A New Genus Species of Melolonthid Beetle（Coleoptera，Scarabaeidae)from Borneo

5 Pages·1993·4.5 MB·English

by  Itoh, T

Elytra,Tokyo,21(1):77-81,May15,1993
A NewGenusandSpeciesof Melo1onthineBeetle
(Coleoptera, Me1o1onthidae) fromBorneo
Takeshi ITCH
Higashi-naruo-cho2-l-l3-2I2,Nishinomiya,Hyogo,663Japan
Abstract Anewme1o1onthid beetle,Wadaiakaoruigen etsp nov. is described
fromBorneo. ItbelongstothetribeMe1o1onthiniandissomewhatrelatedtoStepha-
nopho/1sme1o1ont1/oldes(BRENsKE,l892)fromthePhilippines.
Recently, l had an opportunity toexamine an interestingme1olonthine beetle
from thenorthern and western areasof Borneo. Afteracareful study, I havecome
to theconclusionthat itshouldbelongtoanewspeciesofanewgenus,thoughitis
rather allied to Stephanop/1o/1sme/o/ont/1oldes (BRENsKE, 1892). In this paper, l am
goingtodescribeit underthenameofWadalakao・,u1T.ITCH,gen etsp nov.
Beforegoingfurther, IwouldliketoexpressmyheartygratitudetoProf.Masa-
takaSATofor hiscriticaladviceonthemanuscriptof thispaper,aridalsotoMessrs.
Kaoru WADA, Atsushi KAToH and Masamichi YAGI for their kindness in offering
invaluablematerials.
WadaiaT. ITCH,gen
nov.
Male. Bodyelongateoval andconvex. Dorsal surfacerelatively denselyand
regularlycoveredwiththincreamybrownscales. Clypeusstronglynarrowedanteriad
andremarkably reflexedlikeascreeninapical half. Antenna9-segmented, theclub
composed of 3segments. Menurn trapezoidal, bluntly produced at the centre of
anteriormargin. Eachelytronbearingadevelopedmarginalmembrane fromlateral
margintosuturaledge.
Ventral surface hairyexcept for abdomen. Mesosternum without a produced
process. Abdomen also relatively densely and regularlycovered with thincreamy
brownscalesexcept for themedic-basalareaof2ndsterniteandjustalongsutures;
lateral sidegenerallydevoid ofcreamy brownmaculationscomposedofscalesbut
rarelywithobscureones.
Legs relatively slender. Protibia tridentate, the basal denticle obtuse. Longer
oneofthetwoapicalspinesofmetatibiamuchlongerthantheisttarsalsegment. The
1stto4thsegmentsofeachtarsusprovidedwith tuftsbeneath. Metatarsusslender,a
littlelonger thanmetatibia. Clawwithanapical,anobliquelydirectedmediananda
verysmall basal teeth.
Parameresofmalegenitaliacomposedofapairofouterlargerandapairofinner
78 Takeshi ITCH
・
1 2
Fi9s. 1-2. Wadalakaorulgen etsp nov.;1,habitus(male);2,ditto(female)
smaller branches.
Female. Clypeus lessreflexed, bilobateandclearlyemarginateat thecentreof
anterior margin. Legsmorerobust,metatarsus aboutas longasmetatibia. Claw
withamorestronglybent apicalandaverticalmedianteeth.
Typespecies:Wadaiak;aoru1T.ITCH,sp
nov.
This genus is distinguishable from Stephanopholis BRENsKEand Rhabdopholis
BURMEISTER by the 9-segmented antenna, longer metatarsus, tridentate protibia,
branchedparameresofmalegenitalia,obliquelydirectedmediantoothofmaleclaw,
andso on.
W,adaiakaorutT. ITCH,sp
nov.
(Figs. 1-10)
Bodylength: 22.7-29.6mm.
Male. Bodyblackishbrown torufousblackishbrown.
Clypeusstronglyreflexedat apexandgentlyemarginateat theanteriormargin,
fronto-clypea1sutureclearandbiarcuate. Fronswideandsparselypunctate. Interval
betweeneyes0.70-0.73(0.72onanaverage)timesasbroadasheadwidth. Occipital
Fi9s. 3-10. Wadalakaoruigenetspnov. - 3,Maleclypeus;4,mentum;5,occipitalarea;
6,rightmaxillain lateralview(a),andmaxillarygaleaindorsalview(b);7,membraneof
elytra1margin(El:elytron,Pyg:pygidium); 8, leftmetatibiaandmetatarsusinmale;g,
malegenitaliaindorsalview(a),andlateralview(b)(scale:1mm); 10,clawofforelegin
male(a)andfemale(b).
NewMelo1onthineBeetlefrom Borneo 79
一 、、
'
. / .l.'.''.1・ .. ,・.
5
(a)
(a)
b )
1 0
80 Takeshi ITCH
areacoveredsparselywithfinepuncturesbehindvertex. Antennalclubnearlyaslong
asfootstalkandalso1.7-2.1(1.9onanaverage)timesaslongas4precedingsegments
combined. Maxillarygaleaprovidedwith4denticlesandthelastsegmentofmaxillary
palpusspindle-shaped.
Pronotumconvex,0.59-0.62(0.60onanaverage) timesaslongasbroad;anterior
marginrimmedonlynearanteriorangles; lateralmarginverygentlycurvedalittle
behind themiddle, antero-1atera1margin andpostero-1atera1onestraight; anterior
anglerectangular,posterioroneclearlyobtuse;discsparselypunctate,withanobscure
groove along themedian line. Scutellum triangular inoutlineand rounded at the
apical margin. Eachelytron with4costaeand asutural one, 3rd onesometimes
obscure.
Presternal keel triangular and convex. Metasternum sparsely punctate in the
medianareaanddenselysoat thelateralsides,andmoredenselyhairyat thelateral
sides. Metasterna1medianlineevident. Sixtha1.dominalsegmentv、,holly furnished
withshort thinnerscalesthan theothersegmentsandwithlongbrownpubescence
near theapicalmargin. Pygidiumflatandroundedatapicalmargin.
Metafemurconvex,moderatelyswollenat themiddleand0.30-0.31(0.30onan
average) timesaswideas long. Metatibiamoderatelyswollenat theapexandpro-
videdwitharowofrelativelylongbristlesalongtheinnermarginasinmesotibia.
Female. Dorsal and ventral surfacescovered with thicker scalesthan in male.
Intervalbetweeneyes0.72 timesasbroadasheadwidth. Antennalclubshorter,1.6
timesaslongasthe4precedingsegmentscombined. Pronotum0.61 timesas long
as broad. Eachelytronwith4clearcostaeandasutural one. Scales on abdominal
sternitesthicker thaninmale,eachsimilar toaricegrain. Metafemurmoreswollen,
0.43timesasbroadaslong.
lsfr1加f,on. Borneo.
Holotype: 1 , Mt. Kinabalu, N. Borneo, 21-IV-1988. Allotype: 1 ne ar
,
Keningau, N. Borneo, 13-IV-1988. Paratypes:1 ,samedataas for theholotype;
] Kimanis Road, Keningau, N. Borneo, 2-V-1988, M. YAGI leg ; 1 Crocker
, ,
Range, Sabah, N. Borneo, 29-III-1989; 2 Mt. Bawang, West Kalimantan,
,
Borneo, III-1990, N. NlsHIKAwAleg.
Theholotypeandallotypearedepositedinthecollectionof theDepartment of
Zoology,NationalScienceMuseum(Nat.Hist),Tokyo,2paratypesareinthecollec-
tion of the Zoological Museum (NaturalHistory) of Humboldt University, Berlin,
and theothersareinmycollectionandinProfessorSATo's.
Remarks. Judgingfromreflexedclypeus inthemale, antennal clubcomposed
of3lamellatesegmentsinbothsexes,mesosternitewithoutaprocess,abdomenwithout
obscuremaculationsofscalesat thesides,andsoon, thepresentnewspeciesjsrather
similartoStephanopho/1sme/ole?'t/1oides(BRENsKE)fromthePhilippines.
ThenewspeciesisdedicatedtoMr. KaoruWADA.
NewMe1o1onthineBeetlefromBorneo 81
要 約
伊藤 武: ポルネオ島のコフキコガネ科の1新属新種. - ポルネォ島北部と西部で得られたコフ
キコガネ族に属する1新属新種を記載し, Wadaiakaoru,という新名を与えた. 本不?1は, 強く反り
返った雄の頭盾, ?1性l?llとも触角集1,節が3節からなること, 中胸腹板突起が不明瞭であること,月?側
部にf?11-からなる明 な白斑がないことなどから, Slephanopholis属に近縁であると考えられ, ま
た, アフリカのRkabdoPholis属にも近似しているが,角?角が9節であること, より長い後付節, 前
J ,l 座節に3外11景が存在すること ｮll交尾器の枝分かれした側1- や雄の爪の, 央, 1塩-がやや斜めを向いて
る構造などの特徴から, 別属とするのがよいと考え, 新属を設定した. ;重f名は, 和Il 董氏に献名し
た.
References
BRENSKE, E.,1892. UebereinigeneueGattungen undArtenderMe1oIonthiden. Ent.Nachr.,10:
151-160.
- 1896. NeueMelo1onthidenausAfricaundAsien. Stett ent.Ztg.,57:178-205.
BuRMEISTER,H.,1855. HandbuchderEntomo1ogie 4(2):x十570pp. Reimer,Berlin.
ITCH,T.,1992. AnoteonSte1o/1anopholisme1o1onthoides(BRENsKE,1892)(MeloIonthidae,Melolon_
thinae,Melolonthini). Lame11ic・ornla,8:7-8. (InJapanese)
PERINGuEY, L., 1904. Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and
Scarabaeidae). Tra,ts.S. Ajfii・.Soc.,13:1-293.