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February 2011 Journal of Japanese Botany Vol. 86 No.1 41 めているように，植物学的にはシロバナハギはミヤギノハ名はL. intermedia Nakai var. alba Nakai (1923)であギL. thunbergii (DC.) Nakaiにもっとも近い種類である．る．この学名をL. thunbergiiに組み替えかえて，品種ラミヤギノハギの範囲については諸説あるが，Ohashi et al. ンクでシラハギの学名としなければならない．李昌福「大 (2009) は日本，朝鮮，台湾，中国，インド東部アッサム韓植物圖鑑」(1980) の1869. Lespedeza thunbergii var. に分布する野生種であるとした．この種（広義のミヤギノ intermedia (Nak.) T. Lee には “for. alba T. Lee” の名称ハギ）は著しく多形で，いくつかの亜種に分けられ，そのがあるが，この新組み合わせは正式に発表されていない．中の subsp. thunbergii は狭義のミヤギノハギで，日本，したがって，シラハギの学名として新たにL. thunbergii f. 朝鮮，中国に分布し，ビッチュウヤマハギ，チョウセンヤ alba (Nakai) H. Ohashi & K. Ohashiを提案した．マハギ，ニシキハギなどは同一と考えられる (Ohashi et 　シロバナハギの野生品は朝鮮半島（北朝鮮江原道）か al. 2009)．われわれはシロバナハギはミヤギノハギ（狭義）ら中井 (1923) によってシロバナチョウセンヤマハギとしてに由来する野生白花品であると考えている．報告された．中井（1927）は後にこれをシロバナハギで　シロバナハギをミヤギノハギの種内分類群とする学名はあると訂正した．Lee (1965)も韓国から新産地を追加し Schneider (1907)が最も早く発表していて，Lespedeza た．前川(1938)は三重県で黒川喬雄採集の自生とされる sieboldii Miq. var. albiflora C. K. Schneid.である．大 L. japonica を紹介したが，Akiyama (1988) は自生を疑井(1965b)はこの学名に基づいてL. thunbergii (DC.) 問とした．Hatusima (1967)は福岡市でみたことがあると Nakai var. albiflora (C. K. Schneid.) Ohwi を発表ししているが，これは栽培品の逸出が疑われる．日本ではた．しかし，この変種形容語を生かしたL. thunbergii まだシロバナハギ野生品は見つかっていないと思われる． f. albiflora (C. K. Schneid.)という組み合わせはL. thunbergii f. albiflora (Matsum.) H. Ohashiの後続同名（東北大学植物園津田記念館，となるために，作ることはできない．次に発表された学岩手医科大学薬学部） J. Jpn. Bot. 86: 41–46 (2011) Subhasis Pandaa,* and Munivenkatappa sanjaPPab: The Entity of Enkianthus indicus M. R. Debta & H. J. Chowdhery (Ericaceae) a,*Taxonomy and Biosystematics Laboratory, Post-Graduate Department of Botany, Barasat Government College, Kolkata 700124, Central National Herbarium, Howrah, 711103 INDIA bBotanical Survey of India, CGO Complex, 3rd MSO Building, Block–F, 5th Floor, Sector I, Salt Lake City, Kolkata, 700 064 INDIA *Corresponding author: [email protected] Summary: Enkianthus indicus M. R. Debta & H. (M. R. Debta 40542, CAL; Fig. 1A) and J. Chowdhery (Ericaceae) is a minor quantitative paratype (M. R. Debta 40880, CAL; Fig. 1B), variation of the E. deflexus (Griff.) C. K. Schneid. it was observed that Enkianthus indicus M. R. complex and therefore the former is proposed here Debta & H. J. Chowdhery is a member of the E. as a synonym under the latter species. Photographs deflexus (Griff.) C. K. Schneid. complex. The of holotype and paratype of E. indicus as well as detailed description of E. indicus as described the type photograph and specimen examination of E. deflexus in Indian herbaria are provided to clear by M. R. Debta and H. J. Chowdhery (J. Non- confusion. Timber Forest Products 16(2): 173–174, 2009) was the protologue description of E. deflexus After a critical examination of the holotype (Griff.) C. K. Schneid. (Ill. Handb. Laubholzk. 42 植物研究雑誌　第86巻　第1号 2011年2月 L). A C 0, 8 8 0 4 a bt e D R. M. e ( p y at ar P B. L). A C 2, 4 5 0 4 a bt e D R. M. e ( p y ot ol H A. y. er h d w o h C J. H. & a bt e D R. M. s u c di n s i u h nt a ki n E 1. g. Fi February 2011 Journal of Japanese Botany Vol. 86 No.1 43 Table 1. Comparative account between Enkianthus indicus M. R. Debta & H. J. Chowdhery and E. deflexus (Griff.) C. K. Schneid. E. indicus E. deflexus* E. deflexus** Leaf number and size 5–12 in each 4–8 in each pseudowhorl, 4–12 in each pseudowhorl, pseudowhorl, 30–70 × 15–30 mm 10–90 × 5–35 mm 15–54 × 6–20 mm Leaf apex mucronulate to acute acute usually mucronulate to rarely acute Leaf pubescence densely, minutely pubescent scarsely hispidulous above, leaf indumentum variable, intermixed with scattered densly hispidulous beneath both types observed hispidulous hairs along midrib above, later becoming glabrescent, sparsely hispidulous along midrib or glabrescent beneath Rachis length up to 30 mm long 50–85 mm long 10–50 mm long Seed size 3–3.5 mm long up to 2 mm long 2–4 mm long *As delimited by Debta and Chowdhery. **Based on protologue, types and geographical range of variations in herbarium specimens. 2: 521, 1911). The authors did not cite the paper one pseudowhorl which shows 10 leaves. The of Anderberg (1994) who revised and classified authors put wrong data in E. deflexus without the genus Enkianthus Lour. recognizing four studying protologue, type materials, relevant sections and 17 species based on morphological, references (Hara 1966, 1982, Hsu 1982, anatomical, embryological and cytological Anderberg 1994, Ruizheng and Stevens 2005, data, instead they cited Mabberley (2008) Panda and Sanjappa 2006, Panda 2008) and who followed Anderberg (1994). They did even geographical range of variations observed not consult the specimens deposited in Indian in herbarium specimens (K, CAL, BSIS, DD, Herbaria (particularly CAL, BSIS, DD, BSHC BSHC and ASSAM). Enkianthus indicus is also and ASSAM) where morphological quantitative matched with E. deflexus (Griff.) C. K. Schneid. minor variations are clearly observed, even in a var. chinensis (Franch.) H. Hara in possessing single specimen. nearly glabrous abaxial leaves (some), but not Furthermore they did not consult the type constant abaxially on all leaves. material of E. himalaicus Hook. f. & Thomson Debta and Chowdhery (2009) described and (CAL), a heterotypic synonym of E. deflexus delimited E. indicus (as a new species) without (Griff.) C. K. Schneid. where leaves 5–12 in studying protologue, types and variations in each pseudowhorl (Fig. 2A) instead of 4–8, herbarium specimens to its geographical range apex clearly mucronulate instead of acute (type of its close ally, the Enkianthus deflexus (Griff.) photograph of E. deflexus showed mucronulate C. K. Schneid complex. To clear the confusion, leaf apex, Fig. 2B), leaf pubescence like E. a comparative account among Enkianthus indicus and seeds up to 4 mm long instead of indicus M. R. Debta & H. J. Chowdhery, E. 2 mm as described by Debta and Chowdhery deflexus (Griff.) C. K. Schneid. sensu Debta (2009). Enkianthus indicus as described by & Chowdhery and E. deflexus (Griff.) C. K. Debta and Chowdhery (2009) is clearly matched Schneid. based on protologue, types (K photo !, with the description of E. himalaicus Hook. CAL !) and geographical range of variations in f. & Thomson, a himalayan variation of E. herbarium specimens (CAL, BSIS, DD, BSHC deflexus. Holotype of E. indicus (Fig. 1A) shows and ASSAM) is described in Table 1. Therefore mostly 5–6 leaves in each pseudowhorl except E. indicus M. R. Debta & H. J. Chowdhery is 44 植物研究雑誌　第86巻　第1号 2011年2月 n., s. er k o o H D. J. s ( u c ai al m hi E. A. d. ei n h c S K. C. Griff.) s ( u x e fl e d E. d n a n o s m o h T & k. f. o o H s u c ai al m hiK). us 9, h6 ant& 9 nki7 E1 of h 7 s fit menGrif e speciflexus ( pe yd HolotB. E. 2. K). g. Fi February 2011 Journal of Japanese Botany Vol. 86 No.1 45 proposed here as a synonym under E. deflexus 8620 (BSHC); Lachung to yumthang, 23 May 1987, D. (Griff.) C. K. Schneid. C. S. Raju & B. Mitra 7429 (BSHC); Lachen to Chhaten, no date, R. S. Rao 423 (BSHC); Lachen, 25 AUg. 1982, P. Chakraborty 2291 (BSHC); Lachen to Log bridge, 14 Enkianthus deflexus (Griff.) C. K. Schneid., July 1986, D. C. S. Raju & S. Singh 5897 (BSHC); Tsoka- Ill. Handb. Laubh. 2: 521 (1911) – Rhodora Yuksum trek, 22 June 1993, S. Pradhan 15177 (BSHC). deflexa Griff., Itin. Pl. Khasyah Mts.: 187 (1848) Lachen, 9000 ft, May, 1885, G. King’s Collector s.n. (Lloyd – Meisteria deflexa (Griff.) Nakai in Bot. Botanical Garden Herbarium, Darjeeling); Lachung, 9000 ft, 20 Aug. 1909, Smith & Cave 2592 (Lloyd Botanical Mag. (Tokyo) 38: 39 (1924). Type: BHUTAN. Garden Herbarium, Darjeeling). Between Jaisa and Tongsa, 9000–9500 ft, West Bengal (Darjeeling): Darjeeling, July, 1915, E. A. Griffith 717, 969 (K, photo! CAL!). C. Modder 37D (CAL); Tonglu: 10000 ft, 19 May 1909, I. Enkianthus himalaicus Hook. f. & Thomson H. Burkill 30891 37D (CAL); Darjeeling. 9000 ft, S. Kurz in Hook., J. Bot. Kew Gard. Misc. 7: 125, t. 3 s.n. (CAL 266754); Darjeeling, 30 Oct. 1875, Dongboo s.n. (CAL 266748); Darjeeling, 10000 ft, July, 1882, J. S. (1855). Type: INDIA. Sikkim. Lachen, 8000– Gamble 10416 (CAL); Darjeeling, 9000–10000 ft, 29 May 10000 ft, May, 28, J. D. Hooker s.n. (K, photo !). 1902, J. H. Lace 2251 (CAL); Darjeeling, 28 May 1862, Enkianthus indicus M. R. Debta & H. J. T. Anderson 257 (CAL); Darjeeling, 9000 ft, 4 June1884, Chowdhery in J. Non-Timber For. Prod. 16 C. B. Clarke 35694E (CAL); Tonglu, 9000–10000 ft, 29 May 1902, J. H. Lace 2251 (DD); Tonglu, 10000 ft, 19 (2): 173 (2009), syn. nov. Type: INDIA. West May 1909, I. H. Burkill 30891 (BSIS); Singalelah, 11000 Bengal, Darjeeling, Tonglu to Gairibas, 2900 m, ft, 13 June 1892, G. A. Gammie 79 (Lloyd Bot. Garden 29 May 2006, M. R. Debta 40542 (CAL–holo Herbarium, Darjeeling). !); Darjeeling, below Tonglu, 2850 m, 1 Apr. 2007, M. R. Debta 40880 (CAL–para !). First author (S.P.) is grateful to the Director, Specimens examined: Arunachal Pradesh. West Botanical Survey of India for awarding a Kameng district, Peri La, 24 May 1958, G. Panigrahi research fellowship under “Flora of India 16079 (CAL); Seargaon, 19 May 1958, G. Panigrahi Project”, manuscript correction and arranging 15806 (CAL); Tawang district, SengeDzong, 20 Sept. to send type photograph from K. Thanks are 1969, J. Joseph 40158 (CAL); Jabrang, 8 May1955, K. S. Srinivasan s.n. (BSIS 43357); SengeDzong to SeLa, 11200 also due to Curators of different Indian herbaria ft, 21 May 1957, R. S. Rao 7664 ASSAM); Lama Camp, said above for their help during herbarium West Kameng district, 14 May 1998, A. Sarmah 9932 consultations. (APFH, Arunachal Pradesh Forest Herbarium); Lower Subansiri district, Talle Valley, 10 June 1984, Haridasan Literature cited 1111 (APFH) ; Talle Valley, 17 June 1994, Haridasan & C. Barooa 6904 (APFH); Lohit district, Simbi Hotspring, 9 Anderberg A. A. 1994. Cladistic analysis of Enkianthus Oct. 1984, Haridasan 1581 (APFH). Laou, 7900 ft, 11 Dec. with notes on the early diversification of the Ericaceae. 1999, G. D. Pal 8407 (ARUN, Arunachal Pradesh Field Nordic J. Bot. 14(4): 385–401. Station, Botanical Survey of India). Debta M. R. and Chowdhery H. J. 2009. A new species Sikkim. North district: Chungthang to Lachung, of Enkianthus Lour. (Ericaceae) from India. J. Non- 7000 ft, 25 May 1945, K. Biswas 6963 (CAL); Lachung, timber Forest Products 16(2): 173–174. 9000 ft, 9 Aug. 1892, G. A. Gammie 698 (CAL); Lachen, Hara H. 1966. The Flora of Eastern Himalaya – Results 8000–10000 ft, 17 May 1945, K. Biswas 6661 (CAL); of the Botanical Expedition to Eastern Himalaya Zemu Valley, 9500 ft, 9 July 1909, Smith & Cave 1062 organized by the University of Tokyo, 1960–1963: 234, (CAL); Lachen, 9000 ft, May, 1885, King’s Collector s.n. 647. University of Tokyo Press, Tokyo. (CAL 266757); Lachen, 8000–10000 ft, J. D. Hooker s.n. Hara H. 1982. In: Hara H., Charter A. O. and Williams L. (CAL 266743); Yumthang to Lachung, 3400 m, 1 June H. J. (eds.), An Enumeration of the Flowering Plants 2002, S. Panda 29993 (CAL); Yakche to Phuney, 3100 m, 1 of Nepal 3: 55. Trustees of British Museum (Natural June 2002, S. Panda 29994 (CAL); Lachen, 2950 m, 7 May History), London. 2001, S. Panda 29904 (CAL); Lachen to Chhaten, 2700 m, Hsu T. 1982. Classification, Distribution and Phylogeny 8 May 2001, S. Panda 29907 (CAL); East district, West of of the genus Enkianthus. Acta Bot. Yunnan. 4(4): Chhangu, 11000 ft, June, 1910, W. W. Smith 3302 (CAL); 355–362. Lingto, 11000 ft, 24 Oct. 1875, C. B. Clarke 25549 (CAL); Mabberley D. J. 2008. Mabberley’s Plant-Book: A Portable North district, Dongbong, 2 June 1991, R. C. Srivastava Dictionary of Plants, Their Classification and Uses, ed. 46 植物研究雑誌　第86巻　第1号 2011年2月 3: 307. Cambridge University Press, Cambridge. Panda S. 2008. Taxonomic revision of some selected Enkianthus indicus M. R. Debta & H. J. Chowdhery（ツツ genera of Ericaceae in India. pp. 111–115. Ph. D. ジ科）の正体（S. Pandaa，M. sanjaPPab） thesis, Vidyasagar University, Midnapore (India). 　2009年にインド，西ベンガル州ダージリンから記載さ Panda S. and Sanjappa M. 2006. A new taxon of Enkianthus Lour. (Ericaceae) from Sikkim Himalaya. れたEnkianthus indicus M. R. Debta & H. J. Chowdhery Ind. J. Forestry 29(3): 277–279. のタイプ標本を詳しく調べた結果，これはE. deflexus Ruizheng F. and Stevens P. F. 2005. Enkianthus Lour. In: (Griff.) C. K. Schneid.の異名であることが明らかになった． Mingyuan F., Ruizheng F., Mingyou He, Linzhen Hu, 混乱を防ぐために，E. deflexusとE. indicusのタイプ標 Hanbi Y., Haining Q., Tianlu M., Chamberlain D. F., 本を示した． Stevens P. F., Wallace G. D. and Anderberg A. (eds.), Flora of China 14: 244. Science Press, Beijing and (aインド植物調査所国立中央植物標本館， Missouri Botanical Garden, St. Louis. bインド植物調査所）植物研究雑誌　86: 46–49 (2011) アワモリショウマの新品種チャボアワモリショウマ（ユキノシタ科チダケサシ属）（秋山　忍） Shinobu akiyama: A New Form of Astilbe japonica (C. Morren & Decne.) A. Gray (Saxifragaceae) from Japan 国立科学博物館植物研究部 Department of Botany, National Museum of Nature and Science, 4-1-1, Amakubo, Tsukuba, 305-0005 JAPAN Summary: A new form, Astilbe japonica f. pygmaea って咲くが小さく，花弁は細くあまり目立たな S. Akiyama (Saxifragaceae), is described, which is い．しかしアワモリショウマは，小さな花ながら known as a cultivated plant. 集まって咲く姿は目立ち，その様子から泡盛升麻の名がつけられたという．アワモリショウマの草　チダケサシ属（ユキノシタ科）は東アジアと北丈は Flora of Japan IIb（Ohba 2001）によれば50 米に隔離分布し，東アジアでの種分化が顕著であ～80 cmである．しかし，園芸市場では，那智原り，特に日本においては地域的な固有種や固有変産として，草丈が20～25 cm ほどの小型品がナ種が多く知られている．また，この類はヨーロッチアワモリショウマの名で販売されている．最近，パや日本など世界的規模でしばしば園芸植物としこれよりも草丈が低く高さ9～17 cmほどの矮て栽培され交配種も作出されている．性のものがチャボアワモリショウマの名で販売さ　日本では，アワモリショウマAstilbe japonica れている．その由来などについては不明である． (C. Morren & Decne.) A. Gray，コヤクシマショ　これらのアワモリショウマ小型品の実体を明らウマA. glaberrima Nakai var. saxatilis (Nakai) H. Ohba，ヒトツバショウマA. simplicifolia Makino かにするために，アワモリショウマの変異を調べなどが山野草として栽培される．アワモリショウた．マは日本固有種であり，九州，四国，本州（静岡　国立科学博物館，東京大学総合研究博物館，京県以西）の渓谷の岩上などに生育する．その小葉都大学総合博物館，首都大学東京牧野標本館の標のかたちは他のチダケサシ属植物と比較して，細本151点を検討したところ，アワモリショウマの長く流線型であり，渓流植物ともみられている（田開花時から果実時の草丈は34.1±16.1 cm（最村他 2001）．チダケサシ属の花は円錐花序に集ま小9 cm，最大85 cm）であった．那智を産地と