fmicb-10-01900 August 12, 2019 Time: 16:44 # 1 ORIGINAL RESEARCH published: 14 August 2019 doi: 10.3389/fmicb.2019.01900 Edited by: Ashley C. Banyard, Animal & Plant Health Agency, United Kingdom Reviewed by: Frank van der Meer, University of Calgary, Canada Nicholas Johnson, Animal & Plant Health Agency, United Kingdom *Correspondence: Zhiqiang Wu