A new genus and two new species of Physocrotaphinae from New Guinea (Insecta, Coleoptera, Carabidae)

2004·3.3 MB·English

by  M Baehr

SPIXIANA 27 9 107-113 München, Ol. Juli 2004 ISSN 0341-8391
A new genus and two new species of Physocrotaphinae
from New Guinea
(Insecta, Coleoptera, Carabidae)
Martin Baehr
Baehr, M. (2004). A new genus and two new species of Physocrotaphini from
New Guinea (Insecta, Coleoptera, Carabidae). -Spixiana 27/2: 107-113
Schneien, gen. nov. and two new species, S. monstrosa from Irian Jaya and
S.drumoniifromPapuaNewGuineaaredescribed.Thenewgenusisdistinguished
fromallknownphysocrotaphinegenerabyitsmassivebodyshape,ver\'shortand
stout legs, short and stout antenna with peculiar impressed apical antennomeres,
multisetoseclypeus, deeplyimpressedanteriortransversesulcusofthepronotum,
and large, more or less tectiform elytra. Also Pogonoglossus arfakensis Baehr is
transferred to thenew genus.
Dr. Martin Baehr, Zoologische Staatssammlung, Münchhausenstr. 21, D-81247
München, Germany; e-mail: [email protected]
Introduction numberofundescribed orevenunrecorded species
isperhaps quite substantial, because specimens are
Therathersmall subfamily (ortribe)Physocrotaph- rarely collected and most species seem to occupy
inae (-ini) (according to Moore 1998 this name has ratherrestrictedranges.Both,thedifficultiesinsam-
priority over the more commonly used name Hel- pling and the apparent local distribution, are per-
luodinae)isdistributedintheOrientalandAustral- haps caused by the rather secrete way oflife, since
ian regions from the southern and eastern parts of most species ofPhysocrotaphinae seem to live- or
India, in the north-west, through Sri Lanka, Indo- at least have been found - under bark or in rotten
china, the Indonesian Archipelago to the Philip- wood of rain forest trees. A number of species,
pines, NewGuinea, New Britain, and northern and however, are only known from sampling at light,
north-eastemAustralia, inthesouth.Thesubfamily and hence, virtually nothing is known about their
includes three genera, namely the large genus way oflife.
Pogonoglossus Chaudoir that is distributed over al- From New Guinea, thus far 12 species of the
mostthewholerängeofthesubfamily,themonoba- most speciose genus Pogonoglossus Chaudoir were
sicPhysocwtaphiisParryfromSriLanka(P. cci/lonicus described (Darlington 1968, Baehr 1987, 1995), and
Parry), and Hclluodcs Westwood with two species mostareyetrecorded from feworevenSinglespec-
occurring in Sri Lanka and southern India. The gi- imens. Therefore, the detection of two additional
gantic Holopoucnis godc^fi-oi/i (Fairmaire) from New new physocrotaphinespecies in NewGuinea isnot
Britain, although originally described as a member too surprising. In many respects, however, both
ofthe subfamily Lebiinae, probably belongs also to newspeciesareoutstandingandnotcomparableto
Physocrotaphinae. Unfortunately, the unique spe- anyone of the described species of Physocrotaphi-
cimen was destroyed during World War II, so this nae within and outside New Guinea, except forthe
question is not solvable as long as no new material large Pogonoglossusarfakensis Baehr, and thereforea
is available. new genus is being described to accommodate this
Altogether, less than 40 physocrotaphine spe- and both new species.
ciesweredescribed until now(Lorenz1998),butthe
107
Fig. 1. Schuelea monstrosa, spec. nov. 3aedeagus, parameres, and genital ring. Scale: 1 mm.
Fig.2. Schueleadrumonti, spec. nov. $ stylomeres 1 and2. Scale: 0.5mm.
Measurements remarkably protruding eyes, short and wide, unu-
sually massive mandibles, deep anteriortransverse
Measurements have been taken under a Stereo micro- sulcus onpronotum, and rather tectiform elytra.
scope using an ocular micrometer. Length has been
measured from tip of labrum to apex oi elytra, hence Type species. Schuelea monstrosa, spec. nov., by
measurements may slightly differ from those of other present designation.
authors. Some width/length ratios have been taken in Etymology. Thenew genusisnamed inhonourofMr.
thesamemannerasinBaehr(1988,1993,1995).Itshould PeterSchule, collector ofone ofthe new species.
be noted that length of pronotum was measured from
apex of anterior angles to the most advanced point of Distribution. New Guinea.
base.
Schuelea monstrosa, spec. nov.
Location of types Figs 1, 3
The uniqueholotypes ofthe three species are stored in Types. Holotype: 6, 11.7.1996, 15, Schule/Stuben,West
Museum für Naturkunde der Humboldt-Universität, Papua Fakfak, Mambuni-Buni, Garten (ZSM-CBM).
Berlin (MNHB), Institute Royal des Sciences naturelles
deBelgique, Bruxelles(IRSNB), and intheworkingcol- Diagnosis. Largespecies, distinguishedfromboth,
lection of the author in Zoologische Staatssammlung, S. arfakensis (Baehr) and S. drumonti, spec. nov. by
München (ZSM-CBM). angulate orbits, basally deeply excised external
marginofleftmandible,lessshortandwideprono-
tum with wider base, acute, markedly projecting
Schuelea, gen. nov. anterior angles and characteristicaliy upturned lat-
eral parts of anterior margin, and decidedly tecti-
Diagnosis. Genus of subfamily Physocrotaphinae. form elytra.
InmanycharacterstatessimilartogenusPogonoglos-
sus Chaudoir, in particular with respect to shape Description
and structure of glossa and paraglossae. Distin- Measurements. Length:15.5mm;width:5.5mm.
guished from all other described genera by stout Ratios. Width/lengthofpronotum: 1.33;base/apex
body, very short and stout legs, unusually short of pronotum: 1.12; width of pronotum/width of
antenna bearing characteristic impressions at the head:1.26;widthofpronotum/widthofelytra:0.86;
baseoftheinnerand outersurfacesofantennomer- length/width of elytra: 1.68; length/width of 10*
es 5-11, multisetose clypeus, small though laterally antennomere: 1.2.
108
3 '^ 4
Figs 3,4. Habitus. 3. Schudea monstrosa, spec. nov. Length: 15.5 mm. 4. Scinielea drumouti, spec. nov. Length:
14.5mm.
Colour. Glossyblack,onlytheanteriormargins lateral seta far longer than the inner setae. Mandi-
of clypeus and labrum, and the tips of the palpi bles comparatively short and wide, massive, inner
reddish. borderalmoststraightforalongdistance,onlvnear
Head. Large and massive, but slightly narrovv- apex incurved. Lower lateral margin ofleft mandi-
er than pronotum, ver}' wide between eyes. Frons ble deeply excised at a short distance from base,
with two deep, elongate, irregularly sinuate im- here scrobe absent. Lateral margin above excision
pressions,neckseparatedfromfronsbyaverydeep with a very faint ridge. Right mandible without
sulcus, thusUppersurfacever)'uneven. Eyessmall, such excision, but both mandibles with a short,
semicircular, laterally remarkably projecting. Or- triangulär excision immediately near base. Palpi
bits slightlv shorter than eyes, obtusely angulate, rather short and stout, with sparse and extremely
posteriorly bordered by a ridge, behind vvhich is a shortpilosity.Mentum withtriangulärtooth,with3
very coarsely punctate sulcus. Anterior supraor- elongate setae on either side. Submentum with a
bital setasituated nearinnermarginofeyeatabout transverse row of six elongate setae. Lateral and
middle ofeye, posterior supraorbital seta posterio- lower surface oforbits shortly setose. Glossa rather
mediallyremoved fromeyeandsituatedatanterior short, apicallv transverse, bisetose, paraglossaeapi-
margin of transverse sulcus. Clypeus with four to cally separated from glossa, membraneous, very
fiveelongate setaeon either side, in addition to the narrow,justslightlysurpassingglossa.Laciniamas-
erect pilosity. Labrum 7-setose, one of the parame- si\'e, elongate, with a dense row of verv elongate
dian setae apparentlv doubied at the right side, the teeth. Galea narrow, innersurface slightly concave.
109
Antenna short, just surpassing base of pronotum, someres of median and posterior tarsi, tarsomeres
pilosefrombasalantennomere, scapus rather short almostaswide aslong. Especiallyonlowersurface
andstout,3"*and4*antennomeresmoderatelyelon- and at apex tibiae densely setose. Upper surface of
gate. Antennomeres 5-11 little longer than wide, tarsirathersparselypilose,lowersurfaceofanterior
oval-shaped, depressed. Upperand lowerrimwith fourtarsomeresdenselypilose, 5*tarsomere setose
a narrow, remarkably chagreened field on either underneath. Male protarsus without special vesti-
side that is rather densely pilose. The broad sides ture.
characteristically bi-impressed at base. Upper sur- 6 genitalia (Fig. 1). Genital ring oval shaped,
faceofheadwithsomerugosities and coarsepunc- slightly asymmetric, very strongly sclerotized. Ae-
tures, apparently devoid ofmicroreticulation, rath- degus rather stout, gently curved, lower surface
er sparsely pilose, surface fairly glossy. concave with several irregulär shallow, transverse
Pronotum. Large, though comparatively nar- sulci. Orificiumelongate, almostSymmetrie, situat-
row,littlewiderthanhead,widestatanteriorthird. ed onUpperside,bearingtwoelongate, sclerotized
Apexverydeeplyexcised,anterioranglesmarkedly ligulae at upper surface. Inner sac simply folded,
angulate and projecting. Apical excision laterally rather Symmetrie. Left paramere large, with ob-
withalarge,upright,aboutquadrangularplatethat liquely cut apex. Right paramere with small, nar-
conspicuously Interrupts the excision. Apical mar- row sclerotized upperpart and elongate, little scle-
ginwithoutanypilosity. Lateralborders anteriorly rotized base.
strongly convex, in posterior third with elongate 9 genitalia. Unknown.
sinuation. Basewiderthan apex, inmiddle slightly Variation. Unknown.
concave, laterally straight though oblique. Basal
anglesangulate,almostrectangular,withatinyden- Distribution. Western part ofIrianJaya (now 'Pa-
ticle. Apex not bordered, base laterally bordered. pua'),NewGuinea.Knownonlyfromtypelocality.
Anteriormarginalsetasituatedjustbehindanterior
CoUectingcircumstances. Largelyunknown,though
third, atmargin, posteriormarginal seta situatedat
thespecimenwascollectedin 'garden',presumably
basal angle. Lateral explanation wide and deep,
at low altitude.
lateralmarginconspicuouslyupturned.Medianline
faintly impressed, neither attaining apex nor base. Etymology.Thenamereferstotheoutstandingshapeof
Anterior transverse sulcus deep, about semicircu- the anterior angles ofthe pronotum.
lar. Basal grooves deep, circular, in middle near
base with a rather deep transverse furrow. Surface
remarkablyconvexoneitherside.Puncturationand Schuelea drumonti, spec. nov.
pilosity fairly sparse, microreticulation absent, sur- Figs2, 4
face rather glossy.
Elytra. Fairlyelongate, laterally absolutelypar- Types. Holotype: 9, Coli. I.R.Sc.N.B. Canopy mission
allel, without any sinuation in anterior third. Hu- P.N.G. Madang province, Baiteta, FOG AR 3-10 4.V.
merusevenlyrounded,barelyprojecting.Apexgen- 1995, Leg. OlivierMissa (IRSNB).
tly sinuate, with wide membraneous area. Surface Diagnosis. Largespecies,distinguishedfromS. mon-
remarkably tectiform, in particular towards apical strosa, spec. nov. by rounded orbits, barely excised
third. Marginal Channel very narrow throughout. externalmarginofleftmandible, shorterandwider
Striae lightly impressed, impunctate, intervals gen- pronotumwithnarrowerbase,roundedandfarless
tly convex, with sparse, irregularly uni- to bipunc- projectinganterioranglesandnormal shaped ante-
tatepuncturationandpilosity.3"*intervalwiththree riormargin,andlesstectiformelytra.Mainlydistin-
setiferouspunctures,setaeerectanddistinctlylong- guished from similar and closely related S. arfaken-
er than pilosity. Microreticulationextremely super- sis(Baehr)bylonger,moretectiform,andlessdensely
ficial and fine, visible only under high magnifica- pilose elytra, and slightly narrower base ofprono-
tion,consistingofrathertransversemeshes.Surface tum.
rather glossy. Lateral border not serrate, sparsely
pilose. Marginal setae elongate. Fully winged. Description
Lower surface. Thorax comparatively sparsely Measurements. Length:14.5mm;width:5.2mm.
punctateandsetose, abdomenwithdenserpilosity. Ratios. Width/lengthofpronotum: 1.57;base/apex
Metepistemumelongate,morethantwiceaslongas of pronotum: 0.90; width of pronotum/width of
wide. Terminal sternite in male on either side with head: 1.17;widthofpronotum/widthofelytra:0.83;
3-4 elongate setae near margin. length/width of elytra: 1.70; length/width of 10"^
Legs. Remarkably shortand stout. Protibiasul- antennomere: 1.2.
cate on upper surface. Apart from 1"' and 5'^ tar- Colour. Glossy piceous-black, labrum, anterior
110
margin of clypeus, tips of the palpi reddish, elytra angular,withasmallthoughdistinctdenticle.Apex
slightly lighter than fore body. notbordered,baselaterallybordered.Anteriormar-
Head. Large and massive, slightly narrower ginal seta situated just behind anterior third, at
thanpronotum,verywidebetweeneyes.Fronswith margin, posterior marginal seta situated at basal
two deep, short, irregularlv circular impressions, angle. Lateral explanation fairly wide and deep,
neck separated from frons by a very deep sulcus, lateral margin ratheruptumed. Median line faintly
thus Upper surface very uneven. Eyes small, semi- impressed, neither attaining apex norbase. Anteri-
circular, laterally remarkably projecting. Orbits ortransversesulcusdeep, aboutsemicircular. Basal
slightlyshorterthaneyes,evenlyconvex, posterior- grooves deep, circular, in middle near base with a
lygently surpassing intoneckvvithoutany ridgeor rather deep transverse furrow. Surface remarkably
sulcus. Anterior supraorbital seta situated near in- convex on either side. Puncturation and pilosity
ner margin ofeye atabout niiddle ofeye, posterior fairlysparse,microreticulationabsent,surfacehighly
supraorbital seta posterio-medially removed from glossy.
eye and situated at anterior margin of transverse Elytra. Fairlyelongate, laterally absolutelypar-
sulcus. Clypeus with four elongate setae on either allel, without any sinuation in anterior third. Hu-
side, in addition to the erect pilosity. Labrum 6-se- merusevenlyrounded,barelyprojecting.Apexgen-
tose, the lateral seta far longer than the innersetae. tly sinuate, with wide membraneous area. Surface
Mandibles comparati\'ely short and wide, massive, fairly convex though little tectiorm, not even in
inner border almost straight for a long distance, apical third. Marginal Channel ver\' narrow throu-
only near apex incurved. Lower lateral margin of ghout. Striae moderately impressed, impunctate,
left mandible gently bi-excised at a short distance intervals fairly convex, with sparse, though rather
frombase,scrobeherenarrow.Lateralmarginabove coarse, irregularly uni- to bipunctate puncturation
excision with a conspicuous, elongate ridge. Right and pilosit)^ 3"^inter.^alwith threesetiferouspunc-
mandible without such excision, but both mandi- tures, setae erect but short, difficult to recognize
bles with a short, triangulär excision immediately within the pilosih'. Microreticulation extremely su-
nearbase. Palpi rather short and stout, with sparse perficial and fine, barely recognizable even under
andextremelyshortpilosity.Mentumwithtriangu- high magnification, consisting of rather transverse
lär tooth, with 3 elongate setae on either side. Sub- meshes. Surface glossy. Lateral border not serrate,
mentumwithatransverserowofsixelongatesetae. sparsely pilose. Marginal setae elongate. Fully
Lateral and lower surface of orbits shortly setose. winged.
Glossarathershortapicallytransverse,bisetose,pa- Lower surface. Thorax comparativeh' sparsely
ragiossaeapicallyseparatedfromglossa,membrane- punctateand setose, abdomen with denserpilositv'.
ous, ver\' narrow, but slightly surpassing glossa. Metepisternumelongate,morethantwiceaslongas
Lacinia massive, elongate, withadenserowofvery wide.Terminalsterniteinfemaleoneithersidewith
elongate teeth. Galea narrow, inner surface slightly 3-4 elongate setae near margin.
concave.Antennashort,justsurpassingbaseofpro- Legs. Remarkably shortand stout. Protibiasul-
notum,pilosefrombasalantennomere,scapusrath- cate on upper surface. Apart from P' and 5* tar-
ershortand stout, 3"^ and4'*' antennomeresmoder- someres of median and posterior tarsi, tarsomeres
atelyelongate. Antennomeres5-11 littlelongerthan almostas wide as long. Especially on lowersurface
wide,oval-shaped,depressed.Upperandlowerrim and at apex tibiae densely setose. Upper surface of
with a narrow, remarkably chagreened field on ei- tarsi rathersparselypilose,lowersurfaceofanterior
ther side that is rather densely pilose. The broad four tarsomeresdensely pilose, 5'^ tarsomeresetose
sides characteristically bi-impressed atbase. Upper underneath. Vestiture of male protarsus unknown.
surface of head with some rugosities and coarse 6 genitalia. Unknown.
punctures, apparently devoid of microreticulation, 9genitalia(Fig.2). Stv'lomere1 triangulär, with
rather sparsely pilose, surface glossy. 2-3 nematiform setae at median side of apical rim.
Pronotum. Large,comparativelyshortandwide, Stylomere2ofsemilunarshape, ventro-lateral mar-
thoughbutlittlewiderthanhead, widestinfrontof gin slightly serrate, with four very small ensiform
anterior third. Apex rather deeply excised, anterior setae along margin. Median surface nearapex with
angles rounded, not much projecting. Apical exci- anelongategroovefromwhicharathershortnema-
sion without any special characters. Apical margin tiformsetaarises. Alsowithoneortwonematiform
laterally with short pilosity. Lateral borders anteri- seta arising from median surface nearbase. Lateral
orly rather convex, in posterior third with fairly plate with rather sparse, ver)' elongate nematiform
shallow, elongate sinuation. Base narrower than setae
apex, in middle slightly concave, laterally straight Variation. Unknown.
though oblique. Basal angles angulate, almost rect-
111
Distribution. North-eastern Papua New Guinea. Keyto the species ofthe genus Schuelea
Known only from type locality.
Orbits angulate;lateralmarginofleftmandible
Collecting circumstances. The holotype was cap-
nearbasedeeplyexcised;anterioranglesofpro-
turedby 'canopyfogging'inrainforest,probablyin notum acute, much projecting, base wider than
lowland. It is not known, however, whether the apex, apical margin laterally conspicuously
specimenactuallywasfoggedfromtheuppercano- blade-like upturned; elytra markedly tectiform.
py, or from lowerbranches, or from the trunk of a WesternIrianJaya monstrosa, spec. nov.
tree.
Orbits rounded; lateral marginofleftmandible
Etymology. NamedinhonourofMr.AlainDrumontof nearbasebarelyexcised;anterioranglesofpro-
IRSNB, Brüssels who kindly made accessible to me a notum rounded, far less projecting, base nar-
large sample of New Guinean ground beetles which rower than apex, apical margin laterally not
includes the new species. upturned; elytra more depressed 2.
Largerandwiderspecies,ratio1/wofelytra1.6;
Schuelea arfakensis (Baehr), comb. nov. pronotum with slightly wider base; elytra de-
pressed, barely tectiform, median intervals in
Pogonoglossus arfakensis Baehr, 1987: 363; Lorenz 1998: apical half irregularly bipunctate and -pilose.
483. Vogelkop, westem IrianJaya arfakensis (Baehr)
Types. Holotype: 9, Arfakgeb. Siwi, 800m, 4.6.28, E. Smaller and narrower species, ration 1/w of
MAYRS. G. (MNHB). elytra1.7;pronotumwithslightlynarrowerbase;
elytra less depressed, perceptibly tectiform,
Nspoetcei.esDaulseobteoloitnsgsshtaopteheannedwsgternuuctsurSec,huetlheiasalnadrgiet pmuendcitaanteiantnedrv-aplisloisne.aNpoircatlh-heaalsfteirrrnegPualpaurlayNueniw-
is herewith transferred to this genus. Apparently, Guinea drumonti, spec. nov.
the holotype still is the Single recorded specimen.
Diagnosis. Largespecies,distinguishedfromS. mon-
sirosa, spec. nov. by rounded orbits, barely excised Remarks
externalmarginofleftmandible, shorterandwider
pronotumwithnarrowerbase,roundedandfarless Although the three species clearly belong to the
projectinganteriorangles andnormal shaped ante- subfamily Physocrotaphinae, they differ frommost
riormargin,andlesstectiformelytra.Mainlydistin- other species intheir stout, evenbulkybody shape
guishedfromsimilarandcloselyrelated S. drumon- andtheshortlegsandantennae.Whereas S.arfaken-
ti, spec. nov.by shorter,barelytectiform, and more sis (Baehr) and S. drumonti, spec. nov. in certain
densely pilose elytra, and slightly wider base of charactersdonotexhibitexceptionalmorphological
pronotum. Some measurenemts and ratlos are in- States, S. monstrosa at the firstglance Stands outby
cluded in the tablebelow. itsvery stränge shape ofthe prothorax, butalsoby
its tectiform elytra and the remarkably angulate
orbits. Certainly, S. arfakensis and S. drumonti are
Comparison of measurements and ratlos still closely related, whereas S. monstrosa exhibits a
in the genus Schuelea very stränge and highly aberrant morphological
development.
(w:width,1:length,b:base,a:apex,pr:pronotum,h:head, Together with the two new species, now 14
el: elytra) physocrotaphine species are recorded from New
Guinea and this island more and more proves as
N length w/lpr b/apr wpr/h wpr/el 1/wel one ofthestrongholdsofthesubfamily.Originally,
asBaehr (1988) and Moore (1998) stated, Physocro-
taphinaeisanOrientalsubfamilythatprobablyorig-
arfakensis 1 16.8 1.58 0.94 1.14 0.79 1.60 inated somewhere in southem Asia and so farhad
dmroinismtornotsia 11 1145..55 11..5337 01..9102 11..2167 00..8836 11..7608 its highest degree of taxonomic diversity in Suma-
traandJava(Bouchard1903,Andrewes1937),where-
as the Asiatic mainland and also Borneo and the
Philippine Islands apparently are much less speci-
ose(Jedlicka1963,Stork1986).However,theseState-
ments are based on quite limited knowledge and
have tobe regarded withsome reservation.
112
AccordingtotheopinionofMoore(1998)which Acknowledgements
was supported by larval characters, a sister-group
relation exists betvveen Phvsocrotaphinae and An- I am grateful to Mr. A. Drumont (Bruxelles), Mr. P.
thiinae which means thatboth tribes are next relat- Schule (Denzlingen), and Dr. M. Uhlig (Berlin) for the
ed and probably' stem from a common ancestor. kind loan or gift ofmaterial or t}'pes.
Because Anthiinae are an African group of quite
clearGondwananorigin, this would imply that the
strictly Oriental-Australian Physocrotaphinae like- References
wise belong to the old Gondwanan faunal element
and probably originated on one of the previous Andrevves, H. E. 1937. On the species of Pogonoglossus
terranes of Gondwanan origin that subsequently found inJava and Sumatra.-Bull. See.ent. Fr. 42:
152-156
were attached to the Laurasian continent. Future
phylogenetic examinations should clarify which BaehrN,eMw.G1u9i8n7.eaP(oCgoolneoogpltoesrsau,sCaarrfaabkeindsaies,sHpe.llnuoovd.infareo)m.
physocrotaphine genus is the most plesiot\'pic one — -Dt. ent. Z., N. F. 34: 363-366
which, intum, couldsuggestonwhichlandmassof 1988. Revision of the Australian species of the ge-
Gondwanan origin this subfamily originated. De- nus Pogonoglossus Chaudoir (Insecta: Coleoptera:
pendent on the phylogenetic Status of the genera, Carabidae: Helluodinae). - Invertebr. Taxon. 2:
those terranes that developed either into Sri Lanka — 961-972
(Ceylon) and/or southern India, or into later Su- 1993. A new species of Pogonoglossus Chaudoir
matra and/orJava would come into quesrion. from Australia (Insecta, Coleoptera, Carabidae,
Certainlv, physocrotaphine Stocks subsequent- — Helluodinae). - Spixiana 16: 141-144
ly migrated or drifted to the southeast to reach 1995. A peculiar new species ofthe genusPogono-
those Island arcs that later combined to what is glossusChaudoirfromNewGuinea(Insecta,Coleo-
present New Guinea, and eventually they also ptera, Carabidae, Helluodinae).-Spixiana 18: 255-
258
reached northern and northeastem Australia. The Bouchard, 1903.ATetudedesCarabiquesdeSumatra.
high taxonomical and morphological diversity of -Ann.J.Soc. ent. Fr. 72: 177-180
Physocrotaphinae in the Papuan-Australian area Darlington, P. J. Jr. 1968. The Carabid beetles of New
demonstrates that this Immigration must have oc- Guinea. Part III. Harpalinae continued. Perigonini
curred ratherearly, or that the taxonomic diversifi- to Pseudomorphini. -Bull. Mus. Comp. Zool. 139:
cation at least has proceeded rather rapidly. Only 1-253
the colonisation ofAustralia probably was a rather Jedlicka,A. 1963. MonographiederTruncatipennenaus
recent event which is demonstrated by the occur- Ostasien. Lebiinae - Odacanthinae - Brachvninae
rence of still identical species in New Guinea and (Coleoptera, Carabidae). - Ent. Abh. Ber. Staatl.
northern Australia (Baehr 1988). Mus. Tierkunde Dresden 28: 269-579
Due to thestill lackingknowledgeofthephylo- Lorenz, W. 1998. Systematic ListofextantGround Bee-
tles of the World (Insecta Coleoptera "Geade-
geneticrelationswithinPhysocrotaphinae,atpresent
itisimpossibletodecidewithwhichNewGuinean pinhaaeg,a"Ci:cTinrdaeclhiynpaae.chRihdyaseodainddaeC)a.ra-bTiudtazeiningc,l.pPraiunstesd-
orevenextra-NewGuineanspeciesorspecies-group by the author. 502 pp.
ofthegenusPogouoglossusSchneien isnextrelated,if Moore, B. P. 1998. The enigmatic tribe Physocrotaphini
itiscloselyrelated tothisgenusatall. Inviewofthe Chaudoir (Helluodini auct.) (Coleoptera: Carab-
veryincompleteknowledgeoftheNewBritainHolo- idae). In: In:Ball,G.E.,A.Casale&A.VignaTagli-
ponerusweonlycanspeculateaboutapossiblerela- anti (eds): Phylogenyand ClassificationofCaraboi-
tionship with this enigmatic genus. dea. -Atti Mus. reg. Sei. nat. Torino 1998: 369-380
Unfortunately, no sampling circumstances are Stork,N.E.1986.AnannotatedchecklistoftheCarabidae
recorded foreither speciesofSchuelea, so theirhab- (including Cicindelinae, Rhysodinae and Paussi-
itsare completely unknown. In view ofthe unique, nae)recordedfromBomeo.-Occ.Pap.Syst.Ent.2:
1-24
bulkybodyshapeand theirunusuallyshortmandi-
bles it may be speculated, however, that ecology
and behaviour of the species of Schuelea in one or
another way deviate from those of the species of
Pogonoglossus.
113