Sources of Records for Albania in Flora Europaea

172 Pages·2012·5.21 MB·English

by  Lisa Cheung

Sources of records for Albania
in Flora Europaea

compiled by Peter W. Ball

CONTENTS

1. Introduction - page 2

2. Flora Europaea vol. 1 - page 6

3. Flora Europaea vol. 2 - page 21

4. Flora Europaea vols 3 and 4 - page 40

5. Flora Europaea vol. 5 - page 65

6. Plant collections from Northern Albania by Bruno Schütt - page 78
Part 1: Introduction

The following lists of the flora of Albania were compiled in the period 1961-1968.
During the early stages in the preparation of manuscripts for Flora Europaea volume 1 it
became apparent that there was no reliable source of information for geographical data
for countries in the Balkan peninsula except for Bulgaria. The main Flora covering the
region was Hayek (1924-1931), but Hayek gives distributions based on areas defined by a
combination of phytogeographical considerations and pre-1914 political boundaries.
While most of these areas are wholly within the post 1945 political boundaries some,
particularly Albania, Epirus, Macedonia and Thrace transcend post 1945 political
boundaries and so do not present reliable evidence for the occurrence of a species within
the political areas recognized in Flora Europaea (Webb 1963). This created difficulties
for two countries, Turkey and Albania. D. A. Webb compiled a list for Turkey-in-Europe
(which was part of Hayek’s Thrace), and this was published by Webb (1966). I took on
the task of compiling a list for Albania (see Webb, 1963).
Although Hayek recognized an area he called Albania this area differed significantly
from the post 1919 boundaries of Albania. Hayek’s Albania extended the north-eastern
boundary considerably further east than the post-1919 political boundary, including all of
the mountain range often referred to as “Northern Albanian Alps”. Much of this mountain
range is in Serbia and Macedonia (particularly the part named !ar Planina) and at the
time Hayek was published this portion of the range was much more extensively explored
than the part which remained in Albania. At the southern boundary Hayek drew the
boundary slightly further north than the post 1919 boundary, including this part of
Albania in Epirus, which is primarily part of post 1919 Greece. Following the end of the
first world war several collecting expeditions were sent to Albania resulting in many
additions to the flora which were only partially included in Hayek. A further source of
confusion arises from many pre-1914 collections in which the labels state that the
collection was from Albania when in fact it came from Serbia or Macedonia. Some of
these collections are from areas which are now along political boundaries (especially high
mountain peaks such as Korab, Koritnik and Pastrik) so introducing an element of
uncertainty about which country they came from. During the post-1920 period these high
mountain border peaks were often out-of-bounds for plant collectors so it was not
possible to verify these older reports of the occurrence of a species in Albania.
The list was compiled from published papers, mostly derived from the extensive
bibliography published by Markgraf (1932) together with a supplement Markgraf (1963).
Some post-1945 collections and publications could not be included as they were not
available to me at the time. However some did become available after the publication of
volumes 1 and 2 of Flora Europaea so the list for volumes 3 and 4 is more complete than
those for the earlier volumes. The bibliography indicates (*) those publications which
became available at a later time. For a variety of reasons the list was never completed for
volume 5 of Flora Europaea. The information that had been accumulated by 1968 is
included here, but it does not coincide with the records published in Flora Europaea
volume 5. There are about 50 species included in Flora Europaea volume 5 which are not
in that list. It is likely that these were derived from Flora of Tirana (Tir) and from other
sources that became available after 1970. However the list includes about 90 species
2
reported from Albania but which are not reported for Albania in Flora Europaea. Some of
these appear to be the only report of the species from Albania.
In addition to published records I was given access to unpublished material. The most
important of these was a manuscript by Bruno Schütt which was made available by the
Curator of Botany of the Uebersee Museum, Bremen (BREM). This manuscript is
included as part 6 of this document. Another important addition was a complete list of the
collections made by Alston and Sandwith kindly provided by N. Sandwith. Although
Alston & Sandwith published a proportion of their records this list contained a number of
records which appear to be the only reports of a species from Albania. I was also given
access to complete lists of collections made in Albania during the 1930’s by several
different people and housed at Royal Botanic Gardens, Kew. A few reports of these
collections were published by Turrill (1932, 1939) and Chator & Turrill (1934) but most
are unpublished.
It is important to understand that this list does not purport to represent a comprehensive
list of the flora of Albania. Its main purpose was to ensure that species reported from
Albania in Flora Europaea were based on evidence which could be verified either in print
or by an author examining herbarium specimens. Unfortunately some important
collections of Albanian plants were destroyed during the second World War and later,
particularly those of Markgraf at B. Some material at W was also lost in 1945, and much
material at Budapest (BP) was lost in 1956.
The list is arranged in the sequence published by Hayek (1924-1932), the species being
numbered according to Hayek. Because Flora Europaea followed a slightly different
family sequence from Hayek a few families are out of sequence when they occur in a
different volume of Flora Europaea. For example volume 1 of Hayek is covered by
volumes 1 and 2 of Flora Europaea and volume 2 of Hayek is covered by volumes 3 and
4 of Flora Europaea. Within a family the genera and species are arranged in the Hayek
sequence, new genera being placed at the end of the family and new species at the end of
the genus. Species or infra specific taxa included in the list without a reference are
reported from a minimum of 3 sources by at least two independent authors. Species
reported from one or two sources, or reported by only one author, include appropriate
references. It is important to note that one source may report a species from several
localities and in a few cases a species reported by several different authors may be
restricted to a single locality. Post-1919 collections are assumed to be in present-day
Albania although some are from border mountains. Pre-1919 collections were traced to
present-day Albania as far as it was possible to do so. Where there was a considerable
element of doubt about the locality, records were not included. Baldacci (1899), and
Baldacci (1901) are particularly difficult as Baldacci (1899) collected in areas that are
now mainly in Greece and Baldacci (1901) crossed into Montenegro although most of
these collections appear to be from Albania. Many of the D"rfler and other collections
reported by Hayek (1917) and Hayek (1924) are from boundary areas so some care has to
be exercised with these records and in a few cases it is not possible to be sure which
country a collection came from. Such records are indicated as boundary or border
records, especially when they are the only available record for a species, although many
of these were verified by the unpublished records of Schütt.
Below is a key to the abbreviations used in the list for sources of records together with
references to those publications also mentioned above. Extensive bibliographies for
3
Albania were published by Markgraf (1932, 1963) and these should be consulted for
older sources of information.

List of abbreviations and bibliography.

A&S Alston, A.G. H. & N.Y. Sandwith, 1940. Results of two Botanical
Expeditions to South Albania. Jour. Bot. (London) 78.
Unpublished list by N. Y. Sandwith.
B94 Baldacci, A., Rivista critica della collezione botanica fatta nel 1892 in
Albania. Malpighia 8.
B 94b Baldacci, A., 1894. Contributo alla conscenza della Flora dalmata,
montenegrina, albanese, epirota e greca. N. Gior. Bot. Ital. 1.
B96. Baldacci, A., 1896. Rivista della collezione botanica fatta nel 1894 in
Albania. Bull. Herb. Boiss. 4.
B99 Baldacci, A., 1899. Rivista dalla collezione botanica fatta nel 1896 in
Albania. N. Giorn. Bot. Ital. 6.
B01 Baldacci, A., 1901. Rivista della collezione botanica fatta nel 1897
nell’Albania settentrionale. Mem. Acad. Sci. Ist. Bologna, ser. 9, 5.
B17 Baldacci, A., 1917. Itinerari albanasi. Rome.
BB Baldacci, A. & Beguinot, 1918. Contributo alla flora autumnale et invernale
del dintorni di Vallona. N. Giorn. Bot. Ital. 25.
Bm33 Bornmüller, J., 1933. Zur Flora von Montenegro, Albanien und
Macedonien. Magyar Bot. Lapok 32.
Chator, D. A., & W. B. Turrill, A Contribution to the Flora of Central
Albania. Kew Bull. 1934.
Form. Formanek, E., 1895. Beiträg zur Flora von Albanien, Korfu und Epirus.
Verh. Natf. Vereins Brünn. 33
See Vandas, 1909.
Gr Grimus, V. von, 1871. Beiträge zur Flora Albaniens. Verh. Zool.-Bot. Ges.
Wien 21.
Giu Giusseppi. Specimens at Kew. See Chator & Turrill and Turrill.
H17 Hayek A. von, 1917. Beiträge zur Kenntnis der Flora des albanisch-
montenegrinischen Grenzgebietes. Denschr. Akad. Wiss. Wien, Math.-
natw. Kl. 94
H24 Hayek, A. von, 1924. Zweiter Beiträg zur Kenntnis der Flora Albanien.
Denkschr. Akad. Wiss. Wien, Math.-natw. Kl. 99.
Hayek, A. von, Prodromus Florae Peninsulae Balcanicae. Vol. 1, 1924, Vol.
2, 1927, Vol. 3, 1932 (completed by F. Markgraf).
Held. Heldreich, von, 1879. Ein Beitrag zur Flora von Epirus, geliefert von Herrn
N. K. Chodzes. Verh. Bot. Vereins Prov. Brandenburg 21. Sitzber.
Hep Hepburn, I. Specimens at K. Some published by Turrill and Chator & Turrill
J20 Janchen, E., Vorarbeiten zu einer Flora der Umgebung von !kodra in Nord-
Albanien. Österr. Bot. Zeitschr. 69.
4
Jav. Csiki, E., A. Jávorka & E. B. Kümmerle, 1926. Additamenta ad Floram
Albaniae. Pteridophyta by E. Kümmerle, Anthophyta by A. Jávorka.
Budapest.
M27 Markgraf, F., 1927. An den Grenzen des Mittelmeergebiets:
Pflanzengeographie von Mittelalbanien. Feddes Repert. Beih. 45.
M32 Markgraf, F., 1932. Pflanzengeographie von Albanien. Bibliotheca Botanica
105.
M42* Markgraf, F., 1942. Reisebericht aus Albanien. Notizbl. Bot. Gart. Mus.
Berlin-Dahlem. 15.
Markgraf, F., 1963. Bericht Albanien 1961. Webbia 18.
K48 Kitanov, B., 1948. Contribution á l’étude de la Flora d’Albanie de l’Est. Ann.
Fac. Phil. Univ. Skopje Sect. Sci. Nat. 1.
Pamp. Pampanini, R., 1915. Contributo alla conoscenza della Flora d’Albania. Bull.
Soc. Bot. Ital. 84.
Penn. Pennington. Specimens at Kew. See Chator & Turrill and Turrill.
R. Rechinger fil., K. H., 1939. Zur Flora von Albanien und Mazedonien. Feddes
Repert. Sp. Nov. 47.
Rohl Rohlena, J., 1937. Beitrag zur Flora Albaniens. V#stník Kralovské $eské
Akad. Spolo%nosti Nauk Trída Matemat. P&irod. 1937.
Rod Rodriguez, 1923. Plantes de l’Albanie Moyenne. Bull. Mus. Hist. Nat. Paris
Sch Schütt, B. Unpublished list. See part 6 below.
Tir* Paparisto, K., Xh. Qosja & M. Demiri, 1962. Flora e Tiranes. Tirana.
Turrill, W. B., 1932. A Contribution to the Flora of Central Albania. Kew Bull.
1932.
Turrill, W. B., 1939. Two new Species and New or Important Records. Kew
Bull. 1939.
Vandas, K., 1909. Reliquiae Formanekianae. Brunae (Brno).
Webb, D. A., 1963. The need for check-lists of the flora of certain European
territories. Webbia 18.
Webb, D. A., 1966. The Flora of European Turkey. Proc. Roy. Irish Acad.
65B.
Weiss* Weiss, E., 1866. Floristisches aus Istrien, Dalmatien und Albanien. Verh.
Zool.-Bot. Ges. Wien. 16
5
Part 2: Flora Europaea vol. 1

Pteridophyta

1. Lycopodium selago K48
6. Selaginella selaginoides K48
7. S. helvetica M27
9. Equisetum maximum
10. E. arvense
12. E. palustre
13. E. limosum H24
14. E. ramosissimum
15. E. hiemale
22. Ophioglossum vulgatum M27 M32
24. Botrychium lunaria
26. B. matricariaefolium H24
28. Osmunda regalis M32 Bm33
29. Polypodium vulgare
33. Pteridium aquilinum
34. Adiantum capillus-veneris
35. Notholaena marantae
38. Cheilanthes persica
39. Cryptogramma crispa K48
42. Phyllitis scolopendrium M27 M32
46. Asplenium viride
48. A. trichomanes A. var. trichomanes B. var. csikii Jav
53. A. adiantum-nigrum I. subsp. adiantum-nigrum II. sibsp. serpentini III. subsp.
onopteris
54. A. fissum
55. A. ruta-muraria
56. A. lepidum Jav
57. A. septentrionale
55 x 56. A. x javorkae Jav
58. Ceterarch officinarum
59. Athyrium filix-foemina
61. Dryopteris linnaeana A&S H17(border)
62. D. robertiana B01 Sch
66. D. filix-mas
67. D. villarsii I. subsp. rigida II. subsp. pallidaI
68. D. spinulosa K48
69. Polystichum lonchitis
69 x 70. P. x illyricum
70. P. lobatum
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71. P. setigerum
74. Cystopteris fragilis
75. C. regia
79. Marsilea quadrifolia Sch

Gymnospermae

81. Taxus baccata
83. Juniperus communis I. subsp. communis II. subsp. nana
84. J. oxycedrus
85. J. macrocarpa B96
86. J. foetidissima
89. J. phoenicea Sch Gr?
90. Cupressus sempervirens BB
91. Abies alba
A. borisii-regis A&S
94. Picea excelsa Jav Sch
97. Pinus halepensis B96 M32
100. P. mughus
101. P. heldreichii M32 Bm33
102 I. P. nigra
104. P. pinea M32
105. P. peuce
106. Ephedra campylopoda
107. E. distachya

Betulaceae

109. Betula pendula H24(boundary)
112. Alnus incana
113. A. glutinosa
114. Carpinus betulus
115. C. orientalis
116. Ostrya carpinifolia
117. Corylus colurna A&S
118. C. avellana

Fagaceae

120. Fagus silvatica
122. Castanea sativa
123. Quercus cerris (vars. cerris, austriaca, pseudocerris)
124. Q. macedonica
125. Q. aegilops B94 B94b
126. Q. pseudosuber M27
127. Q. coccifera
7
128. Q. ilex
130. Q. conferta
131 II. Q. lanuginose
136. Q. sessiliflora
137. Q. robur B94 M32
123 x 124 R
130 x 136 Jav

Juglandaceae

139. Juglans regia

Salicaceae

140. Populus alba
142. P. tremula
143. P. italica
144. Salix amygdaloides Sch
145. S. pentandra Bm33
147. S. alba
154. S. arbuscula Jav Sch
155. S. hastata Jav
159. S. elaeagnos
162. S. purpurea I. subsp. purpurea A&S II. subsp. amplexicaulis Jav M27
163 I. S. retusa
165. S. reticulata

Moraceae

166. Morus alba J20 Sch (?cult)
167. M. nigra Sch (?cult)
168. Ficus carica
169. Humulus lupulus J20 Sch

Ulmaceae

171. Ulmus campestris (incl. U. glabra, U. foliacea)
173. U. scabra B01
174. U. laevis H24
176. Celtis australis

Urticaceae

179. Urtica dioica
180. U. membranacea BB H24
181. U. urens
8
182. U. pilulifera J20 Sch
184. Parietaria erecta
185. P. ramiflora

Santalaceae

189. Osiris alba
191. Thesium alpinum B01 K48
194. T. linophyllum M27 H24 (boundary)
195. T. divaricatum
196. T. ramosum A&S Jav
198. T. parnassi
200. T. auriculatum Jav Sch
203. T. bergeri B99

Loranthaceae

206. Loranthus europaeus
207. Arceuthobium oxycedri
209. Viscum laxus

Chenopodiaceae

353. Polycnemum majus B94 J20
356 I. Beta maritime
359. Chenopodium botrys
362. C. vulvaria Jav
363. C. hybridum Jav
365. C. opulifolium
366 I. C. album
368. C. urbicum B96
370. C. glaucum B94b A&S
371. C. bonus-henricus
372. C. foliosum Hep
375. Atriplex hortensis J20
380. A. hastata M27 A&S
381. A. rosea B94
382. A. tartarica B96
384. A. portulacoides
388. Camphorosma monspeliaca B17(1894)
391. Kochia prostrata A&S (sub K. scoparia)
393. K. scoparia H24
401. Halocnemum strobilaceum A&S B96
402. Arthrocnemum glaucum
403. Salicornia fruticosa (incl. f.radicans)
404. S. herbacea
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406. Suaeda fruticosa B96
408. S. maritima
409. S. splendens
413. Salsola kali A. var. kali B17(1894) A&S B. var. tragus B17(1894)
414. S. soda B94b A&S
417. Petrosimonia crassifolia

Caryophyllaceae

442. Herniaria hirsute
443. H. incana
445. H. parnassica
446. H. glabra
448. Paronychia chionaea
449. P. kapela
450. P. cephalotes Penn Sch
457. Polycarpon tetraphyllum J20 Sch
460. Spergularia salina M27 Sch
462. S. rubra
463. S. diandra B96
467. Scleranthus perennis Sch Jav
468. S. neglectus
470. S. uncinatus B01 H17
472.S. annuus I. subsp. annuus II. subsp. verticillatus B01 J20
473. Queria hispanica A&S
479. Minuartia bosniaca Sch
480. M. setacea Hep
485. M. velenovskyi H17
486. M. tenuifolia H17
488. M. mediterranea
492. M. verna I. subsp. montana II. subsp. leptophylla A&S Jav IV. subsp. gerardi
496. M. recurva I. subsp. juressi M27 M32 II. subsp. recurva
498 I. M. graminifolia subsp. clandestina
499 I. M. stellata A&S
503. M. baldaccii subsp. I, II, III.
504 (1st) M. garckeana M27
513. Arenaria cretica B94 B96 (A. gracilis sensu Baldacci)
516. A. rotundifolia var. pauciflora B94 B96 var. pancicii B01 Jav
517. A. biflora
518. A. serpyllifolia
519. A. leptoclados
522. A. conferta
*A. serpentinii A&S
525. Moehringia trinervia
530. M. malyi Jav
532. M. ciliata Jav Sch
10