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NEW A COMBINATION NORTH AMERICAN IN TEPHROSIA (FABACEAE) Guy L Nesom James L Zarucchi 2925 Hartwocd Drive Missouri Botanical Garden Fort Worth, Texas 76 USA. Box 299 109, P.O. [email protected] St. Louis, Missouri 63 66-0299, U.S.A. 1 [email protected] bridos entre Tephrosia chrysophylla y T.florida, que reemplaza a T. xfloridana (Vail) Isely. Se aportan detalles de la tipificacion de Cracca intermedia Small y Cracca floridana Vail. was new Cracca intermedia Small described (Small 1894) as a species intermediate between C. chrysophylla (Pursh) Kuntze (= Tephrosia chrysophylla Pursh) and C. ambigua (M.A. Curtis) Kuntze (= Tephrosia Jlorida Wood). new (F. Dietr.) C.E. Vail (1895) curiously provided C. intermedia with a name, Cracca smallii Vail, because she apparently regarded C. intermedia as invalidated by the earlier Tephrosia intermedia Graham in same Wall. In the publication, Vail described C. floridana Vail, noting that it was "very close" to C. smallii. Isely (1982, 1990, 1998) provided the combination Tephrosia xfloridana (Vail) Isely and used in reference it — to the putative T. chrysophylla T.florida hybrids, even though he correctly recognized Graham's T. intermedia nomen nudum. be to a Tephrosia intermedia Graham was indeed published without a description and illegitimate as a nomen is nudum, Many but does not block the transfer of Cracca intermedia Small to Tephrosia. of the nomina nuda it Don were by among in Wallich's list validated description in (1831-1834), but Tephrosia intermedia not is them. "T. intermedia Grah. in Wall. Cat. 5632" was listed at the end of the paragraph for Tephrosia tinctoria & [W Pers. var. intermedia A.] in J.D. Hooker, Fl. Brit. India 2:112. 1879, so the epithet "intermedia" at the rank under was of species "Tephrosia" not there validated. & Tephrosia xintermedia Nesom comb, nov (Small) G.L. Zarucchi, Craccc The who original hypothesis of hybrid origin of Tephrosia intermedia has been accepted by those have further Wood studied the genus (Vail 1895; Robinson 1899; 1949). Isely (1998, 856-857) noted that "As alleged p. Journal of the Botanical Research Institute of Texas 3(1) 158 and Wood between by (1949), Tephrosia xjloridana probably is a derivative of hybridization T. chrysophylla Whatever widely distributed and consistent in characters to deserve origin, sufficiently T. florida. its it is been from Alabama, Georgia, and Mississippi, but the distribution binomial has collected Florida, listing." It with sporadic and origin as a recurrent hybrid seems a reasonable hypothesis. It occurs in association is its and morphology. the two putative parents is intermediate in ACKNOWLEDGMENTS We Kanchi Gandhi (GH) and Richard Wunderlin (USF) for comments and analysis. are grateful to P. REFERENCES A dichlamydeous -4. Don, 831 -1 834. general history of the plants, vols. G. 1 1 New combinations and one new variety among the genera indigofera, Robinia, and Tephrosia 1982. D. Isely, (Leguminosae). 34:339-341. Brittonia D. 990. Vascular flora of the southeastern United States, Vol. 3, Part 2: Leguminosae (Fabaceae). Univ. of Isely, 1 Chapel North Carolina Press, Hill. and and Leguminosae (Fabaceae) of the United States (exclusive of Alaska 1998. Native naturalized D. Isely Museum. Mi. Bean Science Univ. of Utah, Provo. Hawaii). Life 93-202. of the North American species of Tephrosia. Bot. Gaz. 28:1 Robinson, B.L 899. Revision 1 Club :300-307. Small, J.K. 894. Studies in the botany of the southeastern United States.— II. Bull.Torrey Bot. 21 1 A.M. 1895. A revision of the North American species of the genus Cracca. Bull.Torrey Bot. Club 22:25-36. Vail, Rhodora 93-231 233-302, Wood, The American barbistyled species of Tephrosia (Leguminosae). 51:1 949. C.E. 1 , 305-364, 369-384.