A New Genus with Two New Species of Hygrophilous Trechine Beetles (Coleoptera, Trechinae) from the Daba Shan Mountains, West-central China

2004·7.2 MB·

by  Ueno, S-I

E/、・tてl1,Tokyo,32(2):265-275、November20,2004
ANewGenuswithTwoNewSpeciesofHygrophilousTrechineBeetles
(Coleoptera,Trechinae) fromtheDabaShanMountains,
West-central China
Shun-lobi UENo
DepartmentofZoology,NationalScienceMuseum(Nat.Hist),
3-23-1Hyakunin-cho,Shinjuku,Tokyo,l69-0073Japan
Abstract Anewgenusiserectedfortwohygrophiloustrechinebeetlesbelonging
totheAgono11echusseriesfromtheDabaShanMountainsinWest-centralChina.It isre-
latedtoDoハt ,eot1・ec/nlsS.UENofromnorthwesternYunnan,butisreadilydistinguishedby
mandibular,labialandchaetotaxialpeculiarities.ThenewnamesgivenareDab(;111・coitus
forthegenus,Decce11t1・Iousforthewesternspecies,andD. /1yg1・op1/1/usfortheeastern
species.Thetwonewspeciesarccloselysimilaringeneralappearance,butareincredibly
different、tromeachotherinconformationofthemalegenitalia.
Earlyinthesummerofthisyear,lhadanopportunitytomakeaninvestigationof
thecoleopteranfaunaoftheDabaShanMountainsthatstretchfromwesttoeastonthe
bordersofShaanxi andSichuanProvincesandextendeastwards intothe westernmost
partofHubeionthenorthernsideoftheYangziJiangRiver.CollaboratedwithMasa-
takaSAT0, Ivisitedvariousplacesincludingthirteenlimestonecavesoneithersideof
themountainrange,andcollectedafairnumberoftrechinebeetles,amongwhichwere
foundtwospecimensapparentlybelongingtotheAgonotrechusseries.Theylooklike
Doハt ,eotrec/1usylnae fromwesternYunnaninparticular,butareevidently different
fromitinsomecriticalpoints.0neofthemwasfoundnearthewesternmostpartofthe
mountainrange,andtheotherattheeasternmostpartabout430kmapart fromthecol-
lectingsiteoftheformer.
Thoughsimilar toeachotherat firstsight, thetwospecimens aredifferent in
manydetailsofexternalmorphology,andbesides,incrediblydifferinconformationof
themalegenitalia. Thegenitalic differencealmostattainstothelevel thatcouldbe
consideredasthatofgenericimportance.Atthepresentmoment,however, Iprefer to
regardthemasbelongingtothesamegenusinviewoftheclosesimilarityindiagnos-
tic featuresofexternalmorphology,andamgoingtoerect forthemanewgenustobe
calIedDabatrec1/us.Similardiscrepancybetweenexternalandgenitalicpeculiaritiesis
alsoknown inthegenusDeuveot1-echusS. UENo(1995, p 99,andalso1996)and the
genusLuzonotrec/1usS.UENo(1979,p 26,andalso1987),andcanberegardedasa
uniquediversificationexhibitedbythemembersoftheAgo,7ot1-coitusseriesanditsde-
rivatives.
266 Shun-lobiUENo
Theseinterestingnewtrechineswillbedescribedinthepresentpaperunderthe
names Dabatrechus eccentrlcus and D hyg1ophilus, the former from Mt. Guangwu
Shanlyingat theSichuansideofthewesternmostpartoftheDabaShanMountains
andthelatterfromtheYidaoXiaValleyattheeasternmostpartofthesamemountains
inwesternHubei. Theabbreviationsusedhereinarethesameasthoseexplainedin
previouspapersofmine.
Beforegoinginto furtherdetails, Iwishtoexpressmyheartfelt thankstoDr.
MasatakaSAT0 forhiscollaborationinthefief andtoMr.FANTingoftheAcademia
Sinicaforhisunfailinghelptoourinvestigations.
GenusDabatrechusS.UENo nov
Typespecies: Dabatrechuseccent1・iousS.UENo,spnov.
BelongingtotheAgonotrechusseriesandrelatedtoDeuveotrechusS.UENofrom
northwestern Yunnan, but different from it in thebidentate right mandible, fused
labium,andtheabsenceofsupernumerarydorsalporesonthethirdelytra1stria.
Medium-sizedtrechineswithrelativelyshortbroadbodyeitherconstrictedbe-
tweenprothoraxandhindbody(D hyg1ophilus)ornot;dorsalsurfaceglabrousand
polished,ventereitherglabrous(」Deccentrlcus)orprovidedwithseveralshorthairson
themedianpartofeachsternite(D hygroph11us);microsculptureeithercompletelyab-
sent(D hygrophilus)orpresentonlyinmarginalareasofpronotumasfinetransverse
lines(Deccentrlcus);colourdarkbrownwithpalerpalpiandlegs;innerwingsatro-
phied.
Headtransverse,withdeepentirefrontal furrowsstronglyarcuateorevensuban-
gulateatmiddleandwidelydivergent in frontandbehind; twopairofsupraorbital
porespresentonlinessubpara11e1toeachotherorslightlydivergentposteriad,theante-
riorpairfoveolateandlyingneartotheposteriorpair;eyesfairlylargeandconvexto-
getherwithgenae,eitherclearlyfaceted(Deccentricus)ornot,genaeeitherglabrous
(Dhygrophilus)orwithafewvestigialhairs(D eccentricus); labrumtransverseand
dilatedanteria withtheapicalmargindeeplyandevenlyemarginate.Mandiblesstout
thoughacuteattheincurvedapicalportions,rightmandiblebidentate,devoidofpre-
molar tooth. Labium fusedthoughvestigeoflabialsutureisrecognizable;mentum
longitudinallydepressedoneachside,mentaltoothbroad,eithertruncatedorrounded
atthetip;submentumsexsetose;ligulaporrectatthemiddle,octosetoseasusual;para-
glossaethinandarcuate,extendingmuchbeyondligula; labialpalpus fairlyslender,
penultimate palpomere gradually dilated towards apex and quadrisetose, apical
palpomereslightly longer than thepenultimate. Maxillaeshort,stout and arcuate,
1aciniawithseveralstoutspinesandhairsontheinnermargin;maxillarypalpuswith
shortpenultimatepalpomerewhichiswidelydilatedtowardsapexandbearsvestiges
ofafewminutehairs,apicalpalpomereelongatedsubconica1,about13timesaslong
asthepenultimate.Antennaesubfiliform,notreachingthemiddleofelytra.
Pronotumsubcordateandconvex,eithergraduallynarrowedposteriad(D eccen-
NewTrechincBeetles、fromtheDabaShanMountains 267
tricus)orratherstronglycontractedatthebase(Dhyg1,〔)ph11us),whichiseitherbriefly
or vestigially subpedunculate; sides completely bordered, more or less strongly
roundedinfront,eitherstraight(Deccentricus)orbrieflysinuatejustbeforehindan-
gles(Dhyg1-〔)phihls),withtwopairofmarginalsetae,ofwhichtheanteriorpair isin-
sertedataboutoralittlebeforethewidestpartandtheposterioronealmostonhind
angles; frontanglescompletelyroundedoff,hindangleseitherobtuse(Deccent1-1cus)
orsubrectangular(Dhyg1,oph11us);nopostangularcarinae.Scutellumsmall.
Elytraovate,evidentlywider thanforebody,withdistinctbutroundedshoulders;
dorsumconvex;sidesbordered,prehumera1bordereithercompletetobasalpeduncle
(Deccentrlcus)ordiminishingantero-internallythoughreachingthesiteofthebase
ofstria5;striaevanishedwiththeexceptionofinnertwoorthreeandapicalhalfof8,
theformerofwhich(atleastthesuturalone)arecoarselypunctateorcrenulateonthe
disc;scutellarstrictodistinct,apicalstriolealsodistinct thoughshortand freeatthe
anteriorend;oneor twosetiferousdorsalporespresentonor onthesiteofstria3,
none on stria 5; preapica1 pore either present (D. eccent1・Ious) or absent (D
hyg1,oph11us);twoapicalporespresentasusual;marginalumbilicateporesregular, four
poresofthehumeralsetrangedalmostequidistantly.
Ventral surfacesmooth,either glabrousorbearingseveral shorthairs oneach
sternitebetweenapairofparamediansetae;analsternitebisetosein(S.Legsofmoder-
atelength;protibiaepubescentthoughratherminutelyontheanterior face,eithersim-
ple(D hygrophilus)or vestigiallygroovedontheexternal face(D eccentricus); tarsi
thin; in , twoproximal tarsomeresofeachprotarsusmoreor lessdilated, inwardly
producedordenticulateatapices,andfurnishedbeneathwithadhesiveappendages.
Malegenitaliasurprisinglyvariableinconformationaccordingtospecies.Aede-
agusgutter-like,widelymembraneousondorsum frombasetoapical lobe,withthe
basalpartopenonbothdorsalandventral sides,sometimesformingakindofring,
thoughthebasalextremityiscompletelyclosedandbearsasagittalaileron;apicallobe
eitherveryshort(Decce11tricus)orlongandnarrow(Dhyg1-ophilus);innersacscaly
at theapicalpartandbearingone(D. り/ ,gl,oph11us)or two(Deccent1・Ious)copulatory
pieces,spoon-likeandhyalineinthe former,palp-likeandheavilysclerotizedinthe
latter.Stylesnarrowat theapicalparts, leftstyleobviouslylongerthantherightand
bearingreducedventralapophysis,eachprovidedwithfour,relativelyshortsetaeatthe
apex.
Range. Known so far only from the Daba Shan Mountains in West-central
China.
Notes. ThoughsimilarinmanyrespectstoDeuveotrechus,thisnew genusmay
notbedirectlyrelatedtoitinviewoftheradicaldifferenceinsomefundamental fea-
tures,and theoverallsimilaritybetween thetwogeneramayhavebeenbroughtabout
throughparallelevolution. 0ntheotherhan similarunusualdivergenceofgenitalic
conformation is also known in Deuveotrechus. Indescribing Deu、l,eotrechusylnae
(UENo,1996,p.14,figs.1-3) fromtheDiancangShanMountainsinwesternYunnan,
Inoticedthatthemalegenitaliaofthenewspecieswerestrikinglydifferent fromthose
268 Shun-lchiUltNo
ofthetypespecies,Dyuae(DEuvE,1992,p.171,figs.1,12;UENo,1995,p.100,figs.
4-5),andthat thepeculiarmodificationofmalegenitaliashownby D yinaewasto
someextentsimilar tothatshownbythebontocgroupofthegenusLuzonotrechus,
whichmighthavearemoterelationshiptothegeneraoftheAgonotrechusseries.The
discoveryofDabatrechusseemstogiveanotherexampleofunusualdiversification,
suggestingthatthegenitalicdivergencemaynotbeexceptional intheAgonotrechus
series.
Aswillbedescribedundertheheadingsofthenewspecies,theknownmembers
ofDabatrechusareunusuallyhygrophilousandarefoundincoexistencewithbembidi-
ines. Thisisquiteexceptional for theapterousmembersoftheAgonotrechusseries,
mostofwhichareeitherhumico1ous,muscico1ousorsaproxy1ophi1ous.
DabatrechuseccentricusS.UENo,sp nov
(Figs.1-3)
Length: 4.10mm(fromapicalmarginofclypeustoapicesofelytra).
Bodyshort andbroad, glabrousandpolishedonbothdorsumand venter;mi-
crosculpturemostlyvanishedbutperceptibleasfinetransverselinesinmarginalareas
ofpronotumand insmall fragmentary portions along frontal furrows. Colour dark
brown,shiny,lighter inbasalhalvesofelytra1disc,whichisweakly iridescent;palpi
andlegslightyellowishbrown.
Headwide,depressedabove,withdeepfrontal furrowswhicharenotangulateat
middleandmorewidelydivergentposteriadthananteriad;supraorbital areasgently
convex, with twopair ofsupraorbitalpores onlinessubpara11e1toeachother and
closely locatedoneafter theother,theanteriorpair beinglocatedslightlybehindthe
mid-eyelevel;eyesmoderatelyconvexanddistinctly faceted;genaeobliqueandonly
slightlyconvex,two-thirdsaslongaseyes,andpracticallyglabrousthoughbearinga
fewvestigialhairs;neckverywide,withtheanteriorconstrictionsharplymarkedat
thesides;palpishort thoughlabialpalpusis fairlyslender;mental toothbroad,trun-
catedat thetip;antennaefairlystout, reachingbasalthree-eighthsofelytra,scapeas
longasandabout15timesasthickassegment3,whichisslightlyshorter thanseg-
ment4,pediceltheshortest,five-seventhsaslongassegment3,segments5-8gradu-
allydecreasinginlengthtowardsapex,eachaboutthreetimesoralittlelessaslongas
wide,segments8-10equal inlengthtooneanother, terminalsegment thelongest,a
littlelongerthanscapethoughtwo-thirdsaswideasthelatter.
Pronotumrelativelylarge,transversesubcordate,muchwiderthanhead,evidently
wider thanlong,widestatabout two-thirdsfrombase,andmorerapidlyandstrongly
narrowedtowardsapexthantowardsbase,withthebasalpartbrieflysubpedunculate;
PW/HW134,PW/PL129,PW/PAca.157,PW/PB137;sidesratherwidelyre?exed
beforemiddle,narrowlysoinbasalthird,stronglyroundedinfront,almoststraightly
convergentbehind, andverybrie ysinuateataboutbasalsixthatthesidesofpostan-
gular denticles; apex slightly arcuate forwards, front angle on each side widely
NewTrechineBeetlesfromtheDabaShanMountains 269
Fig. 1. Dabat,-ec1/useccent,・IousS.UENo,sp nov., fromMt.GuangwuShan
,
roundedoff;basewiderthanapex,PA/PBca.087,gentlyobliqueoneachsideinside
hindangle,which formsanobtusedenticleslightlyproducedlaterad;dorsumconvex,
withvague,irregularlytransversestriations;medianlineshallow,reachingneitherapex
norbase;apicaltransverseimpressionmat-defined;basaltransverseimpressionsuper-
ficial anduneven. witha foveoleoneachsideofmedian line; basal foveaesmall but
270 Shun-lobiUENo
fairlydeep;basalareaabovebasalpedunclenarrowandsmooth.
Elytraovate,widerthanpronotum,widestslightlybeforethemiddle,withsmall
basal partsand relatively pointedapices;EW/PW152,EL/PL2.78, EL/EW 141;
shoulderswidely roundedinclusiveofarcuateprehumera1borders,whicharerather
widelyborderedandcompletetobasalpeduncle;sideswidelybordered inanterior
partsbutnarrowlysobehindmiddle, feeblyarcuatetonearapices,whicharenarrowly
roundedandrelativelypointed,preapica1emarginationslight;dorsumconvexthough
depressedonthediscbeforemiddle,steeplydeclivousatthesidesthoughtheapical
declivity israthergentle;only twoinnerstriaepresent,stria1almostcompleteand
coarselypunctatebeforemiddle,stria2shallower thant,nearlysmooth,andobsolete
near base and inapical two-thirds; apical striole short though sharply impressed,
hardlyarcuateinanteriorpart;intervalsflatevennearsuture,apicalcarinaobtuse;two
setiferousdorsalporespresentonthesiteofstria3ataboutbasalsixthandslightlybe-
forethemiddle;preapica1porepresentonapicaldeclivityjustbehindtheleve1ofthe
terminusofapicalstrioleandalittlemoredistant fromapexthanfromsuture.
Ventral surfaceglabrousandsmooth. Legsmoderateinlength,withthin tibiae
andtarsi;protibiaestraight,gentlydilatedtowardsapices,eachwithavestigialgroove
ontheexternal faceandafewshorthairsontheanterior face;tarsishort,tarsomere1
longer thantarsomeres2-3 combined butshorter than tarsomeres 2-4 combined in
mesotarsus,aslongastarsomeres2-4combinedinmetatarsus;in ,protarsomeres1
and2widelydilatedandstoutlyproducedinwardsatapices.
Malegenitalorgansmall thoughmoderatelysclerotized.Aedeagusone-thirdas
longaselytra,elongate,hardlyarcuateexceptattheapicalpart,nearlyparallel-sided
but ratherwidelydilatedattheapicalpart,andwidelymembraneousondorsum;basal
partnotcurvedventrad,being formedbytwonarrowlobesdeeplyemarginateateach
ventralsideandcompletelyconnectedattheproximalextremity;sagittalaileronsmall
butprotrudent,hyalineexceptforthecore;apicalpartdilatedinbothdorsalandright
lateralviews,curvedventracしandabruptlyendinginaveryshortapicallobe,whichis
directedtothele量androundedattheextremityindorsalview,smallandobtuseinfeft
lateral view; left aedeaga1wall notdilatedat theapical part;ventralmarginnearly
straightatmiddle,moderatelyemarginateattheapicalpart.Innersacminutelyscalyal
theapicalpart,bearingapairofpalp-likesclerotizedcopulatorypiecesjustinsideapi-
calorificeattherightdorsalandleftventralsides,eachofwhichisimperfectlyarticu-
latedatthemiddleandfoldable.Stylesslender.
Femaleunknown.
Typespecimen. Holotype: (5、,17-VI-2004,S.UENoleg.Depositedinthecol-
lectionoftheDepartmentofZoology,NationalScienceMuseum(Nat.Hist),Tokyo.
Type1o-ality. Mt.GuangwuShan,Zhaipoonthenortheasternside,1,780min
altitude,inNanjiangXianofSichuan,SouthwestChina.
Notes. Thesingleknownspecimenofthisextraordinarynewspecieswastaken
at thesideofaforestryroadonthenortheasternslopeofMt.GuangwuShan,which
liesat thesouthernsideofthewesternmostpartoftheDabaShanMountainsinNan-
NewTrechineBeetlesfromtheDabaShanMountains 271
3
ミ
ミ
・
0
5
Figs.2-5. MalegenitaliaofDabal,coitusspp;leftlateralview(2,4),andapicalpartofaedeagus,dorso-
apica1view(3,5). - 2-3.Deccen1t-1cusS.UENo,spnov.,fromMt.GuangwuShan(theleftcopu-
1aorypieceis folded). - 4-5. D /n,g1・op/1i/usS.UENo,sp nov., fromtheYidaoXiainBaokang
Xian.
272 Shun-IchiUENo
jiangXianofeasternSichuan.Theunpavedroadismadethroughabroadleavedforest
andhasashallowditchatthemountainside,whichistedbytricklesandeitherwetor
keepingaverylittle,slowlymovingwater.
At thecollectingsiteofthetrechinebeetle, thebottomof theditchwasthinly
coveredwithstagnantwaterandscatteredwithfallenleaves.Twospeciesofbembidi-
ineswerefoundhereandtherefrombeneathwet fallenleaves,andthetrechinebeetle
wasalsofoundfrombeneathafallenleafhalf-immersedinthewater.Itwasnotpartic-
ularlyagilewhenexposedjustlikesaproxy1ophi1ousmembersofthegenus-group.
Since furthersearchesdidnotyieldanymorespecimens,wewentbacktothat
particularspotonthenextdayandspentmanyhoursforseekingadditionalmaterial,
butalloureffortsendedinvain.
Dabatrechus/lygrophilusS.UENo,sp
nov
(Figs4-6)
Length: 4.15mm(fromapicalmarginofclypeustoapicesofelytra).
Closelysimilar ingeneralappearancetoD ecce11trlcus, but different fromit in
manydetails,aboveallintheshapeofpronotum.Incrediblydifferent fromDeccentr1_
cusinconformationofmalegenitalia,aswillbedescribedlater.
Bodyconstrictedbetweenprothoraxandhindpart,glabrousandpolishedondor_
sum but partiallypubescentonventer;microsculpturecompletelyvanished.Colour
verydarkbrown,shinyandnot iridescent,headandprothoraxalmostblack,elytra
darkbrownwithreddishbasalareas;clypeusandscapereddish;palpiandlegsyellow_
ishbrown.
HeadsimilartothatofDeccent1・Ious,buttheeyesarelargeranddevoidoffacets,
thegenaearelessoblique,the frontal furrowsaresubangulateatmiddle,andthean_
tennaeareshorter; frontal furrowssubangulateatmiddle,andmorewidelydivergent
infrontandbehind;supraorbitalareasrelativelyflat,bearingtwopairofsupraorbital
poresonlinesslightlydivergentposteriad;eyeshemispherica1lyconvextogetherwith
genae,butthe facetsaresodegeneratedexceptalongperipheries,thattheconvexsur_
facesofeyesaresmoothandshiny;genaeconvexattheanteriorportionsincontjnua_
tiOnoftheconvexityofeyes,rapidlyconvergentbehindtowardsneckconstrictjon,and
Completely glabrous;neckconstrictiondeeper thaninD. eccentricus;mental tooth
broad, roundedat thetip;antennaealittleshorter thaninD.eccentricus, reaching
basalthree-tenthsofelytra.
Pronotum cordate, obviously narrower thanin D. eccentricus, widest atabout
two-thirdsfrombase,andmorestronglycontractedatbasethanatapex;PW/Hw1.23,
PW/PL1.11,PW/PAca.141,PW/PB160;sidesrathernarrowlyborderedthrough_
out,Stronglyarcuatein front,lesssobehind,brieflybutdeeplysinuateataboutbasal
two-thirteenths,andbrieflyparallelinbasalarea;apexslightlyarcuate forwards,wjth
frontanglesroundedoff;baseobviouslynarrowerthanapex,PA/PBca.1.14,almost
straight atmiddle,andbrieflyandalittleobliquelyemarginateateachlateral on
NewTrechineBeetlesfromtheDabaShanMountains 273
Fig. 6. Dabat,,echushyg1・op/1ilusS.UENo,spnov., fromtheYidaoXiainBaokangXian
,
basalpeduncleextremelyshortandhardlyrecognizable;hindanglesnearlyrectangular
thoughbluntatthecorners;dorsumstronglyconvexandsmooth,steeplydeclivousin
marginalareas;medianlinefineexceptnearbasaltransverseimpression,whichiscon-
tinuous, somewhat sulciform and smooth, laterally merging into very small basal
foveae;basalareaverynarrow, formingacollar.
Elytrasubova1rather thanovate,evidentlywiderthanpronotum,widestslightly
beforethemiddle,andalmostequallynarrowed towardsbases andtowardsapices;
274 Shun-IchiUENo
EW/PW1.63,EL/PL2.60,EL/EW1.44;shouldersrounded,lesssalientthaninD.ec-
centricus;prehumera1bordersobviouslymoreobliquethaninD.eccentrlcusandonly
veryslightlyarcuate,withverynarrowmarginal gutter,whichdiminishesanteriorly
andreachesthesiteofthebaseofstria5;sidesratherwidelyre exed at thehumeral
partbutverynarrowlysoinapical two-thirds,gentlyarcuateforthemostpart,and
widelyandconjointlyroundedatapices,eachwithoutappreciablepreapica1emargina-
tion;dorsumstronglyconvexthoughsomewhatdepressedat theanteriorpartof the
disc,verysteeplydeclivousatthesidesbutrathergentlysoat theapicalpart;three
innerstriaedistinctlyimpressedonthedisc,allcoarselycrenulate,stria1almosten-
tire,2-3obsoleteinbothbasalandapicalareas;apicalstrictoshortthoughsharplyim-
pressed;intervalsslightlyconvexnearsuture,apicalcarinaobtuse;stria3withasingle
setiferousdorsalporeat two-fifteenthsfrombase;intheholotype,vestigeofasecond
poreperceptibleonstria3at four-ninths frombase, thoughnotsetiferous;preapica1
poreabsent.
Ventralsurfacemostlyglabrousandsmooth,buteachsternitebearsseveralhairs
atthemedianpart.LegsasinD.eccentricus,buttheprotibiaearedevoidofvestigial
grooveontheexternal face,andthetwoproximalprotarsomeresin areonlyweakly
dilatedandminutelydenticulateinwardsattheapices;protarsomere1in elongate,
1.5timesaslongaswide,protarsomere2aboutthree-fourthsaslongasl,alsoelon-
gateinthesameproportionasthelatter.
Malegenitalorganstrikinglydifferent fromthatofDeccentricus,alittlelarger
andalittlemoreheavilysclerotized.Aedeagustwo-fifthsaslongaselytra,twistedto
theright,lightlyarcuateasawhole,widelyconcaveattheleftsidebeforemiddle,and
thoroughlymembraneousondorsum,withtherightwallobviouslyloweredatthebase
ofapical lobe;basalpartstraightlyextended,openonbothdorsalandventralsidesbut
thelateralwallsaremuchhigher thaninD. eccentricus; basal orificedeeply emar-
ginateatthesides;sagittalaileronelongate,whollyhyaline;apical lobeelongate,pro-
trudingrightapically,andnarrowlyroundedatthetipindorsalview,graduallytapered
towardsbluntextremityandsomewhatreflexedattheterminalportioninlateral view;
ventralmarginnearlystraightatmiddleinprofile.Innersacrathercoarselyscalyatthe
apicalpart,bearingalargehyalinecopulatorypiece,whichisabout two-seventhsas
longasaedeagus,spoon-like, rightdorsallyconcaveinapicalhalf,andobliquelylying
justinsideapicalorifice.Stylesfairlywideinbasalhalvesandwithshortnarrowapi-
calparts,leftstyleobviously longerthantherightandbearingshortventralapophysis,
eachprovidedwith four,rathershortapicalsetae.
Femaleunknown.
Typespecimen. Holotype: 11-VI-2004,M.SAT01eg.Depositedinthecol-
,
lectionoftheDepartmentofZoology,NationalScienceMuseum(Nat.Hist),Tokyo.
Type10_ality. YidaoXia,580minaltitude,atHoupingCunofHoupingZhenin
BaokangXian,westernHubei,West-centralChina.
Notes. Inordinarytrechinetaxonomy,thisnewspeciescouldberegardedasbe-
longing toagenusdifferent fromDabatrechus,mainly inviewofthemarkeddiffer-