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NOTES ON ERITRICHIUM (BORAGINACEAE) IN NORTH AMERICA PDF

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Preview NOTES ON ERITRICHIUM (BORAGINACEAE) IN NORTH AMERICA

NORTH AMERICA ON NOTES ERITRICHIUM (BORAGINACEAE) IN of Alaska Museum of the North University 907 Yukon Drive Fairbanks Alaska 99775-6960, U.S.A. a and of selection how Wight which of the Bering Strait region, is plant cushion-forming nail he had been led asttay that believe canne to however, Hulten (1973) saw ami then Hnlten (1968). Later, (1902) it fmm the exerted within or barely included with flowers by Wight who applied E. chamissonis to the plants he decide by two Fust, about errors. was brought thinking change in cushion stems and Hulterfs of leaves. PopoVs tllustratton of aretirn- DC, (1953) because named (Cham.) lhata cushion form should be E. onrlioides specin^o the type believed Hulten and second, . des- showed a densely pulvinate plant (Plate 25, fig. 1), (IWL de Candolle descnption of odds with the original which ctonissonis had long flowering stems, is at Po^v unknowmglytllus- had determined that and Petrovsky (1980) As to the assumption, Jurizev first hadgonennrecognmeduntfl that pulvtnatespecies,not£ritrlchi»umtioides,butone t«tedadistinet. densely tepubllcationofE.,schuhtschor„mJurtzsevSrW.Petrovsky,whichonleafmorphologyalotw«re^^^^ rable from both or chamissonis. E. aretioides E. uretiofc CamioUe ofEritrichiun. by de As descriptions second assumption, the to the “ made he clear that E. nevertheless, te and two not well differentiated; brief, the species are « ^ no er E anttois chumissonn at of the flowers plant with the shorter stems of the two. In fact, ^rfaceofthetuft,whereasEuretioldeshasfloweimgsteins,mlUto«^ flowering the length ot in the densely compact, from chamissonis but differ E. account. flowers. Hulten had no reason doubt Wight’s to n- chased on sneci- ncnni awi i., De Candolle (Prodromus 10:125. 1846) De 1816 or 1817. voyage of the Rurife during the Eschscholtz mens Chamisso or by originating LE, collected at ad <«nrti-Laurentii fre- unambiguously to St. Candolle and referred unequivocally ^ and thus localities “inKamtchatkaa^ t«mBeering(Cham.O”but continued then new spe two his specimens, or syntypes, were the basis for 18 & 4.442. Linnaea ) had Schlechtendal, y reported (Chamisso ^ ^ earlier Chamisso in his e were regar Chamisso ously not named by y all of the plants so reference to Lawre was y- St. viUosa M. account was his clear that the locality for Kamtchatka. provided by G, then images digital with excellent AreviewofthetypesheetatG-DCwasundertaken,firs Kamchatka, from collector The^one G-DC ^ a visit to G. Mounted on the type sheet at This speci- ^^^^ription). ^ j li ac h. unknown, by a colleague at ( provided de Candolle to ^ ®cn strong with and its IS labeled Myosotis Ledeb., y resem- villosa instead clearly exertis,” it pauciiio racemuhs flescription of Eritrichium chamissonis: “. '•««-lte.hBtT«as7(1):3 352 Why bles E. villosum (Ledeb.) Bunge. This specimen cannot be considered as E. chamissonis. de Candolle would make Kamchatka specimen own a direct reference to the is a curious contradiction of his description. The other specimen on the type sheet, also labeled Myosotis villosa, is a specimen that matches the de- scription of Eritrichium chamissonis as the flowers barely exceed the cluster of sterile, leafy shoots, and it is clearly attributed to Chamisso. However, one Lorenz meaning this labeled Lawrence is “St. insel” St. Island,. This specimen, while a perfect match with the description of de Candolle, is in conflict as to location. However, the Rurik visited both St. Lawrence Bay, Chukotka, and St. Lawrence Island, Alaska, during voyages. its De Candolle took the locality Lawrence Bay directly from what Chamisso Chamisso and (in Schlechten- had dal 1829) written under Myosotis villosa. Yet, neither of the two specimens on the type sheet from is St Lawrence Bay. That de Candolle add Lawrence failed to St. Island to his description, despite the label on the sheet before him, must have been a lapsus calami. It is otherwise impossible to reconcile the discordance. An herbarium LE sheet bearing at Chamissio’s signature breve, labeled Myosotis and from Law- villosa, St. rence Bay, would seem be what to is referred to in the description of Eritrichium chamissonis, but it is clearly not consistent with inasmuch as the flowering stems it, are tall, leafy, rising well above the basal leaves, thus closer in habit to E. villosum or E. aretioides than to E. chamissonis. That sheet cannot be considered as the basis forE. chamissonis. Another LE sheet has been labeled, incorrectly, as holotype by Ovchinnikova The specimens (2008). are attributed to Eschscholtz. Ovchinnilova (2008) reported the locality as “Sin. Laur[enti]”; the handwriting St. on the original labels attached Who to that sheet not easy known is to decipher. wrote that note not (VY. is Petrovsky, pers. comm.). There no LE G-DC evidence is at or that this material was seen by De Candolle. Not LE found at is a sheet with specimens of Eritrichium chamissonis from Lawrence Island from which St. the^specimen on G-DC the type sheet at might have been G-DC The taken. source of that specimen remains at Nevenheless, de Candolle did have him before and most he certainly based his description of Eritrichium chomis^is on the Alaskan specimen. De Candolle did not no designate can be holo- a type, therefore, there The type. specimen, a syntype, on the type G-DC sheet from at Lawrence here designated St. Island, Alaska, is as lectotype for E. chamissonis DC. Eritrichium chamissonis DC., Prod. 10:125. 184 mixed olJes with M. andXhom vIBostt, a. Lawrence St, Bay, Chukotka, St two were ItKalities given by Chamisso, nameM. the imdesmustbeb^ synw« ate, on one or more of the and a lectotypification is necessary. Ovchinnikova (2008) an cited Eschscholtz specimen from Lanrcntit Lawrence St. Island Clns. St. “ “ ' “miouks. Although t her incorrect usage of the term ‘hololype' , notlec^^rrelT-trS^r^t^^^ fotn«i name needs a .he unre a e to aretioides. Another from is the Academy label printed in St Petersburg (LE) with a Myrnmuarntt^des, Sinus UurentiLbutacolfectot St. not tecord^l'Thf^^^^^ hB would be better as the type M. for orehoides. Wheteas ha« would a specimen LE Chamisso seen by at 353 G-DC no Chamisso specimen LE from which one might have been taken or the there at at standing, is greater specimen G-DC was determined by Chamisso represents Chamisso’s intentions. Since the at better one that furthermore connects Chamisso to de Candolle, that specimen is designated here as lectotype. and ACKNOWLEDGMENTS Andrei Yu. Razzhivin, Sy- my Komarov Petersburg, V. Thanks colleagues at the Botanical Institute (LE), St. to Chamisso and Eschscholtz specimens there; to on YN. Petrovsky information the and especially to for tin, making Manager Geneva possible for and Fumeaux, Collections at (G), for it Nicolas Laurent Gautier, Curator, am my and specimens. grateful to re- much view types associated 1 me accomplish so during short visit to to has James Reveal who my and generally improved the text of this note. very kindly corrected errors viewers authorship of the and Kanchi Gandhi clarified the read and commented on the manuscript. Reveal carefully names and descriptions, author of the Eritrichium DC, the Eritnchium names: A.P. de Candolle or the father, DC, de Candolle or A. the son. notwithstanding the page heading that credits A. Dep Chamisso, A.L von and von Schleci 1829. D.F.L. . rationem dicunt. Linnaea 4:435-496. enumerato coi vegetabilis sive regni DeCanooue, 1846. Prodromus systematis naturalis A.P. normas digest. methodi naturalis specierumque plantarum hucusque cognitarum, juxta and neighboring HultEn, E. 1968. Flora of Alaska an HuLTfN, E. 1973. Supplement to flora of Alaska nomenclature for McNhu code botanical et al. 2012. International of Congress opted by the 18th International Botanical ^ A world of Asiatic North It species in iginaceae) genus Ovchinnikova 2008. Conspectus of the Eritrichi S.V. Russia 1:17-36. Russian] [in 8^38-245. BA Arctic Flora USSR. tin .nd Jurtze., eds. ln:T0lma.he», A.I. Jurtzev,B.A. AND V.V. Petrovsky. Eritrichium. Moscow & Leningad. Russian] [in Nauk. Akad. 9:482-521 USSR Popov, M.G. Flora 1 . 1953. Shishkin, ed.. Eritichium. B.K., In: 29:407-414. Club Torrey Bot. Wight, W.F. 1902. The genus Eritrichum in North Americia. Bull.

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