1 Guinea Fowl Coronavirus Diversity has Phenotypic Consequences for Glycan 1 and Tissue Binding 2 3 Kim M. Bouwman1, Mattias Delpont2, Frederik Broszeit3, Renaud Berger2, Erik 4 A.W.S. Weerts1, Marie-Noëlle Lucas2, Maxence Delverdier2, Sakhia Belkasmi2, 5 Andreas Papanikolaou1, Geert-Jan Boons3, Jean-Luc Guérin2, Robert P. de Vries3, 6 Mariette F. Ducatez2*, Monique H. Verheije1* 7 8 1 Department of Pathobiology, Faculty of Veterinary Medicine, Utrecht University, 9 3584 CL Utrecht, The Netherlands 10 2 IHAP, Université de Toulouse, INRA, ENVT, 23 Chemin des Capelles, 31076 11 Toulouse, France 12 3 Department of Chemical Biology & Drug Discovery, Utrecht Institute for 13 Pharmaceutical Sciences, Utrecht University, 3584 CG Utrecht, The Netherlands 14 * corresponding authors M. Verheije: