A new species of Heisteria (Olacaceae) from Mesoamerica

5 Pages·2000·2.8 MB·English

by  Quírico Jiménez

1
Bull.nat.Hist.Mas. Lond. (Bot.)30(1): 1-5 Issued29June2000
A
new species ofHeisteria (Olacaceae) from
Mesoamerica
IWc r««i»aAKT^Tae.UH-^-ArwLwaie^i*™
QUIRICOJIMENEZ
InstitutoNacionaldeBiodiversidad,Apdo. 22-3100, SantoDomingodeHeredia, CostaRica - JUL 2000
SANDRA KNAPP
DepartmentofBotany, TheNaturalHistoryMuseum, CromwellRoad, LondonSW75BD
Synopsis. A new species ofOlacaceae from Mesoamerica, Heisteriapovedae, is described and illustrated. Its range of
variationisdescribedandrelationshipstootherMesoamericanmembersofthegenusbrieflydiscussed.
While preparing treatments of the family Olacaceae for Flora {Standley 68338), 'palo de baston' {Steyermark 33360); El Salva-
MesoamericanaandtheManualdeplantasdeCostaRica,itbecame dor,Ahuachapan: 'ojodepava' {Sandoval& Chinchilla281).
clearthatthewidespread species treatedas Heisteria macrophylla Etymology. ThisspeciesisdedicatedtotheCostaRicanbotanist
Oerst. inSleumer's (1984)FloraNeotropicaaccountofthefamily LuisJorgePoveda, inrecognitionofhis greatknowledgeofCosta
includedatleasttwoelements. ThetypeofH. macrophyllacomes Ricanplants,andinhonourofhismanycontributionstotheknow-
fromtheareaalongtheRioSanJuanneartheCostaRica/Nicaragua ledgeandconservationofplantsinMesoamerica.
border,andmaterialfromthelowlandAtlanticslopeforestsmatches
thetypequitewell(seebelow).Themajorityofspecimensidentified Distribution. Heisteria povedae is endemic to Mesoamerica,
as H. macrophylla are significantly different from the type, and occurringfromChiapastoPanama. Itisgenerally foundatmiddle
despite being the most widespread component ofH. macrophylla elevations, (200-)800-1500(-2000) m, in lowland forest,
sensuSleumer,donothaveaname. Wethereforecoinanewname premontaneforestandoccasionallyintheoakforestsoftheCordillera
forthis widespread,common specieshere. deTalamancainCostaRica.Heisteriapovedaeismorecommonon
thePacificslope,butsomehavebeencollectedontheAtlanticslope.
Heisteria povedae Q. Jimenez & S. Knapp, sp. nov. Type: Costa (Fig. 2).
Rica,Guanacaste,EstacionCacao,ParqueNacionalGuanacaste,
Specimensexamined.
11(X)m, 10°55'32"N, 85°28'02"W, 8February 1995,A. Azofeifa
25 (INB!-holotype; BM!,CR!,F!,MO!-isotypes). MEXICO.Chiapas: Palomeque,Mt.Ovando, 1400m, 16February 1969,
HernandezM., R. 528 (MO, NY); VolcanTacana, Union, 14(K) m, 17-23
Fig. 1. March 1939, Matuda, E. 2787 (MEXU); LaGrada, Escuintla, 12 August
Frutices vel arbores parvae; ramuli valde alati albo-punctati vel 1947,Matuda,E. 16807(F);Cacaluta,Escuintla, 15August 1947,Matuda,
striati,alisincaulibusmaturioribuspersistenibus;fohainstatusicco E. 16875 (F); Corcega, Pueblo Nuevo Com., 15 April 1948, Matuda, E.
viridi-olivaceapunctulata;fructusnigerglobosinitidileviterporcati; 1178675586((FF);MLEasXUNu)v;esA,rMrto.yoOvdaendlao,Cu1c1hi5l0lma,,a1r5riNboavdeembFeinrc1a9P4r9u,sMiaat(udSaW,dE.e
calyxfructiferpatenterubermargine sinuatus; pedunculi fructiferi Jaltenango), 1500 m, 22 February 1951, Miranda, E 6965 (US); Mun.
crassi. Acacoyagua,Mt.Ovando,trailtoc.5kmaboveFincaLaMagnolia,3kmN.
Shrubsorsmalltrees,1-5m;barkpale.Branchletsstronglyangular, omf,L2o9sMCaacyao19a8n5d,1T3hkommaNs.,oWf.Ac&acVoilylaagseunaoro,nJr.oLad3t6o6O3lo(nMdrEiXnUa,,1N2(YX)>;-1n5e(aXr)
markedlyflattenedand strongly 2-winged, occasionally4-winged,
RanchoofFincaPrusia,Mun.AngelAlbinoCorzo,24(X)ft,23January1968,
the wings persisting to older stems, pale olive green, often white Ton,AlushShilom3566(F,MEXU,MO);alongtheRioCuztepequesnear
striped. Leaves 6-20 x 2.5-6.5 cm, eUiptic to narrowly elliptic, Finca Cuztepeques, 2400 ft, 26 March 1968, Ton, Alush Shilom 3865 (F,
shapeveryvariableevenwithinasinglecollection,usuallydryinga MEXU,MO);MonteOvando,1950m,9January1987,VazquezT,M.3899
verypaleolivegreen,stronglypelluciddottedbelow,matteonboth (NY).
surfaces; venation drying bright pale yellow or white, all orders GUATEMALA.Chimaltenango:lowerandmiddleSWslopesofVolcdn
clearly visible; apex acuminate, occasionally acute; margins revo- Fuego,aboveFincaMontevideo,alongBarrancoEspinazoandtributaryof
clumt,e,rsulgioghstelyanudndpuallaeteg,repaelnera;lobnagseenatciurteelteongctuhn.eaItnef;lopreetsicoelensce0.a5f-e1w.- R(Fi,oNPYa)n.taElsecouni,nt1l2a0:0-F1in6c0a0Mmo,nt2e0rrSeeyp,teS.mbselrop1e94o2f,VSotlecye^rmdaerkF,ueJg.oA.,512104807-
flowered glomerule of 5-7 flowers; bracts glabrous, pale green; 1260m,5February 1939,Standley,P.C.64570(F).Quezaltenango: Finca
Pireneos, below Santa Maria de Jesus, 1350-1380 m, 11 March 1939,
pedicels 1-2 mm. Flowers white, the buds ellipsoid; calyx Standley, P.C. 68199, 68338, 68396 (F); along old road between Finca
cupuliform, the lobes c. 0.25-0.5 mm, either mere teeth from the Pireneos&Patzulin, 200-1400m,9February 1941,Standley, P.C. 86950,
calyxrimormoreorlessequallingthetube;petalsc.2mm,glabrous 86959,86992(F);alongQuebradaSanGeronimo,FincaPireneos,lowerS.-
within;filaments 1-1.5mm,liguliformtopetaloid.Fruit7-9x7-10 facing slopes of Volc^ Santa Maria, between Santa Maria de Jesus &
mm,oftenwiderthanlong,globose,stronglyridgedandsomewhat Calahuache, 1300-2000m, 1-2January 1940,Steyermark, J.A. 33360(F,
shiny,black;fruitingcalyx1.9-2.5cm,unlobed,themarginssinuate, NY);offHwy.95,c.4milesS.ofSantaMariadeJesus,W.sideofVolcanSta.
spreading or slightly cupping the base of the fruit, fleshy, red; Maria,29June 1973,Stone,D.E.3492(F,MEXU,MO,NY);FincaSt.John
peduncles5-9mm,verythick. ofMrWhitehead,c. 5kmS.ofSantaMariadeJesus,SEslopesofVolc^
Santa Maria, 4300-5500ft, 17 May 1966, Walker, J.W. 443 (F, NY, US).
Commonnames. Guatemala,Quezaltenango: 'aritodemontaiia' SanMarcos: LaTrinidadc. 2kmfromFincaArmeniaaboveSanRafael,
©TheNaturalHistoryMuseum,2000
SQ-bO-VD-- lOOl
Q.JIMENEZANDS. KNAPP
A
L~^
'/K-
5cm
«.TeU>^
Fig,1 Heisteriapovedae(habitbasedon/INBio 149(MO)&Fuentesetal.67(MO);fruitbasedonDavidseetal.28398(MO)).
1100-1250 m, 12 July 1977, Croat, T.B. 40873 (MO); above Finca El EL SALVADOR. Ahuachapan: Mun. San Fco. Menendez, Hacienda
Porvenir on Todos Santos Chiquitos, lower S.-facing slopes of Volcan SanBenito,alN.deCampana(ElImposible), 13°49'N,89°56'W,23Febru-
Tajamulco,1300-1500m,7March\9AQ,Steyermark,J.A.37080(F).Solola: ary 1982,Sandoval&Chinchilla281(MO(B,LAGU));SierradeApaneca,
cercaniasdePatulul,240m,December 1929,GarciaSalas,J. 1418(F);S.- inregionofFincaColima,17-19January\922,Standley,P.C.20185(F,MO,
facingslopesofVolcanAtitlan,aboveFincaMoca, 1000-1250m,20June US).
1942, Steyermark, J.A. 47903 (F). Suchitepequez: SW lower slopes of NICARAGUA.Boaco:CerroMombachito,4kmNEdeCamoapa,700-
Volcan Zunil, between Finca Montecristo & Finca Asturias, SE ofSanta 1000m, 1 February 1979,Grijalva,A. &Araquistan,M.60(MEXU,MO);
MariadeJesus,1200-1300m,1February1940,Steyermark,J.A.35281(F); MonteAzul,SanJos^delosRemates, 1000-1200m, 12°38'N,85°43'W, 12
FincaMoca,3000--U00ft, 10January 1935,Skutch,A.F.2109(BM,F,NY, February1983,More«o,f:P.20309(MO);NEdeMombachito,11May1982,
US). Zacatepequez: Acatepeque,4300ft, March 1892, Donnell-Smith, J. Sandino,J.C.2829(MO);upperSWslopeofCerroMombachito,S.ofroad
2479(US);VolcdndeAgua,9000ft,November 1889,Heyde&Lux(JDS) betweenBoacoandCamoapa,900-1000m,c. 12°24'N,85°33'W,3October
4446(MO,NY). 1979,Stevens, W.D.etal. 14589(MEXU,MO,NY);upperW.slopeofCerro
NEWSPECIESOFHEISTERIA
Q.JIMENEZANDS. KNAPP
*Utley3218(F).Cartago:vicinityofPejivalle,600-850m,January 1940, watershed, 1400m, 10°18'N,84°48'W, 18-21 March 1973,Burger. W.C. &
Skutch, A.F.4581 (F, MO, NY, US). Guanacaste: Estaci6nCacao, Parque Gentry,J.L. 8765(F);E.ofLasCrucesand5-6kmS.ofSanVito,property
Nacional Guanacaste, 1100 m, 10°55'32"N, 85°28'02"W, 2 June 1990, ofMrRobertWilson, 1100-1200m,8°47'N,82°58'W, 15-16January 1967,
Acevedo,D.5,Alvarado, C. 15(CR);Tilar^,March 1942,Acosta,M.A. 14 Burger, W.C. &MattaU.,G.4464(BM,F,MO);E.ofQuepos,Pacificslope
(CR); Estaci6nCacao, ParqueNacionalGuanacaste, 1100m, 10°55'32"N, ofTalamancas, 150-250 m, 9°29'N, 84°03'W, 19 February 1977, Burger,
85°28'02"W,23September 1994,Alverson. W. & Christy, CM. 2769(CR); W.C.etal. 10607(F);justE.ofMonteverdeonPacificwatershed,1300-1450
AreadeConservaci6nGuanacaste,biologicalstationonW.flankofVolcdn m,10°18'N,84°48'W,24February1971,Burger. WC.etal. 10706, 10847(F
Cacao, 1100m,c. 10°55'32"N,85°28'02"W,23July 1994,Alverson, W.S. & MO,NY);foothillsofCordilleradeTalamanca,vicinityofHelechales,along
Flores, T.S. 3105 (CR, MO); Abangares, la Sierra y Rio Caiias, 1000 m, RioGuineal, 1100-1200m,9°04'30"N,83°05'W,29March 1984,Davidse.
10°22'00"N, 84°55'00"W,Bella, E. 4959(INB); E. slopesofVolcdnMira- G. &HerreraCh., G. 26242(F);CordilleradeTalamanca,areaaroundRi'o
NW
valles (Cerro laGiganta), abovetownofRioNaranjo,c. 800m, 10°42'N, Canasta,9.5airlinesmiles ofAguasCaliente,betweenCerroFrantzius
85°07'W, 8 April 1973, Burger, W.C. & Gentry, J.L 9143 (F); Estaci6n &CerroPittier,1500-1600m,9°02'N,82°59'W,6September\9M,Davidse,
Cacao,ParqueNacionalGuanacaste, 1100m, 10°55'32"N,85°28'02"W,29 G.etal.28398(MO);Monteverde,comunidad, 1400-1500m,28November
October1989,Chacdn,I.2520(CR);ParqueNacionalGuanacaste,Estaci6n 1976,Dryer, V. 1004(MO),7Febniary 1977,Dryer. V: 1167(F);SanLuis,
Cacao, Liberia, 1100 m, 10°55'45"N, 85°28'15"W, 18 November 1990, Monteverde,1100m,10°16'33"N,84°47'45"W,MDocQmhtr1991,Fuentes,
Chavez, C. 341 (MO);EstacidnMaritza,ParqueNacionalGuanacaste,600 Zetal.67(INB,MO);Monteverde,lowermontaneforest, 1400m, 14July
m,10°57'40'N,85°29'40'W,29August1990,Carballo,G.215(CR);Estaci6n 1990,Gentry,A.etal.71672(MO);LaTigra,LasMellizas, 1280m,August
Maritza,ParqueNacionalGuanacaste,600m, 10°57'40"N,85°29'40"W,29 1983,Gomez,L.D.etal.21971 (BM,MO);SanLuis,Monteverde, 1100m,
August 1990, Chavarria, U. 178 (CR); Estacion Cacao, Parque Nacional 10°16'33"N, 84°47'45"W, 10 September 1994, Gonzalez, J. 400 (INB);
Guanacaste, 1100m,10°55'32"N,85°28'02"W,8February1995,Chinchilla Monteverde,1300-1450m,10°18'N,84°48'W,10June1985,Grayum.M.&
33 (INB);Parque Nacional RincondelaVieja, SEslopesofVolcan Santa Hammel, B. 5389(CR); Monteverde, lowercommunity, 1350-1400m, 25
Maria,aboveEstaci6nHaciendaSantaMaria,900-1200m,10°47'N,85°18'W, Febniary 1985,Haber, W.A. 1376(MO), 29August 1985,Haber, W. etal.
27-28January 1983,Davidse, G. etal. 13369(CR,MO);EstacidnCacao, 2532 (CR), 18 December 1988, Haber, W. 8905 (INB), 20October 1990,
ParqueNacionalGuanacaste, 1100m, 10°55'32"N,85°28'02"W,9February Haber. W. 10089 (INB); Monteverde, lowercommunity,1350 m, 10°20'N,
1995,Gamboa, B. 51 (INB);ParqueNacionalRincondelaVieja,Liberia, 84°50'W, 20January 1986,Haber, W.A. & Wolf. J.A. 4284(MEXU, MO);
CordilleraGuanacaste, Estaci6n Las Pailas, sendero Rio Blanco, 1200 m, nearCerroChivo,S.ofCotonRiver, 1600-1800m, 17July 1983,Hazlett,
10°47'50"N,85°21'10"W,5November1992,Garia,D.32(MO);Rinc6nde D.L. 5240(F);roadfromPensionFlorMarchtoMonteverdeCloudForest
laViejaNational Park, PuestoSantaMaria,pathtoMirador, c. 800m, 27 Reserve, 1400-1500m, 10°18'N,84°48'W,8January 1987,Hill. S.R.etal.
January 1983,Garwood,N.etal.717(BM,F,MO,NY);ridgestoN.ofRio 17686 (NY);just E. ofMonteverde on Pacific watershed, 1300-1450 m,
LasFlores, c. 1 kmE. ofRioTenorio, haciendaMontezuma,475-500m, 10°18'N,84°48'W,21July 1977,Lawton.R.O. 1201(F);MonteverdeCloud
10°40.5'N, 85°04.5"W, 24January 1985, Grayum, M. & Herrera, G. 4854 ForestReserve,Hoge'sWood, 1320m, 18July 1979,Koptur,S. 155(MO).
(CR, MEXU); S. &E. slopesofridgeSEofQuebradaZopilote, lowerSE SanJose:CantondeAcosta,ZonaProtectoraCerrosdeEscazii,cuencadel
slopeofVolcdnSantaMaria,850-940m, 10°46.5'N, 85°18'W,25January RioTabarcia,faldaW.deCerroCedral,porsenderoaLondres, 1600-1700
1986,Grayum, M. etal.6243 (CR,NY);EstacionCacao,ParqueNacional m,9°50'52"N,84°04'40"W, 14May 1994,Morales,J.F2758(MO).
Guanacaste, 1100m, 10°55'32"N, 85°28'02"W, 12July 1996,Gonzalez, J. PANAMA.Code:footofCerroPilon,abovevalledeAnton,2000ft,28
1101 (INB); Parque Nacional Guanacaste, estacion Maritza, sendero a la March1969,Rorter,D.M.etal.4676(MO).Colon:26kmfromTransisthmica
cimadeVolcanOrosi,600m,10°57.6'N,85°29.6'W,2July19S9,lINBio142 HighwayonSantaRitaRidge,NWfacingslopes,500m,9°26'N,79°57'W,
(MO);ParqueNacionalGuanacaste,EstacionMaritza,senderoalacimadel 21 October 1981, Knapp. S. et al. 1711 (MO). Chiriqui: trail from Paso
VolcanCacao,600m,10°57.6'N,85°29.6'W,3July1989,IIINBio119(MO); AnchotoMonteLirio,uppervalleyofRioChiriquiViejo, 1500-2000m, 16
ParqueNacionalRinc6ndelaVieja,900-1200m,8May1972,Janzen.D.H. January 1939,Allen, P.H. 1589(F,MO);OjodeAgua,propertyofRatibor
401(CR);ElSilencio,Tilar^,850m,Jimenez,A.s.n.(F);Monteverde,1450 Hartmann,vicinityofSantaClara(betweenVolcan&RioSereno), 1520m,
m,7February1966,Kern,P.M.801(NY);SantaElena-Tilar^nroad,c.2km 8°51'N,82°45'W,17June1987,Croat,T.B.66286(US);roadfromVolcanto
fromSantaElena,c. 1300m, 17February 1984,Khan,R. etal. 1096(BM, RioSerano, 16kmfromRioSereno,29June 1977,Folsom,J.R4048(MO
MO);Estaci6nCacao, ParqueNacional Guanacaste, 1100m, 10°55'32"N, x2);Hartmanfinca,nearCerroPando,c.2000-2200m,8°52'N,82°45'W,22
85°28'02"W,30May-1 June 1990,Maas, P.J.M. 18(CR);VolcanTenorio, August1982,Hamilton, Cetal.816(MOx2);nearCostaRicanborder, 13
1200m,8January 1993,Quiros,M. 1076a(CR);EstacionMaritza,Parque km by road S. of Rio Sereno, Finca Hartmann, 1400-1500 m, 8°50'N,
Nacional Guanacaste, 600 m, 10°57'40"N, 85°29'40"W, 18 August 1993, 82°45'W,14May\99\,Hensold,N.&McRherson,G. 1050(MO,US);lower
Ramirez, R. 53 (INB); Parque Nacional Rinconde laVieja, senderode la slopesofCerroPelota, 1800-2000m, 8°53'N, 82°43'W, 10October 1981,
TomadeAgua,a3kmdelaEstacion, 1000m, 10°46'05"N,85°17'40"W, 17 Knapp,S. 1481 (MO);forestsandedgesofcafetalesofFincaOjodeAgua,
September1990,Rivera,G.628(MO);ParqueNacionalRincondelaVieja, 1300m, 8°5rN, 82°46'W, 14October 1981, Knapp, S. 1578 (MO); Santa
EstacionBiologicaSantaMaria,tracktoAguaFria,780m,10°46'N,85°17'W, Clararegion,27kmNWofElHatodelVolcan,oncoffeefincaofRatibor
19 June 1996, Short, M.J. & Stafford, RJ. 151 (BM, CR, INB); Parque HartmanncalledOjodeAgua,5000-5300ft,18July1975,Mori,S.&Bolten.
NacionalGuanacaste,VolcanCacao, 1000m, 30July 1987,Smith, J.F. & A. 7215 (MO). Panama: alongtrail totopofCerroCampana, 13 October
Frost,E.425(CR,F);EstacionMaritza,ParqueNacionalGuanacaste,600m, 1974,Mori, S. &Kallunki.J. 2463(MO);CerroCampana,c. 10kmSWof
10°57'40"N, 85°29'40"W, 31 August 1990, Solomon, J 19111 (INB); El Capira, on trail to summit, 870-1000 m, 7 December 1974, Mori, S. &
Silencio,nearTilar^,c. 750m, 13January 1926,Standley, RC. & Valerio, Kallunki. J. 3565 (MO). Veraguas: vicinity ofEscuela Agricultura Alto
J.44596,44602(US);QuebradaSerena,SEofTilar^,c.700m,27January Piedra,nearSantaFealongtrailtotopofCerroTute,2800-3200ft,3April
1926,StandleyP.C.&Valerio,J.46178,46196(US);NaranjosAgrios,600- 1980,Antonio,T.4003(MO);0.6milebeyondEscuelaAgn'colaAltoPiedra,
700m,29January 1926,Standley,RC.&Valerio,J.46495(US);3kmN.of 730m,4April1976,Croat,T.B.&Folsom,J.R34069(NY);aboveSantaFe
RioNaranjo,nearContinentalDivide,500m, 10°52'N,85°04'W,5January beyond Escuela Agricola Alto Piedra, 1.8 miles beyond fork in road on
719k7m5,STaWyloofr,BrJi.b1ri8,101300(-F)2;5T0ilma,r^4n,Ma20yJ1u9l8y31,9G2o0,meVzal,erLi.oD,.Je.t1a8l.(2U0S4).30Li(mMoOn):; CPaecrirfoicTustleo,peW,.siodfeSoafntCaerFer,obTeutyeo,nd5AAlptroilde19P7i6e,draC,ro6a0t,0-T8.0B.03m4,1c9.98°(3M0O')N;,
vicinityofGuapiles,300-500m,12-13March1924,StandleyRC.37331(F, 81°07'W, 18October1985,McRherson,G.7188(MEXU);NWofSantaFe,
4US6)1;36(mNiYl)e.sPinulnatnadrfernoams:moCuetrhroofPEasntdroe,llariRdigveesr,a1b9ovAeprRilio19C5o2t,6Sntoarkn,dHR.Ei.o 119k7m5,fMroormi.EsSc.u&elaKaAlglruinckoi,laJ.Al4t7o7d8e(PMieOd,raN,Ys)l;opaelsoonfgCsetrereopTturatiel,t2o4sFuembmriutaroyf
MN&eagGyrdom1,e9z11,10,0L0B.u-Dr1.c8h10,601Dm9.,(48F6)°;1556F'i(NnF,c)a;82jLu°as4st5'EC.rWu,ocfe1sM9,-o2nS1taenFveeVbirrtduoeadroeynJ\Pa9avSca2i,,fciB.car1wr3ait0ne0gremsrh,,e2dK3., C3e2r0r0ofTt,u4teJca.n3uakrmya1b98o1v,eSEyssctumeal,aKA.gr&icAonltaonAilot,oPT.ie3d0r3a3ne(aMrOS)a.ntaFe,2800-
1300-1450m, 10°18'N, 84°48'W, 29 October-2 November 1975, Burger, Specimens here described as Heisteria povedae were called H.
W.C.&Baker,R.9599,9619(F,MO);c.2kmSEofMonteverde,onPacific acuminata(Humb.&Bonpl.)Engl,byBurger(1983).Herecognized
NEWSPECIESOFHEISTERIA
them as different to H. macrophylla Oerst. (s.s.), differentiating Heisteriaacuminata,theotherwidespreadspeciesoiHeisteriain
thembyhabitat: withplantsofH. macrophyllas.s. growinginwet Mesoamerica,especiallyinPanama,differsfromH.povedaeinits
lowlandforests andthoseofH. acuminata(sensuBurger)growing ellipsoidfruit,andlonger,thinnerfruitingpedicels.Theleavesof//.
indriermoreuplandformations. Sleumer(1984)lumpedthesetwo acuminata generallydry adarkergreenishbrownthan thoseofH.
taxa, stating however 'Specimens with leaves smaller than usual povedae, and the branchlets are not markedly winged. Heisteria
occurinPanamaexclusivelyonitsPacificside'.Thepastdecadesof acuminataoccursinlowlandforestsfromtheOsapeninsulainCosta
collecting, particularly in Costa Rica, have produced abundant RicatonorthernSouthAmerica,andhasbeenknowninMesoamerica
specimensshowingthesetwoentitiesasdistinct.Heisteriapovedae as H. longipes Standi, or H. cyanocarpa Poepp. Material of H.
is generally (as recognizedby Burger) aplantofhigherelevation, acuminata from Popaydn in southern Colombia may not match
somewhatdrierforests, thanH. macrophylla s.s., whichappearsto materialfromMesoamerica(J0rgensen,pers.comm.),andthecor-
beconfinedtothelowlandrainforestsontheAtlanticslopenearthe rect name for the material currently known as H. acuminata in
CostaRica/Nicaraguaborder.TheleavesofH.povedaeareusually Mesoamericamaychangeinfuture.
(butnotalways)smallerthanthoseofH. macrophylla s.s.,anddry
apaleolivegreenwithprominentyellowishvenation.Thestemsof
H. povedae are strongly flattened and winged atthe tips, with the Acknowledgements. Wethanktheherbariacitedinthetextforpermis-
wingspersistingontoquitelargebranchlets,andaremarkedlywhite siontoborroworexaminetheirspecimens;MargaretTebbsforpreparingthe
punctulate or striate. Stems ofH. macrophylla s.s. can be slightly illustration;NormanRobsonforcheckingtheLatindescription;HarryTaylor
winged when very small, but the angles rarely persistbeyond the ofthePhotographicUnitintheNHMforhelpingwiththedistributionmap;
firsttwoleaves. andPeterJ0rgensonfordiscussionsonthe systematicsofHeisteria anda
SpecimensfromLimonprovinceinCostaRicaarefoundwithin carefulreviewofthemanuscript.
thedistributionrangeoiHeisteriamacrophyllas.s.andmayrepres-
ent hybrids with that species. They have the typical yellowish
venation,wingedstemsandsubglobosefruitofH.povedae,butthe REFERENCES
leavesaregenerallylargerthanthosefromPacificslopespecimens.
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