A new species of genus Eriborus Förster (Hymenoptera: Ichneumonidae) parasitizing Holcocerus insularis Staudinger (Lepidoptera: Cossidae)

2006·0.93 MB·

by  Mao-Ling Sheng

©EntomologicaFennica.16June2006
Fungus gnats (Diptera: Bolitophilidae, Diadocidiidae,
Keroplatidae and Mycetophilidae) new to Finland
AlexeiPolevoi,JevgeniJakovlev*&AlexanderZaitzev
Polevoi,A.,Jakovlev,J.&Zaitzev,A.2006:Fungusgnats(Diptera:Bolitophi-
lidae, Diadocidiidae, Keroplatidae and Mycetophilidae) new to Finland. —
Entomol.Fennica17:161–169.
Thirty-sevenspeciesoffungusgnatsnewtoFinlandarereported.Elevenofthese
are reported in Fennoscandia for the first time: Diadocidia fissa Zaitzev,
Macrocera estonica Landrock, M. nigricoxa Winnertz, M. pusilla Meigen,
Boletina pallidula Edwards, Mycetophila morata Zaitzev, M. ostentanea
Zaitzev, Trichonta nigritula Edwards, T. subterminalis Zaitzev & Menzel,
NeoempheriawinnertziEdwardsandNeurateliasintenisiLackschewitz.There-
cordsarebasedonoriginalmaterialcollectedinlarge-scaletrappingprojectsin
SouthernandEasternFinlandmainlyinold-growthforestsduring1997–1998.
DetailedinformationonFinnishfindings,anddataonthegeneraldistributionof
thespeciesaregiven.Severalspeciesareknownwithonlyone(typematerial)or
afewpreviousrecordsrangingfromNorwaytoSakhalin.Fortwopoorly-known
species,NeurateliasintenisiLackschewitzandRymosiapinnataOstroverkhova,
newfiguresofmalegenitaliaarepresented.
A. Polevoi, Forest Research Institute, RU-185910, Pushkinskaya 11, Petro-
zavodsk,Russia;E-mail:[email protected]
J.Jakovlev,FinnishForestResearchInstitute(METLA),VantaaResearchUnit,
P. O. Box 18, FI-01301 Vantaa, Finland; *correspondent author’s e-mail:
[email protected]
A.Zaitzev,MoscowCityPedagogicalInstitute,DepartmentofBiology,Faculty
of Chemistry and Biology, 111568, Chechulina 1, Moscow, Russia; E-mail:
[email protected]
Received22June2005,accepted11November2005
1.Introduction treated as subfamilies in the family Myceto-
philidae (Edwards 1925). At present all these
Fungusgnatsisahighlydiversegroupofnemato- families are included into the superfamily
cerousDipteraincorporatingca.1500speciesin Sciaroidea(Väisänen1984,Matile1990,Hippa
the Palaearctic fauna and comprising about one &Vilkamaa2005).
tenthofalltheDipteranspeciesrecordedinFin- In Finland, the fungus gnat fauna has been
land.Thegrouphastraditionallybeenconsidered fairly well studied. The first data on Finnish
to include five families in Europe – Bolito- mycetophilids were published by Carl Lund-
philidae, Diadocidiidae, Ditomyiidae, Keropla- ström(Lundström1906,1909,1912,1913,1914)
tidae and Mycetophilidae – which were earlier and Richard Frey, whose data were included in
162 Polevoietal. (cid:127) ENTOMOL.FENNICAVol.17
thefirstFinnishDipterachecklist(Frey&Storå nuu (Ostrobotnia kajanensis, Ok). Atotal of 51
1941) and in Krogerus (1960). During 1960’s, sites of old-growth forest stands were studied.
RistoTuomikoskiandWalterHackmancollected Thepooledmaterialconsistsofover270,000in-
extensive material of mycetophilids throughout dividuals and more than 500 species of fungus
the country, mainly for taxonomical studies gnats.ThematerialisdepositedattheZoological
(Tuomikoski1966,Hackman1970,1971,Gagné MuseumoftheFinnishMuseumofNaturalHis-
1981).Thatmaterialwasutilizedinacheck-list tory, Helsinki. The sites where species new to
ofFinnishDiptera(Hackman1980)comprising FinlandwerefoundarelistedinTable1.
485speciesoffungusgnats.Sincethen,additions InTaandSt,thematerialwascollectedbyM.
anddeletionstotheFinnishfaunaoffungusgnats Kuussaari during 1998 in 19 old-growth forest
have been summarized by Silfverberg (1981, stands.Ineachstandthetrappingwasdoneusing
1986,1991,1996,2001)andPolevoi&Jakovlev fivestandardflight-windowtraps,onelighttrap
(2004).Thelatterreviewbroughtthetotalnum- andtwobaittrapsbaitedwithfermentedsugarso-
beroffungusgnatspeciesintheFinnishfaunato lution.Thetrapswereoperatingfrom4Mayto15
589. Octoberandwereemptied8–9timesduringthe
However, the list by Polevoi & Jakovlev season.Thepooledsamplewas31,745individu-
(2004) is still far from complete. In Russian als.
Karelia with rather similar nature, 616 species InKb,thematerialwascollectedbyM.Tietä-
havebeenrecordedafterintensivestudiesduring väinenintheNorthKarelianbiospherereservein
aperiodofjustabouttwentyyearsduring1977- themunicipalityofIlomantsiusingoneMalaise
2000 using rearing of fungus gnats from trapineachoftheninesampleplotsplacedinold-
macrofungi and collecting with netting and dif- growth forest stands. The traps were operating
ferent kinds of traps (Jakovlev 1995, Polevoi from14Julyto29Septemberandwereemptied9
2000). We believe that at least a hundred addi- timesin1997,andfrom11Juneto2Octoberand
tionalspeciesshouldbefoundinFinland.More- emptied17timesin1998.Thepooledsamplewas
over,thenumberofyetunknownspeciesmaybe 67,103individuals.
evenmuchhigher,asrecenttaxonomicalworks InOk,thematerialwascollectedbyM.Kuus-
(e.g. Zaitzev & Økland 1994, Polevoi 1995, saariduring1997in23old-growthforestpatches
2001, Zaitzev 2003, Polevoi & Hedmark 2004) in the municipalities of Kuhmo and Sotkamo.
have revealed many species new to science in ThetrappingdesignwasthesameasinTaandSt.
Fennoscandia. Thetrapswereoperatingfrom3Juneto8Octo-
Thispaperisbasedonoriginalmaterialcol- berandwereemptied6–7timesduringthesea-
lected during 1997-1998 in different parts of son.Thepooledsamplewas177,739individuals.
SouthernandEasternFinlandandintroduces37
speciesnewtoFinland.Sixteenofthesespecies
werealreadymentionedasknownfromFinland 3.ThelistofspeciesnewtoFinland
intheonlinedatabase“FaunaEuropaea”(Chan-
dler2005).However,theserecordsarebasedon Higher taxonomy follows Fauna Europaea (De
unpublishedinformationanddonotcontainde- Jong2005).Thespeciesrecordedbyusasnewto
taileddatae.g.oncollectinglocalitiesanddates. Fennoscandiaaremarkedbyasterisk(*).
2.Studyareas,materialandmethods 3.1.Heterotrichagroup
Thematerialwascollectedwithtrappinginold- Sciarosoma borealis Chandler, 2002. Sciaroso-
growthforestsinthreedifferentregionsinSouth- maChandlerisagenuswithaproblematicsys-
ernandEasternFinland.Thestudyareaswerelo- tematicpositiondescribedonthebasisofasingle
catedinthebiogeographicalprovincesofEtelä- speciesfromNorwayandRussianKarelia(Chan-
Häme (Tavastia australis, Ta), Satakunta (St), dler 2002) and later reported from Germany
Pohjois-Karjala(Kareliaborealis,Kb)andKai- (Kallweit&Jaschhof2004)andfromCzechRe-
ENTOMOL.FENNICAVol.17 (cid:127) FungusgnatsnewtoFinland 163
Table1.Thestudysites,withinformationontheirlocality.Province=biogeographicprov-
ince;Municip.=municipality;Coords.=coordinates(E27grid),Year=samplingyear.
Studysite Province Municip. Coords. Year
Ahvenus St Parkano 6888:3295 1998
Elimyssalo Ok Kuhmo 7129:3667 1997
Honkavaara Ok Kuhmo 7091:3632 1997
Iso-Saarijärvi Ta Ruovesi 6882:3337 1998
Jauhovaara Ok Kuhmo 7106:3603 1997
Jonkeri Ok Kuhmo 7093:3636 1997
Kolmikoura Ta Ruovesi 6868:3364 1998
Kotavaara Kb Ilomantsi 6998:3721 1997–1998
Kotiset Ta Lammi 6794:3396 1998
Kuivikkovaara Ok Kuhmo 7125:3654 1997
Kujanki Ok Kuhmo 7086:3618 1997
Lakeisnevankangas Ta Virrat 6920:3344 1998
Lauvuskylä Ok Kuhmo 7089:3626 1997
Luhtavaara Ok Kuhmo 7152:3610 1997
Lymylampi St Ikaalinen 6862:3319 1998
Metsäopisto Ta Kuru 6866:3320 1998
Multinharju St Ikaalinen 6872:3311 1998
Musturi Ta Ruovesi 6865:3362 1998
Nälkähittenkangas St Parkano 6892:3295 1998
Näveri Ok Kuhmo 7104:3599 1997
Pellinkangas Ok Kuhmo 7135:3594 1997
Petäjäjärvi Ta Kuru 6871:3320 1998
Pirhu Kb Ilomantsi 6993:3723 1997–1998
Pitkävaara Ok Kuhmo 7162:3617 1997
Puntari Ok Kuhmo 7090:3616 1997
Puukkohonka Ta Lammi 6792:3395 1998
Rengassalo St Parkano 6897:3285 1998
Riihisuo Ok Kuhmo 7102:3610 1997
Sudenpesänkangas Ta Lammi 6789:3403 1998
Susimäki Ta Ruovesi 6864:3355 1998
Särkijärvi Ok Kuhmo 7111:3622 1997
Teerisuo Ok Kuhmo 7091:3608 1997
Tulisuo-Varpusuo Ok Kuhmo 7164:3616 1997
Urpovaara Ok Sotkamo 7089:3602 1997
Vepsä Ok Kuhmo 7091:3615 1997
Vesijako Ta Padasjoki 6806:3399 1998
publicandFinlandwithnolocality data(Chan- and specimens from Russian Far East (Zaitzev
dler 2005). It was not included in any existing 1994). Reported from Finland with no locality
familyuntilthenewconceptofSciaridae(Hippa data(Chandler2005).Material:1#,Ok:Jonkeri,
&Vilkamaa2005)withSciarosomaplacedinthe 27.VIII.–10.IX.1997; 1 #, Kb: Kotavaara, 15.–
familySciaridae,subfamilySciarosominae.Ma- 29.IX.1997.
terial:3 ##, Kb:Kotavaara, 22.–29.VI.1998;1
#,Ok:Luhtavaara,15.–27.VII.1997.
3.3.FamilyDiadocidiidae
3.2.FamilyBolitophilidae *DiadocidiafissaZaitzev,1994.Typespecimens
fromVologdaprovince(Zaitzev1994)werethe
Bolitophila caspersi Plassmann, 1986. Previ- only known specimens until now. Material: 2
ously known only from Swedish type material ##,Kb:Kotavaara,8.–15.IX.1997.
164 Polevoietal. (cid:127) ENTOMOL.FENNICAVol.17
3.4.FamilyKeroplatidae 3.5.FamilyMycetophilidae
3.4.1.SubfamilyKeroplatinae 3.5.1.SubfamilyGnoristinae
TribeKeroplatini Boletina digitata Lundström, 1914. Distributed
KeroplatustuvensisZaitzev,1991.Ararespecies inEuropeandMiddleEast.InFennoscandia,pre-
ofthetestaceusgroup.DescribedfromTuvaRe- viously known from the type locality, viz.
public (Zaitzev 1991) and later reported from Kandalaksha, Murmansk province, Russia
RussianKarelia(Polevoi1995)andfromCzech (Lundström 1914) and Sweden (Plassmann
Republic and Finland but with no locality data 1979).AccordingtoPlassmann(1979)andCata-
(Chandler2005).Material:1#,Ok:Pitkävaara, logueofPalaearcticDiptera(La(cid:1)tovka&Matile
12.–25.VIII.1997. 1988)thisspecieshasalsobeenfoundinFinland,
but was not mentioned in the Finnish Diptera
TribeOrfeliini check-list (Hackman 1980) nor in the updates
Pyratula perpusilla (Edwards, 1913). A Euro- published afterward. Chandler (2005) gives the
pean species; in Fennoscandia, previously re- Finnishrecordwithnolocalitydata.Material:5
corded from Sweden (Plassmann 1974, 1978) ##,Kb:Kotavaara,22.VI.–8.IX.1998.Wehave
andNorway(Økland&Zaitzev1997).Material: alsofoundtwomalesofthisspeciesinthecollec-
1 #, Ok: Pitkävaara, 26.VIII.–8.IX.1997; 1 #, tion of the Finnish Museumof Natural History,
Ok:Tulisuo-Varpusuo,8.–24.VIII.1997. Helsinki: one male from Ob: Uleåborg (V.
Wuorentaus) and another from Le: Malla (R.
3.4.2.SubfamilyMacrocerinae Frey).
BoletinajamalensisZaitzev,1994.Described
*MacroceraestonicaLandrock,1924.Thisspe- from Siberia (Zaitzev 1994) and later found in
cies is apparently widely distributed in the NorthernEurope(Økland1996,Hedmark1998,
Palaearcticregionbutitwasnotearlierreported Polevoi 2000). The Finnish record was earlier
from Fennoscandia. The record from Finland publishedwithnolocalitydata(Chandler2005).
(Chandler 2005) is given with no locality data. Material:1#,Kb:Kotavaara,15.–29.IX.1997;1
Material: 5 ##, Kb: Kotavaara, 14.–25.VIII. #, same place, 2.–8.IX.1998; 1 #, Ok:
1997. Luhtavaara, 17.–30.VI.1997; 2 ##, St: Lymy-
*MacroceranigricoxaWinnertz,1863(=M. lampi, 26.V.–12.VI.1998. We have also found
tuscaLoew).Thisspeciesiswidelydistributedin twomalesofthisspeciesinthecollectionofthe
Western and Central Europe. It has also been Finnish Museum of Natural History, Helsinki,
foundinIsrael(Chandler1994a)andinCentral both from Ka: Vehkalahti, 5.VI.1971 and
Asia (Zaitzev 1994). The Finnish record is the 9.X.1971.
firstonefromFennoscandia.Material:3##,Kb: * Boletina pallidula Edwards, 1925. A rare
Kotavaara, 15.–29.IX.1997; 3 ##, same site, European species not previously reported from
25.VIII.–8.IX.1998. Fennoscandia. The record from Finland (Chan-
* Macrocera pusilla Meigen, 1830 (= M. dler2005)isgivenwithnolocalitydata.Material:
nana Macquart). This species is widespread in 35##,Kb:Kotavaara,17.VII.–21.IX.1998;1#,
Europe,includingNorthAfricaandMiddleEast Ta:Kotiset,4.–17.VIII.1998;1#,Ok:Kujanki,
(Chandler 1994a) and has also been reported 31.VII.–13.VIII.1997.
from Central Asia and Japan (Zaitzev 1994). It BoletinatiroliensisPlassmann,1980.Species
wasnot,however,earlierfoundinFennoscandia. describedfromAustria(Plassmann1980a),later
Material:9##,Kb:Kotavaara,17.VII.–20.VIII. foundinSweden(Hedmark1998)andinseveral
1998. northernlocalitiesinRussia(Zaitzev1994).The
record from Finland (Chandler 2005) is given
with no locality data. Material: 1 #, Ok:
Kuivikkovaara, 15.–29.IX.1997; 1 #, Ok:
Pellinkangas, 10.–28.IX.1997; 2 ##, Ok:
ENTOMOL.FENNICAVol.17 (cid:127) FungusgnatsnewtoFinland 165
Riihisuo,12.IX.–7.X.1997;1#,Ok:Urpovaara, Exechia similis La(cid:1)tovka & Matile, 1974.
12.IX.–8.X.1997. This species is closely related to E. spinuligera
Ectrepesthoneura tori Zaitzev & Økland, Lundström, which is widely distributed in Eu-
1994. A rare species described from Norway. rope,andhaspossiblybeenoverlookedinFenno-
Also known from Germany (Kallweit & Plass- scandia.ReportedfromRussianKarelia(Polevoi
mann1999),Sweden(Kurinaetal.2005),Great 2000).Material:7##,Kb:Kotavaara,25.VIII.–
Britain and Czech Republic (Chandler 2005). 8.IX.1998.
Material:1#,Kb:Kotavaara,11.–15.VI.1998. ExechiasubfrigidaLa(cid:1)tovka&Matile,1974.
Tetragoneura sylvatica Curtis, 1837. This ThisspeciesiscloselyrelatedtoE.frigidaBohe-
speciesiswidelydistributedinEurope.InFenno- man, which is a widely distributed Trans-
scandia it has been found in Norway (Økland palaearctic species. It has been described from
1996, Økland & Zaitsev 1997) and Sweden Mongolia (La(cid:1)tovka & Matile 1974) and then
(Kurinaetal.2005).ReportedfromFinlandwith beenreportedfromdifferentlocalitiesinRussia
nolocalitydata(Chandler2005).Material:1#, (Zaitzev2003).IthasalsobeenfoundinNorway
Kb: Kotavaara, 29.VI.–2.VII.1998; 1 #, Ok: (Søli1994)andSweden(Hedmark2000).Mate-
Teerisuo,12.IX.–2.X.1997. rial:1#,Ok:Vepsä,11.IX.–6.X.1997;3##,Kb:
Kotavaara,8.–29.IX.1997;4##,Kb:Kotavaara,
3.5.2.SubfamilyMycetophilinae 20.VIII.–8.IX.1998.
Exechiopsis (Exechiopsis) pseudindecisa
TribeExechiini La(cid:1)tovka&Matile,1974.Thisspeciesisappar-
Allodia (Brachycampta) angulata Lundström, entlywidelydistributedinthePalaearcticregion.
1913.ArareEuropeanspecies.Sincetheoriginal InFennoscandia,itwasfirstrecordedfromNor-
description fromSwedish Lapland (Lundströrm way(Kjærandsen1993,Økland&Zaitzev1997),
1913), it has been reported from Great Britain and later on from Sweden (Plassmann 1980b),
(Chandler1977),Denmark(Chandler&Petersen and from Karelia (Polevoi 2000). The record
2001), Germany (Plassmann & Schacht 1999, fromFinland(Chandler2005)lackslocalitydata.
2002), France (Chandler 2005) and Vologda Material: 4 ##, St: Ahvenus, 25.VIII.–12.X.
province in Russia (Zaitzev 2003). Material: 2 1998;1#,Ok:Vepsä,13.–26.VIII.1997;1#,Ta:
##,Kb:Kotavaara,30.VII.–20.VIII.1998. Iso-Saarijärvi, 15.IX.–14.X.1998; 4 ##, Ok:
Anatella ankeli Plassmann, 1977. This spe- Jonkeri, 27.VIII.–10.IX.1997; 8 ##, Kb: Kota-
cies is known from several European localities vaara,30.VII.–2.X.1998;7##,Ok:Luhtavaara,
(Chandler1994b,2005,Zaitzev2003).InFenno- 25.VIII.–9.IX.1997;4##,Ok:Näveri,25.VIII.–
scandia,thespecieshasbeenreportedfromNor- 2.X.1997;1#,St:Nälkähittenkangas,25.VIII.–
way (Kjærandsen 1993) and Russian Karelia 16.IX.1998; 2 ##, Ok: Pellinkangas, 25.VIII.–
(Polevoi2000).TherecordfromFinland(Chan- 9.IX.1997; 2 ##, Ok: Puntari, 27.VIII.–10.IX.
dler2005)isgivenwithnolocalitydata.Material: 1997; 3 ##, St: Rengassalo, 12.VIII.–13.X.
1#,Ok:Pitkävaara,26.VIII.–8.IX.1997. 1998;1#,Ta:Sudenpesänkangas,17.–31.VIII.
Brachypeza (Paracordyla) obscura Win- 1998;2##,Ok:Jauhovaara,11.–24.VIII.1997.
nertz,1863.Thisspeciesiswidelydistributedin Rymosia pinnata Ostroverkhova, 1979. De-
Palaearctic(Zaitzev2003).InFennoscandia,the scribed fromSiberia and Russian Far East. The
speciesisknownfromRussianKarelia(Polevoi originalfiguresofOstroverkhova(1979;p.175,
2000). The Finnish record (Chandler 2005) is figure35:2),thoughbeingsketchy,allowreliable
given with no locality data. Material: 1 #, Kb: identification, especially due to the outstanding
Kotavaara, 17.–23.VII.1998; 1 #, Ok: Puntari, parameres resembling the wing of a bird. The
2.–15.VII.1997. male genitalia are illustrated in Fig. 1. The re-
Brevicornu occidentale Zaitzev, 1988. De- cords following the original description have
scribedfromNorthAmerica(Zaitzev1988)and been from Sweden (Hedmark 2000) and from
later found in Norway (Økland 1996) and Rus- Finland with no locality data (Chandler 2005).
sian Karelia (Polevoi 2000). Material: 1 #, Kb: Material: 1 #, Ta: Kolmikoura, 24.VIII.–
Kotavaara,23.–26.VII.1998. 15.IX.1998.
166 Polevoietal. (cid:127) ENTOMOL.FENNICAVol.17
Fig.1.RymosiapinnataOstro-
verkhova,malegenitalia.–a.Left
gonostylus,internalview.–b.
Gonocoxitesandrightgonostylus,
ventralview.–c.TergiteIXand
cerci,dorsalview.–d.Aedeagus
andparameres,ventralview.–e.
Aedeagusandparameres,lateral
view.Scalebar0.1mm.
TribeMycetophilini Russian Karelia (Polevoi 2000). Material: 11
*MycetophilamorataZaitzev,1999.Aspeciesof ##, Kb: Kotavaara, 14.VIII.–17.IX.1997; 1 #,
thevittipesgroup.Itwassofarknownonlybythe Ta:Musturi,11.–24.VIII.1998.
typematerialfromEasternPalaearctic–Sakhalin * Trichonta nigritula Edwards, 1925. The
and Altai (Zaitzev 1999). Material: 1 #, Kb: specieshasbeenreinstatedbyP.Chandler(1992)
Pirhu,6.VI.1994;1#,Ok:Honkavaara,11.IX.– after having been synonymized with Trichonta
5.X.1997. vitta(Meigen)byR.Gagné(1981).Ithasprevi-
* Mycetophila ostentanea Zaitzev, 1998. ously not been known from outside the British
KnownbythetypematerialfromNorth-Western Isles. Material: 3 ##, Kb: Kotavaara, 29.VI.–
Russia, Vologda province (Zaitzev 1998, 2003) 08.IX.1998.
andbyarecordfromCzechRepublic(Chandler *TrichontasubterminalisZaitzev&Menzel,
2005).Material:2##,Kb:Kotavaara,17.VII.– 1996. This species was described from Russian
11.VIII.1998. Far East (Zaitzev & Menzel 1996). It is closely
MycetophilapecinaiLa(cid:1)tovka,1963.Besides related to T. terminalis (Walker) and T. facilis
thetypematerialfromBohemia,thisspecieswas GagnéandmayhavebeenoverlookedinEurope.
reported from Sweden (Plassmann 1980b) and ItmustbenotedthatT.funebrisWinnertz,which
other European countries (Chandler 2005). The was considered a synonym of T. terminalis
Finnishrecordgivenbythelatterauthorcontains Walker by Edwards (1913), resembles T.
no locality data. Material: 1 #, Kb: Kotavaara, subterminalis,byhavinglargetriangularventral
8.–17.IX.1998; 1 #, Ok: Särkijärvi, 18.– cleftonthegonocoxites(Dziedzicki1915;table
29.VII.1997. XI, figure 149). However, the figure of T.
TrichontaapicalisStrobl,1898.AEuropean terminalisbyR.Gagné(1981;p.48,figure93)in
species;inFennoscandia,previouslyknownonly the revision of the genus, is distinctly different.
fromSweden(Plassmann1980b).Material:1#, Material:1#,Kb:Kotavaara,20.–25.VIII.1998.
Kb:Kotavaara,25.VIII.–2.IX.1998.
TrichontaeximiaGagné,1981.Thisspeciesis 3.5.3.SubfamilyMycomyinae
common in North America. In Eurasia it was
known from the Himalayas (Gagné 1981) and MycomyafuscataWinnertz,1863.Knownfrom
ENTOMOL.FENNICAVol.17 (cid:127) FungusgnatsnewtoFinland 167
* Neoempheria winnertzi Edwards, 1913.
Known from Europe and Iran. No previous re-
cordsfromFennoscandia,withtheexceptionof
onefromFinlandwithnolocalitydata(Chandler
2005).Material:1#, Ok:Teerisuo,16.–29.VII.
1997; 3 ##, Ok: Urpovaara, 16.VII.–11.VIII.
1997.
3.5.4.SubfamilySciophilinae
*NeurateliasintenisiLackschewitz,1937.Previ-
ouslyknownonlybytypematerialfromEstonia
(Lackschewitz1937)andbyanadditionalrecord
Fig.2.NeurateliasintenisiLackschewitz,malegenita- from Czech Republic ((cid:2)ev(cid:3)ík 2001). Reported
fromNorth-WesternRussiaandFinlandwithno
lia.–a.Ventralview.–b.Dorsalview.Scalebar0.1
mm. localitydata(Chandler2005).Malegenitaliaare
illustrated in Fig. 2. Material: 2 ##, Kb: Kota-
Europe and North America. In Fennoscandia, vaara, 23.VII.–6.VIII.1998; 1 #, St: Nälkä-
previously reported fromNorthern Sweden and hittenkangas,15.–31.VII.1998.
Murmanskprovince(Väisänen1984).Material: PhthiniasetosaZaitzev,1994.Ararespecies
2##,Ok:Teerisuo,12.IX.–2.X.1997. known so far only from Fennoscandia: Russian
MycomyanornaVäisänen,1984.Previously Karelia(Zaitzev,1994),Norway(Økland1996)
known by the holotype from Northern Sweden and Sweden (Hedmark 1998). Material: 2 ##,
(Väisänen 1984), and additional records exist Kb:Kotavaara,23.VII.–11.VIII.1998.
from South-Eastern Norway (Økland 1996), SciophilamodestaZaitzev,1982.AHolarctic
Germany and Finland (Chandler 2005). The species;widelydistributedinNorthAmerica.In
Finnish record (Chandler 2005) has no locality the Palaearctic region mostly recorded from
data.Onthebasisofthepresentmaterialthisspe- northern areas including Russian Karelia
ciesappearstobecommoninold-growthforests (Zaitzev1994).Material:3##,Kb:Kotavaara,
in Southern Finland. All individuals were col- 22.VI.–6.VIII.1998.
lectedwithbaittraps.Material:1#,St:Ahvenus, SciophilanonnisilvaHutson,1979.AHolarc-
25.VIII.–16.IX.1998; 17 ##, Ta: Vesijako, ticspecies.WidelydistributedinEurope(Chan-
7.VII.–12.X.1998; 13 #, Ta: Iso-Saarijärvi, dler 2005), recorded also from Azerbaidjan
15.VII.–13.X.1998; 4 #, Ta: Kolmikoura, (Zaitzev1994).InFennoscandia,thespecieshas
31.VII.–24.VIII.1998;13#,Ta:Kotiset,7.VII.– previously been found in Norway (Økland &
12.X.1998; 11 #, Ta: Lakeisnevankangas, 10.– Zaitzev1997)andSweden(Kurinaetal.2005).
21.VIII.1998; 1 #, Ok: Lauvuskylä, 13.– Material:1#,Kb:Kotavaara,13.–17.VII.1998.
26.VIII.1997; 88 ##, St: Lymylampi, 12.VI.– SciophilapersubtilisPolevoi,2001.Thisspe-
13.X.1998; 48 #, Ta: Metsäopisto, 24.VIII.– cieshaspreviouslybeenknownonlybythetype
17.IX.1998; 2 ##, St: Multinharju, 10.VIII.– material from Russian Karelia (Polevoi 2001).
16.IX.1998; 20 ##, Ta: Musturi, 24.VI.– Material: 6 ##, Kb: Kotavaara, 11.VI.–8.IX.
15.X.1998; 102 ##, St: Nälkähittenkangas, 1998.
12.VIII.–16.IX.1998; 1 #, Ok: Pellinkangas,
25.VIII.–9.IX.1997;6##,Petäjäjärvi,30.VII.–
Acknowledgements. We are grateful to Timo Hokkanen
17.IX.1998; 37 ##, Ta: Puukkohonka, 24.VI.– (North Karelian Environment Centre, Joensuu), Raimo
12.X.1998; 3 ##, St: Rengassalo, 15.VII.– HeikkiläandGergelyVarkonyi(KainuuRegionalEnvi-
13.X.1998;1#,Ok:Riihisuo,12.–25.VIII.1997; ronmentCentre,ResearchCentreoftheFriendshipPark,
Kuhmo)forprovidinguswithlong-termtrappingmaterial.
1#,Ta:Susimäki,21.VIII.–15.IX.1998;32##,
WealsothankJuhaSiitonen(FinnishForestResearchIn-
Ta:Sudenpesänkangas,7.VII.–21.IX.1998;1#,
stitute,Vantaa)andtherefereesforvaluablecommentson
Ok:Elimyssalo,28.VII.–7.VIII.1997. themanuscript.Thispaperispartoftheproject“Finnish
168 Polevoietal. (cid:127) ENTOMOL.FENNICAVol.17
fungus gnats (Diptera, Mycetophilidae, etc.): faunistics, gens group from Northern Europe. — Annales
habitatrequirementsandthreatstatus”.Thefinancialsup- EntomologiciFennici37:3–7.
portfromtheFinnishMinistryofEnvironmentisacknowl- Hackman,W.1980:AchecklistoftheFinnishDipteraI.
edged. Nematocera and Brachycera (s.str.). — Notulae
Entomologicae60:17–48.
Hedmark,K.1998:Svampmyggor–nyaarterförSverige
och Finland (Diptera: Mycetophilidae s. lat.). —
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