5 Oct 1966, HPB (2 HPB). John- lighttrap, EGR& LeDoux(8 EGR; 2 KLS).
son City, Pedemales R., 27 Aug 1967, HPB Frio Co. Pearsall, 26 Jun 1970, J. W. Smith
(1996H7P,B)1.9 BAousgqu1e9C7o0.(113miTANMUCl)i.fto2n,m6iJWul (m1iTSAFMrUed)e.ricGkislbleusrpgi,e2C3o.JuPned1e9m78a,leWsDRS.,(44
Iredell, 1 Jul 1966 (2 TAMU). Morgan, WDS). GoliadCo. Goliad, San Antonio R.,
SteeleCr., 23 Dec 1974, 30 Dec 1976, WDS 23 Jul 1962, H. R. Burke (1 TAMU). Small
(4WDS). BrazosCo. Bryan, 16-21 Jul 1965, creekEGoliad, 29 Mar 1974, HPB(2 HPB).
28 May-12Jul 1966, lighttrap, J. C. Schaff- Gonzales Co. Gonzales, Guadalupe R., 28
ner (17 TAMU), 25 Jun-8 Aug 1988, EGR Mar 1974, HPB(57 HPB). PalmettoSt. Pk.,
(7 TAMU; 4 EGR). Cedar Cr., Oct 1971, 7 Jun 1969, Board & Hafemik(61 TAMU),
R. G. Phelps (1 TAMU). College Station, 14Jun 1969, M. W. Sanderson(125 INHS),
2
VOLUME NUMBER
93, 1 59
30 May 1977, light trap, EGR (6 LSUC; 20 25 May 1969, HPB (4 HPB). Menard Co.
NMNH; 50 UMRM; 39 HPB; 1 1 EGR; 25 Menard,May 1946, L.J. Bottimer(6CNCI).
KLS), 30 May 1983, 27 Apr 1988, CBB (3 Montgomery Co. Conroe, W. Flc. San Ja-
LSUC; 5 CBB), San Marcos R., 31 May cinto R., 25 Jun 1967, HPB (2 HPB). The
1983, CBB (2 CBB). Grimes Co. Navasota, Woodlands, 1-10 Jul 1978, R. Tumbow (2
Navasota R., 31 Aug 1967, HPB (9 HPB). UGAM), 24-30 Jun 1979, J. E. Wappes (1
NavasotaR.,Hwy 158,15Aug 1962, Reeves TAMU). Nacogdoches Co. Douglas, Ange-
& Burke (1 TAMU). Guadalupe Co. Shertz, lina R., 10 Aug 1969, HPB (1 HPB). Nolan
25 Jun 1985, light trap, D. W. Sundberg (2 Co. Sweetwater, 10 Jul 1937 (1 TAMU).
LSUC; 12 CBB). Hamilton Co. Hico, N. Nueces Co. Corpus Christi, 26 Jun, 1 Jul
Bosque R., 22 Jun 1978, WDS (45 WDS). 1970, light trap, C. W. Griffen (6 TAMU).
Leon R., 6 mi N Hamilton, 22 Jun 1978, Palo Pinto Co. Brazos R., Hwy 4, 30 May
WDS(133 WDS).HarrisCo. Houston, Buf- 1973, B. Stark (2 HPB), 29 Jul 1982, WDS
W
falo Bayou, 16 Jul 1985, JRD (6 JRD; 2 (3 WDS), 4 mi Mineral Wells, 27 Jun
KLS). Seabrook, 19 Aug 1970, light trap, 1978, WDS (63 WDS). Dark Valley, 29 Jul
WDS (1 WDS). Havs Co. Buda, Onion Cr., 1970, D. McNeely (2 WDS). 4-5 mi SW
17 Sep 1960, H. Chambers (8 TAMU). San Graford, 25 Aug 1981, WDS (53 WDS; 15
Marcos, San MarcosR., 30 Sep 1964, 9 Aug KLS). Parker Co. Dennis, Brazos R., 24-25
1969, HPB (4 HPB). San Marcos R. 12 Jul Jun 1989, EGR (1 TAMU). Robertson Co.
1967, light trap, G. S. Tulloch (1 TAMU), CampCr., 1 Jun 1962, lighttrap, V. House
27 Aug 1974, J. Kolb (2 NMNH). Hidalgo (1 TAMU). Runnels Co. Ballinger, Colo-
Co. Bentsen-Rio Grande St. Pk., 18 Jun rado R. Rte 83, 19 Dec 1989, R. S. Zack (5
W
1969, Board & Hafemik (1 TAMU), 9-10 WSUC). San Patricio Co. 6 mi Edroy, 1
Jun 1975, light trap, R. Tumbow (6 HPB; Jul 1970, C. W. Griffen (4 TAMU). Lake
8 UGAM), 17-18 May 1979, 4 Oct 1986, Corpus Christi St. Pk., 9 Jun 1969, Board
15-16 Oct 1988, EGR(2NMNH; 3 TAMU; & Hafemik (5 TWAMU). San Saba Co. San
2 EGR). SantaAnaNat. Wldlf Ref 23 Oct Saba R., 5 mi San Saba, 22 Jun 1978,
,
1987 (2 EGR). Hill Co. Blum, Nolan Cr., WDS (2 WDS). Somervell Co. Glen Rose,
23 Dec 1974, WDS (1 WDS). Kaufman Co. Paluxy R., 23 Dec 1974, WDS (8 WDS).
TrinityR., Hwy 34 nrRosser, 26 Aug 1987, Tarrent Co. Benbrook, Trinity R., 27 May
1 Oct 1988,JRD(20JRD; 7 KLS). Kendall 1971, 13 Jun 1974, WDS (50 WDS). 5 mi
Co. Boeme, Cascade Caverns, 20 Jul 1981, S Benbrook, 21 Jun 1978, WDS (7 WDS).
W. E. Steiner (2 NMNH). Mission, 17 Jul Tom Green Co. Concho R., 14 mi E San
1981, light trap, W. E. Steiner (5 NMNH). Angelo, 12 Jun 1961, H. R. Burke (2
Lavaca Co. Navidad R., 10 & 12 mi N Hal- TAMU). Travis Co. Austin, Onion Cr., 19
lettsville, 23 Apr 1961, 28 Jul 1961, H. R. Aug 1970, 4 Jul 1978, HPB (5 HPB). Vic-
Burke (2 TAMU). Limestone Co. Groes- toria Co. Raisin, Coleto Cr., 29 Mar 1974,
W
beck, Navasota R., 18 Oct 1969, HPB (2 HPB (5 HPB). sandy creek Inez, 29 Mar
HPB). LiveOak Co. 12 mi N Mathis, 7 Aug 1974, HPB (12 HPB). San Antonio R., 16
1971, light trap, W. H. Tyson (1 NMNH). mi S Victoria, 20 Aug 1962, H. R. Burke
Whitsett, Atascosa R., 25 Aug 1962,J. Haf- (4 TAMU). Victoria, 26 Jun 1965, P. J.
emik (1 TAMU). Llano Co. Castell, Llano Spangler (3 NMNH). Washington Co.
R., 1 Oct 1960, H. R. Burke (2 TAMU). Washington-on-the-Brazos St. Pk., 12 Jun
McCulloch Co. San Saba R., 16 mi S Brady, 1969, M. W. Sanderson (21 INHS). Whar-
13 Jun 1963, D. C. & K. A. Rentz (363 ton Co. Wharton, 19 Aug 1970, light trap,
CASC; 26 KLS). McLennan Co. Bosque R. WDS(1 WDS). WilliamsonCo. SanGabriel
nr Waco, 14 May 1961, B. McDaniel (2 R., 6 mi E Georgetown, 1 Oct 1960, H. R.
TAMU). Medina Co. Hondo, Hondo Cr., Burke (5 TAMU), Taylor, 16 Jun 1967, J.
60 PROCEEDINGSOFTHE ENTOMOLOGICAL SOCIETY OFWASHINGTON
Fig. 8. Distribution ofStenelmis occidentalis. Star represents type locality. Arizona record is questionable;
see section on distribution.
E. Hafernik (2 TAMU). Wilson Co. Cibolo F. Knowlton (1 HPB). Mexico. Coahuila
Cr., 8 May 1980, JRD (34 HPB). Zapata Rio San Rodrigo, E El Remolino, 26-27
Co. Falcon St. Pk., 1 1 Jun 1975, light trap, May 1969, HPB (3 HPB). Villa Fuente, 26
R. Tumbow (7 UGAM). Falcon St. Rec. May 1969, 19 Aug 1970, HPB (27 HPB).
Area, 5 Jun 1983, light trap, CBB & D. A. Zaragoza, 26 May 1969, HPB (14 HPB).
Rider (25 LSUC; 47 CBB; 16 KLS). Utah. Distribution(Fig. S).—Stenelmisocciden-
Box Elder Co. 5 Oct 1971, D. W. Shaw (1 talis is the most common and widespread
HPB), 5 Jul 1972, R. Newell (2 HPB). Deep species ofStenelmis restricted to areas west
Cr., 3Jan 1971, D. W. Shaw(2 HPB). Field- ofthe Mississippi River, and one ofonly a
ing, Malad R., 14 Aug 1954, A. Gaufin (4 few found in Mexico. It is also the only
HPB). Cache Co. Logan, 24 Jul 1974, G. F. species known to occur in the northwestern
Knowlton (1 USUC). Duchesne Co. Ft. Du- U.S. (MT, OR, UT), an area formerly be-
chesne, Uinta R., 31 Mar 1973, 5 Oct 1974, lieved to be devoid ofthe genus. Although
B. P. Stark (14 HPB). Grand Co. Arches there are large areas in its range from which
Nat. Mon., Moab, 12 Jul 1964, J. T. Doyen we have not seen specimens, we suspect a
(1 HPB). Uintah Co. Uinta R., Hwy 40, 23 lackofcomprehensivecollectingisthecause
May 1974, B. P. Stark (2 HPB). Ashley Cr. ofitsapparentabsence. Twospecimensfrom
nr Vernal, 1 Apr 1973, 5350 ft, B. P. Stark Yuma, Arizona were found in the Hom-
(7 HPB). Wayne Co. 14 mi S Hanksville, LeConte Collection, MCZC. Since these
25 Jul 1968, at light, H. F. Howden (1 venerable beetles have never been men-
CNCI). Weber Co. Ogden, 10 Aug 1960, G. tioned in the literature, and we are unaware