SOME DATA ON THE BIOLOGY OF COMMON BREAM, ABRAMIS BRAMA (L., 1758

23 Pages·2015·3.41 MB·English

ACTA ICHTHYOLOGICA ET PISCATORIA
Abbrev.I:c hAtchPtyiaos L3c 1a( t13.)- :2 6,2 A0nn0.1
ZbignNEJAi,e Anwd rzKeOjMP OWSKI
Fish biology
SOMED ATAO NT HEB IOLOGOYFC OMMONB REAMA,B RAMIS
BRAMA (L1.7,5 F8R)O,M T HEM IF,:DZYODWRAZTEE RS
NIEKTODRAEN EZB IOLOGLIEIS ZCZAAB,RA MIS JJRAMA (L.1,7 58)
ZW ODM IF;DZYODRZA
DepartomfBe inotl ogicRaels oMuaArrgcirenisec, u UlntiuvreaorlfSs zictzyeP coilna,n d
Growrtaht e o(nfb rlayes nemgdet ahs uranedmb eanctks
calculcatoinodnirste)il,oag nto,in vaed slieznategfi ,tr hsa tn d
matuwreirdtyee terimnti hnceeo dm mborne paomp ulation
inhabMiit(,idnzgya od derlzateras,eii act uiantt hveeid c inity
ofS zczecin.
Il,;fTRODUCTION
Alownigtt hhr eo atchhce,o mmborne iamst hmeo sitmp ortfianssthp eciine s
freshwarateears of Poolfban rde.iam n BhiaobPliootlgyii nsghi rswe atlearwtsei lvle ly
knowTnha.em pllies rte loepfvu anbtl iicnatcilopunadspe besyFr isl uk 19(61o39n)5 7,
differento fa sbpianeodcfil tsoshg eyor fti hbeers e samt oicthnke V istulaK arpLiang­oon.
ska-Walus (g1r9o6wo1tf)bh r ediamen2 s 0cM raisbulerakdie ahsne, sr t uodnifi essh
lengtrha-dsriceualsla etb ieoionnfaspg hart iipc iumlparo rfotrman ecteh odoofgl roogwyt h
ratec ablaccuklW oat jin(oo1n 9sa6.b,4 ) i nvesitndi egtatathsieeeld a soinnba rleiamt y
growZtahw.i( s1z9as7 t0u)db iieodla onmgdiy g ratibornesiam ns eovfce oraaPslot ma­l
eralnakieanBs ry.l i(n1s9k7a1 )ac owmrporteemh oennosgiorvnfeape ch u nanddir teyp ro­
ductoifbo rnei andm i ffewraetbneotrd Iinea sd.d iwtoirootnfha, yt teinast isoeonrf i es
papebryMs ar ciak a(b,1c, 9w) 7h 4i ch cfoocnufrcsoi,mbs oebtlihyo logfiischaelr ianeds ­
relpateerds peocndt iiffveqerusee,ns tt iotnbosr ergamer loiatwPneto dhl wiasthe rs.
Brehama asl wpaylsa aynie mdp orrotilannfiet s heorfti hdeeos w nssterceotafim o n
thRiev eOrd ranadi tess t(uNarye auur1b 92Z6i;m d1ar9s4P 1�;c z1a9l6sb3kP; a� czalska
andKr aczki1e9w7Ki2oc;mz p oanwdsP kiie nkowsTkhise p 1e9icv9ise2e cr)soy. m mon
int hMei �dzywoatdearrszws ee lMli.� dzyioasdd erlzarteea iaint ch leo wpeartro fth e
Odras,i tuuatpesdto rfSe zamc zanedcc ionn siosftt wmioajn ogrr i vberran cthheRese :­
galanidct ahW ee stOedrrnai ,n tercboynac n oemcptsleyedsx ot fme imn borran canhde s
canMaolsosttf. h Mei �dzyaroeidaasr b zree hama bihteanitcit,eus n derstthantadhtae b le
4 Zbigniew Neja, Andrzej Kompowski
specdioemsi niatne sc aatccchoeufosnr,t---- iotnnhga e v eraget-hmaonr oe4ft 0 ht%eo tal
annuaclo mmecractifrcaohlmt he (arKeoam po1w9s9k9i) .
Int hdeo wnstrpeartoam ft he anOddi rteass tumaryo,s tt hroeef l eivnanfotrm ation
perttaoti hnbesr esatmo icnkhsa btihStezi cnzgLe acgionPo onm,e raBnayi,aan nL dak e
D!l_bNieeu.b (a1u9r2 Z6u)k,o w(s1k9i6P 2�)c,z a(l1s9ak6,ab3 ) P,� czaalnsKdkr aa cz­
kiew(i1c9z7K 2o)m,p ow(s1k9i8 2,a sw1 e9al8slZ8 a)l achanodWw is�ksi(k 1i9d 9e8a)l t
with vqaruieosutosifb o rnebsai mo laongdy fisihnte hroariseeeasS s of. a trh,Me i �dzy­
odrbzree hama s sbteuedbniy Ke odm pow(s1k9oi8n 2l)Ay st. h ossteu dhiade sc bare­en
rioeudint raa tdhiesrpt eanrti toidm ieist,ne emjeuds ttiorfi eepdte haetanm d b roaden
theisrc oSpuecan.h e ffowrta dse emneedc esisnary voifte hfawec tth thate n oticeena­ble
vironmentalt akiimpnplgra docuvere timhnleegan dstet c a(dTer zebi1at9o9cw9os)uk lid
influtheenb cieo looftg hpyeo pulianqtu ieosnt ion.
The pwroerswkea nasti mate sdt udsyoimnsege leacstpeeodcft t hMsei �dzyodrze
brebaimo logy.
MATERIAALN DM ETHODS
Thmeat eriparlosc,ei snts hesidts u wdeyrc,eo llect1e9d9 2wi-thp1ir9ni9 m7ar ily
fromc ommecracticahle sb yfie sffheecrtomefted hFn ei sheCromoepne r"aRteigvalei ca"
inGryfi now hou sedde ep-fywkatene erwt ist4 h- m-wihnigganshd 1 5-mm comdeesnhd
sizRea.n dsoammp lfreosmt hocsaet chest oytoiafel li3 dn5ed3di vai duals, supplemented
withi n2d1i1v ,mi odsujtauvlleysn iclaeusg,h te xdpureirnigfim sehntianlg l OwOixthl OaO
cm/mm1 0m esshi szceo noepat ,1 b5 etwme end-iwsitnangc ec/o5d emmmne dss hi ze
smatlrla wla,s eoantf2d 0 an d2 2m mm esh tsriammzenele (tKso mpoanwdsN keij a
199(9T)a ble 1).
Thien divciaduuwgaehlrtmse e asu(rteoldte anlgTL t ahn,sd t andlaredn SLg)tt ho,
Im man dw eig(wh1)et d1o Fgo.l lodwiisnsgeo cftt hbieoo dcnya vtihtfieys ,whe rsee xed
andt heMiaris ecram laet uriwtayds e stteargfrmeoi mthn ees dt atoefg onads,t ow eighed
0.g1.T hgeu tfitsewhde rree wei(wg2)h.Fe idsa hgw ea dse terfrmoimnann euda rli ng
couonnts sc acloelsl,fre ocmtt helede ft osfti hbdeoe d bye ltohwle a tleirnnaeelar, the tip
oft hpee ctfionran,ad el x aminuendda em ri crorfiecahdGeer ro.wr tawhta ees s timbayt ed
baccka lculTaotwiartodhn eisns.td ,hc ea usdcaalrl aed (iVum,sm )e,n lar1g7e.td5i mes
witthhme i crofichvess .atr nedalardeden (rgS tcLhm, r) e latiwoandsse htiepr amfitir nsetd
(Fi1g) .T. h ree latwiaodsne sshcirpi btehfodel lwoiwthtirhne dgei ffemraethnte matical
modeelascp,hr oduac irneglg atoiofivdtte tol hyee m pidrati(acn a 2l=7 9):
1.al ineeqaru atSLio= 0n.:2 8V+0 0 4.·6 6r25= 40 ;. 9749;
2.ab inomSiL=a -l0:. 020+ 00 1.·2V9705.·2V1r2+0= 10 ;. 9;751
3.an exponeeqnutaiStalLi= 0o.n3:·9v 0 8·92614; r 2= 0 .980.
'fab1l e
Liosfmt at ercioalll aencadtn eadl pyesrefosr med
Tvooefa nalysis
Dataenl do caolfci attcyh Fishgienagr( L rSLeanc,ngm ge)t t..., i:::Q.)h sQ.)Q.)i::: ::I § ... gJ i:3:i:::bf)Q.) "O-«l"'§bf)i:::()� - ..... e«l�Q.) s <!)i::: � "Cls"Clcoro§bf)§)<' ·"Cl·..s-0i3E<!) .p i:::€ ."@i."gc:,bf)�i:::0 Q) s
� �
Commerccaitaclh es
05N o1v9 9W2i,d uchowa fyknee t 28.5-3l9l. 51 1 10 11 9
l DOe 1c9 9W2i,d uchowa " 35.2-363. 33 3 3 3
26A p1r9 9N3a,m yslin " 29.7-431.78 17 9 17 17
27S e1p9 9O3d,r aZachodni"a 30.7-3151. 81 1 2 11 11
28O c1t9 9M3i,� dzyodrze*" 11.0-1182. 41 2 12 - -
10M a1r9 9W4i,d uchowa " 25.0-3171. 81 1 9 11 11
26A p1r9 9M4i,� dzyodrze· " 31.0-4234. 42 4 14 24 24
25O c1t9 9M4i,� dzyodrze*" 22.3-3317. 73 7 33 37 37
27M a1r9 9S5i,e kii erk " 8.8-252.33 2 3 5 23 7
17A u1g9 9B5e,r lCiannsakli " 30.5-373. 23 - 3 3
28M a1r9 9C6i,e Cpalnya l " 14.0-2141. 91 1 1 11 11
12A p1r9 9T5a,mP ao morzans,k,a 20.9 1 1 - I -
18A p1r9 9G6ry,fi nsCkain al ,, 27.5-348. 78 l 8 8
24A p1r9 9C6i,e Cpalyn al " 7.1-247.29 7 2 59 72 20
30A p1r9 9S6t,aR reag alica " 8.2-231.88 1 8 18 18 5
30A o1r9 9G6r vfinCsaknia l " 20.4-3951. 59 1 45 91 79
TOTAL 7.1-4335.383 53 221 3412 45
Exoericmatecnhteasl
10M 199D3l,u Cgain al scoonpe t 5.5--6.75 7 6 - -
14 ay 1J9u9nG3 ryfi,n skCia nal ,, 6.7 1 1 1 - -
21J u1n9 9M4a,lR ae galictar amnmeetl1 2.0-222. 12 2 - -
13J u1l9 9P4o,d kCoawnaa l " 20.3 1 I l - -
31M 199B5o,b rCoawyn al " 26.1-393. 83 3 - -
12J ayu1 !9 9D5l,uC gain al ,, 12.0-182. 32 2 - -
13S e1p9 9G5r,y finCsaknia l " 23.1 1 1 1 - -
09O c1t9 9D5l,uC gain al " 17.0-212. 32 2 - -
16M ay1 9,9P 6odkCoawnaa l " 11.7-40.35 3 3 - -
23J u1i9 9S6t,aR reag alicas mall t2r.a4w-l3 .21 - 2 - -
06A u1g9 9S6t,arR ae galica " 2.9 2 - 2 - -
27A u1g9 9D6lu,g Cia nal trammneelt 20.9 1 1 1 - -
29u gA1 99S6t,aR reag alica " 30.3 1 1 1 - -
29A u1g9 9S6t,aR reag alicsam atlrla wl 3.4 1 - 1 - -
27S e1p9 9D6l,u Cgain al trammenle t 10.7 1 1 1 - -
13M 199D7l,uC gania l small t4r.a9w-l6 .70 - 7 - -
23M ay 199D7l,u Cgain al " 3.9-5.194 - 14 - -
03J ayu1 i9 9S7t,aR reag alica " 1.9-2.178 ·- 1 - -
03J u1i9 9D7lu,gC ia nal " 1.9-31.345 -· - - -
02S e1p9 9D7lu,g Cia nal " 3.8 1 - 1 - -
02S e1n9 9S7t aRreag alica ,, 4.4-4.67 - 6 - -
TOTAL 1.0-402.151 25 58 - -
GRANTDO TAL 1.0-4536.483 78 279 3412 45
Nod etadialatebado c uattl cohc ality
••15i3n divmiadkuiaunlpagsr ,a ndsoamm pplieco kuefrtdo mat otoaf7l 0 7w,e rmee asured
6 ZbignNieeajwA, n drKzoemjp owski
45
40 ....... . ..
..
35 ..�..,•s .. • - �,;..... ·
30
E
� 25
SL= 0 .3V90·89 26
20 R2 =0 .980
n= 2 79
15
_A••.
.
to
/
�;
0 20 40 60 80 100 120 140 160 180
V( mm)
Fi1gR..e lationsh1i7p.m Sabxge ntwiseficeeanrdl a ed( iV)ua snb do dlye n(gSLt)h
ofb refraommt h Mei �dzyodrze
Thbee fistwt a osb tained wietquhat itaohsene v, i thdiebrnyidc t hesid g choeesftfi ­
cieondfte termi(n2r)aF.to tirho rines a tshoeenx ,p onceurvnept lioatl w(aFusis gfoe.rd 1)
thes ubsebqauccekan ltc ulTathlieeo nnogsffit. sh h d iiffne yreearonsflt i wfea rse con­
strufrcotmthe esd c aclauedr aald iincu osn segcrutoizwvotenh( e Vnsm) e,a siuanrm eand­
neird enttiotc hauatls foedrt hteo stcaalrl aedc iourrse,s ptootn hdleie nnoggfa tg hi ven
indivTihdreua adwlia.uths se dni vibdyte hadec tusaclar laedo iftu hsie n diveixdamu­al
ineTdhr.ea toibot awiansseu db sequuseeansdact olryr efacctttioobor enm, u ltibpyl ied
ther adoifaug si vgerno zwothn Teh.le e ncgtho rresptootn hdceio nrgr reacdtwieaudss
refraodmt hpel ot.
Thgeo nado-isnodm(iattcFiheucesl itnodnGe Sx/io, rt hCel arikn dGShe)x ,w ere
calcfruloatmte hfoder mula:
GS! = G-100
Wn
wheGr,ge o:n waedi (gghw)1to ; wr 2: t owteail(g 1w)ho trg u twteeid(g wht2 ).
Whedne tertmhilene inatngfi gtr hms attu rtihtfieys w,hi gtohn sahdosw atil netga s
Maiesrc aslteaII wg eera es sutmboee md a ture.
Somdea otnta h e boifco olmomgybo rne am 7
RESULTS
SLv .sTL reiloantship
Althtohusegt han dlaredn (gSLt=h l .hca.bs)e euns eidnm o sbtr esamt udsioemse,
authorresp otrthteeod t laeln( gTLt= hl .)Tt..o r enthdeet rw doi mencsoimopnasrt ahbel e,
relationsthhiepwm a ssbt eutdwTieheeredne . l atiocnanbs ehd ieps crwiibtaehl d i near
equaTLt i=1o n. 212+51 ·.SL3 2w7h4is chho awv se ryg oofidtt ot heem pirdiactaaasl,
evidebnyic tcesod e ffiocfdi eetnetr mri2 n= a0 t.i9Ho9on7w,.e vteherex, p oneeqnutai­al
tiTLo n= l .420·496·4pSL5r oduac esbdte ittlfielt(r 2r = 0 .99I9ct)an b. e t herceofonr­e
clutdhethadetr elatbieotnwstehhseietnp an danarddt hteo lteanli gsth slciurvgihltilnye ar.
Frlyne ght
Tab2ls eu mmarisdeatsao n n umbejrusv eobnfri el(amea gger o0u)ip n d ifferent
len(gStgLhr) o iuntp hsee x perimenOtna lM1 a0cy at c1ah9t e9os3to.,af6 jl u veinnidlie­
vidumaelass uri5n5g-6 m5m w erceau gwhittt hhs ec onoepTt h.e sicra lesan anl­acked
nuarli nSgi.m iwlearrs ec aolfe s2t 1hi en divmiedausaurl3is9n -gm6 m0,c auwgihttah
smaltlr aiwnMl ay (132 31a)9n 9de7 x,c foerpa st i nignldei vmiedausau5lr7m i mnth ge
scaolfwe hsi schho wace lde varilsyani nbulraeil on ngt heed gAelt.lh oisned ivwiedruea ls
ofm oroerl etshsse am ea goef1 y earh,e ntchee irp rloevniigndtfoehrms at oinot nhs ei ze
attabiytn hbeerd e iamnt h peo puluatnidsoetnru d dyur itnhge iyre oarfirfl s itfe .
Table 2
Lenfrgetqhu ednicsyt riojbfuu vteibnorinelfrsaeo m mt hMei �dzywoatderrzse
Samppleirnigo d
SL( cm)
Ma1y9 93 Ma1y9 97 Ju1i9 97 Au1g9 96 Se1p9 97
2.0 - - 84 - -
2.5 - - 68 - -
3.0 - - - 2 -
3.5 - - 1 1 -
4.0 - 2 - - 1
4.5 - 2 - - 6
5.0 - 2 - - -
5.5 2 12 - - -
6.0 2 3 - - -
6.5 2 - - - -
Total 6 21 153 3 7
Al argseam ploef1 5m3e asurineddi vpiidcuokauelatdsrt andofrmo m7 0j7u venile
brecama uwgihthtas matlrlao wn3l J1u9l9yw7 a fosun dt oc onsoiffis stmh e asuring
19-27( mmomdv aallo ufe mm 2)2.T hseam plceo ntaailanns eoed x ceptliaroginena ­lly
dividual3 4mm ema.Ms ousortfi ndtgeh lei sccataeol fthe oss sem ablrle aml olfeff o n
8 ZbigniNeewj Anad,r zKejo mpowski
captTurhee. restcaaoilfane2 es7md m l onign divsihdouw4ae-lds6 c lethreiictrae usd,a l
radlieunsbg etihan bgo0 u.mt3m .T husd urtihn1eg- 1m.o5n othf lsiifen hcaet( cMha y),
thien diveixdaumaihlnasged rd o twaontt aani anv erlaegnoegf 2th 2m m.
A smaslampll ec,o nsiosf1t 0ii nndgi vciaduuwgaihltttsh h s em atlrlai wnAl u gust
199an6d i ne arlSye pte1m9b9ae7lr l otwoae sds umtheat th jeu veonfith lepe osp ulation
studwieeradeb tloge r otwo2 9-4mm7 ithne s echoanoldfsf u mmTehre.si cra les showed
13-s3c3l etrhiceta eusd,a lr anrgaiwdniigtu0 hs.i 3n4 -m0m..80
Lineagrr owth
Tab3ls eh omwesan l engathttsa ibnye dMth iey dzybordreziamend i ffeyreearonsft
lirfee,c onsfrtorumbc atcekd- srceaaNldoe tse.w oirsact ohnys idleernargbanthlg reee ­
coridnei dn divoifid dueanlatsgi Secp.ae lc ifitchlaeel nlrgyant,ghr e e consfotrrut chtee d
firysetoar f l iwfea 3s. 9-c9m.t,5h c eo efficoifve arnita atcicoonu fonrt1 i8n.og6f %t he
mean( 6.c1m7)I .nt hsee coyneodarf l itfeh,fie s h gtro6 e.w8 -c1m5t,.h m9ee alne ngth
andc oeffiocfvi aernitba etii1no0gn. c3m8an d1 5.7r%e,s pecItnti hvseeu lbys.e quent
yeartsh,le e nrgantghbe e caemveew ni dtehrde:i fferbeentcwtheee eup np eanrd l ower
valoufte hr ean gwea 5s. c6mi nth fier ysetanar d i ncrteo9a .s10e2,d.1 66,.an 1d1, 8 c.6m
inth es ecotnhidrfo,du ,r athn,fid ft yhe arr,e specOtnti hvoeet lhhyean.rd t ,hc eo efficient
ofv ariadteicornew aisateghtde h ,eu vsi deanr ceilnatrgie vdeu ocfvt ariioanb ility.
Meanan nuall enignthc rewmeernfoetu snt do m onotondieccarwiletlayhsa eg e
(Tab3l)et, h umsa kitnhvgeo Bne rtagrloawthnf cfyurv et ofi tw eltlhe em pirdiactala .
Wheens timLatooti,hnv ege hriygc ho rrelcatoieoffinc( ire n=t w 0a.os9b 9t9a)ti hnceeo dr,­
relactoieoffinco ifte henest t iKmanadtt0 eb de ir=n - g0 .999.
Thfoel lowpiarnamge tweerrse arrived at:
L� 9=. c35m K;= 0 .09t=9o-9 0;. 037yr.
Toe stitmhaoptsareeam etmeerasln,e ngithnas gg er oIup-sXw Ie rues eLde.n gitnh s
oldgreoru wpesrd ei sregaarstd hseeamd p,ls eiw zaets o som all.
Weig-hltenrgetlha tiaonnwdse hiiggphr to wth
Thwee ight-rleelnagtthfoi ortn hsMehi i�pod drbzree waamds e scrwiibttehhd e
zy
following exponential equation:
w= O .Ol23166·3r8SL2=; 0 .9.9 7
Bys ubsttihtmeue tanli enngga ttthaisn eddi ffieynre earonsflt i (fefr oTma b3l)ie n to
thee quatthieoc no,rr espmonedanwi eniggw hetroseb ta(iTnae3bd)l B.eys ubstituting
Loo= 59c.m3th, e w eiogfwh oot= 509g7w .a3cs a lculated.
Somdeao tnta h bei oolcfoo gymm on bream 9
Conditn io
ThFeu l(tKoa1nn) Cd l ar(kK2 c)o ndictoieofnfiw ceirecena tlsc usleatpeadrfo art ely
eacihn diveixdamuianTleh dre.e swuelrtgesr oubpye dans det xh sep ranidna gu tumn
valwueersce o mpa(rTead4b )lT.eh seum meanrd w invtaelrwu eerse disdruteeog arded
thes maslamlp lsei zTehc.eo nditionv sfic.so lhee ffincrgietelhntat iiopsnr sehsieipntn e d
Fgis2.-af.
Ins pritnhfegem, a lsehso weKd rtahntegov e ary from1 .6t7o22 . 7t4h8em, e an
1
valbueei 2n.g1 F9i02g.a. d epiarc attsch oenrs idienrcarobeflKa1 es w ei tfihs lhe ngth.
The rwaindogefve a liunde isff erent cmloassltsi ekfrseo lmtry he faesc tuthl tatth ededi f­
fernetb ream cfeamauilgnseh pstr wienregei tfuhleslrp, a wnionrsg p,eh netn,dc ieff erent
matusrtiatogyfte hseg iorn afrdosmM: a isecras lteaV gIte oV IITIh.ii scs o nfirbmyae d
mucnhar rorwanegroe ft hCel arcko effic(iKes2ne,t e 2FbiT)gh.K.e 2 r angiens prwiansg
15.56-2(.m1e6an13:. 8(6T9a)4b )lA.ess hobwny F i2gb.th, e c oeffigcriesewln otw ly
witthheiSL n r angoef1 3-an2d5e quaslllodywe lcyr etahseerde after.
Ina utumtnh,fee m alKev aluvearsiw eidt h1i.n8 73-(2m.e5an29:.5 1 2t7h)e,
1
autumnv albueeistn hgus so mewhlato wtehrat nhs ep roinnegTs h.me e anfe malK2ei n
autumnw a1s. 8i9.4oe,n. ls,yl ighhtilgyth heanir ns prianndgw itanh ar rorwanegroe f
variati(oTna4sb) l.e
Inm alKe1s r,an geidns prwiintg1h .i9n3 0-3.m0e4anv6 a,l (ut2eh. e2b 8e7i)n g
hightehrant hatth feeim na lTehswe.i desto fKs cvaalttueanerdst hhei gheexsttr eme
1
valwueersre e coirrtdt ehdom sael meesa sur2i6n-gc3 m4( Fi2gc.T) h.se am el ength
claspsreosd uaclesthdoeh ighveasltu eKs2an dot fh ehiirgh evsatr ia(bFii2lgdi.)t .y
Ass hobwynT ab4l,e v K2ar iienmd a lweist 1h.i8n8 2-t2hv.ea0 l7bu7ee,is thun sgh igher
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Ag(ery .)n Li L2 L3 L4 Ls L6 L1 Ls L9 Lio L11
a 7.28
1 21 b 5.1-8.9
C 1.12
a 6.09 10.19
2 33 b 4.0-68..20 -13.4
C 1.10 1.46
a 6.44 11.091 5.34
3 17 b 4.4--8.86. 3-1141..67 -19.7
1.14
C 1.72 2.16
a 6.20 10.011 4.221 7.89
4 23 b 4.6-87..74 -1140..76 -1173..85 -22.7
C 1.12 1.65 1.84 2.03
a 5.66 9.94 14.911 8.972 2.21
5 52 b 3.9-87..50 -1150..86 -2114..95 -1264..24- 29.9
C 1.06 1.61 2.23 2.41 2.71
a 6.02 10.301 5.191 9.702 3.522 6.48
6 26 b 4.2-77..84 -1120..51 -1242..47- 3107..08 -2314..68- 35.7
C 0.89 1.48 2.77 3.77 4.07 3.90
a 6.23 11.081 6.182 0.442 4.342 8.143 0.81
7 24 b 4.6-8.75. 5-1143..39 -1168..58- 2205..30- 2320..91- 2344..22- 36.8
C 1.07 1.55 l4.3 2.53 2.75 2.79 3.17
a 6.14 10.811 5.362 0.432 4.212 7.963 1.353 4.09
8 14 b 4.4-97..58 -1151..28 -2115..17 -1298..4--63 203..05 -2373..73- 3306..03- 39.5
C 1.30 1.90 2.43 3.68 3.23 3.05 2.51 2.86
Tabl(ec o3n t.)
Age( ry). n L1 L2 L3 L4 Ls L6 L1 1..,T 3 L9 Lio Ln
a 5.79 10.421 4.751 9.182 3.592 6.943 0.613 3.473 5.46
9 15 b 4.8-76..48 -1122..24 -1169..26- 1293..39- 2217..47- 2351..55- 2387..45- 3309..72--4 0.5
C 0.78 1.52 2.12 2.31 2.52 2.91 3.27 3.56 3.39
a 6.31 10.641 5.101 8.712 2.822 5.992 9.453 2.413 5.453 7.40
10 11 b 3.9-88..47 -1131..53 -1149..76- 1282..09- 28.6 2240..22--233541...920- 2397..15- 3318..38--4 1.5
C 1.28 1.63 2.71 3.30.20 9 3.39 3.27 3.73 3.19 3.21
a 7.15 10.881 5.071 9.882 4.082 8.123 0.983 3.873 6.683 8.534 0.30
11 6 b 5.8-88..62 -1142..35 -1250..3l- 1293..57- 2247..19- 2371..14- 2393..78- 3327..50--4 304..25 --4317..92 --43.4
C 1.22 2.16 2.77 3.35 3.18 3.15 2.99 3.36 3.35 3.04 2.64
12 1 a 4.80 8.90 11.901 7.101 9.402 2.802 7.202 9.003 1.703 4.003 5.30
13 1 a 5.50 8.30 15.302 1.102 6.302 8.603 2.103 4.803 7.203 7.703 8.70
14 l a 7.00 10.301 3.601 7.202 1.202 5.802 8.203 1.30 32.603 5.503 7.00
a 7.10 11.001 5.651 9.302 1.902 5.552 7.103 0.803 3.803 5.903 8.30
15 2'b 5.9-81.03. 0-1142..80- 1186..65- 20.0 2241..30--226-2..88 - - - -
C 1.70 1.41 I.20 0.99 1.27 1.77 - - - - -
a 6.17 10.381 5.081 9.282 3.202 7.153 0.573 3.303 5.483 7.413 9.11
Tota2l4 7b 3.9-96..58 -1150..81 -1232..97- 1360..20- 2304..28- 2345..27- 2357..95- 2399..15-- 3410..35--4 315..93 --43.4
C 1.15 1.63 2.24 2.384. 11 3.26 3.03 3.29 3.19 2.95 2.65
V 18.601 5.701 4.901 4.701 3.401 2.00 9.90 9.90 9.00 7.90 6.80
M 6.17 4.21 4.70 4.20 3.92 3.95 3.42 2.73 2.18 1.93 1.70
W1 3.99 20.676 7.581 46.0266 2.2443 4.1632 9.2862 2.8180 05.14188 8.17306 7.95
n 247 226 193 176 153 101 74 50 36 21 10
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