BIOMES OF THE EARTH GRASSLANDS Michael Allaby Illustrations by Richard Garratt Grasslands Copyright © 2006 by Michael Allaby All rights reserved. No part of this book may be reproduced or utilized in any form or by any means, electronic or mechanical, including photocopying, recording, or by any information storage or retrieval systems, without permission in writing from the publisher. For information contact: Chelsea House An imprint of Infobase Publishing 132 West 31st Street New York NY 10001 ISBN-13: 978-0-8160-5323-0 ISBN-10: 0-8160-5323-5 Library of Congress Cataloging-in-Publication Data Allaby, Michael. Grasslands / author, Michael Allaby ; illustrations by Richard Garratt. p. cm.—(Biomes of the Earth) Includes bibliographical references and index. ISBN 0-8160-5323-5 1. Grassland ecology—Juvenile literature. 2. Grasslands—Juvenile literature. I. Garratt, Richard, ill. II. Title. III. Series. QH541.5.P7A38 2006 577.4—dc22 2005005615 Chelsea House books are available at special discounts when purchased in bulk quantities for businesses, associations, institutions, or sales promotions. Please call our Special Sales Department in New York at (212) 967-8800 or (800) 322-8755. You can find Chelsea House on the World Wide Web at http://www.chelseahouse.com Text design by David Strelecky Cover design by Cathy Rincon Illustrations by Richard Garratt Photo research by Elizabeth H. Oakes Printed in Hong Kong CP FOF 10 9 8 7 6 5 4 3 2 This book is printed on acid-free paper. From Richard Garratt: To Chantal, who has lightened my darkness. CONTENTS Preface ix Acknowledgments xiii Introduction: What are grasslands? xv CHAPTER 1 GEOGRAPHY OF GRASSLANDS 1 Where grasslands occur 1 Temperate grasslands and tropical grasslands 3 Prairie 4 Steppe 7 Pampa 10 Veld 12 Tropical grasslands of South America 14 Savanna 17 Australian grasslands 20 Upland grasslands 22 CHAPTER 2 GEOLOGY OF GRASSLANDS 26 Movement of continents 26 Alfred Lothar Wegener and continental drift 28 How mountains rise and wear away 31 Grassland soils 37 How soils are classified 39 Laterite 40 Water and grasslands 41 CHAPTER 3 GRASSLAND CLIMATES 45 Why there are seasons 45 Continental and maritime climates 48 How climates are classified 49 Dry seasons and rainy seasons 51 The Dust Bowl 55 Monsoons 57 Adiabatic cooling and warming 59 El Niño 61 Convection and storms 64 Lapse rates and stability 66 Tornadoes 67 CHAPTER 4 HISTORY OF GRASSLANDS 71 Evolution of grasslands 71 Grasslands and past climate changes 73 How forest can change into grassland 76 The transformation of New Zealand 78 CHAPTER 5 LIFE ON THE GRASSLANDS 81 What is grass? 81 How grasses work 84 Photosynthesis 85 Prairie grasses 89 Pampas grasses 92 Steppe grasses 94 Savanna grasses 95 Grassland trees and shrubs 99 Grassland herbs 100 Grassland insects 103 Mongooses, prairie dogs, marmots, ground squirrels, and pocket gophers 110 Snakes and lizards 114 Grazing animals 119 Hunters of the grasslands 124 Hunter and prey: The evolutionary arms race 130 Grassland birds 130 Coping with drought 136 Coping with heat and cold 139 Hibernation 141 CHAPTER 6 GRASSLAND ECOLOGY 143 How the plant eaters help the grass 143 Food chains and food webs 144 Ecological pyramids 147 Do predators control their prey? 151 How herding provides safety in numbers 153 Mammal migrations 155 CHAPTER 7 PEOPLES OF THE GRASSLANDS 158 Peoples of the prairie 158 Homesteaders and the way the prairie was transformed 160 Indians and gauchos: The peoples of the pampa 165 Farmers of the pampa 167 Peoples of the African savanna 169 Traditional life on the steppe of Central Asia 171 Genghis Khan, the Golden Horde, and Mogul emperors 173 Aboriginal peoples of the Australian grasslands 177 CHAPTER 8 USES FOR GRASSLAND 179 Cereal farming 179 The origin of cereals 180 Cattle ranching 182 The origin of cattle 184 Sheep farms of Australia and New Zealand 186 The origin of sheep 189 Upland sheep farming 190 Forestry 191 Biofuel production 194 Biofuels and the greenhouse effect 195 CHAPTER 9 BIODIVERSITY AND GRASSLANDS 198 What is biodiversity? 198 Why it matters 199 Protecting grassland species 201 The Biodiversity Convention 202 CHAPTER 10 THREATS TO GRASSLAND 205 Conversion to farmland 205 Conversion to forest 207 Overgrazing and soil erosion 209 Climate change 211 The greenhouse effect 212 Expansion of towns and roads 214 CHAPTER 11 MANAGING THE GRASSLANDS 218 Lessons from the Dust Bowl 218 Ranching on equatorial grasslands 220 Farming tropical grasslands 222 Dry farming 224 Grassland restoration and conservation 225 CONCLUSION 228 What future for the grasslands? 228 SI units and conversions 231 Soil classification: Orders of the soil taxonomy 235 Glossary 239 Bibliography and further reading 253 Index 257 PREFACE Earth is a remarkable planet. There is nowhere else in our solar system where life can survive in such a great diversity of forms. As far as we can currently tell, our planet is unique. Isolated in the barren emptiness of space, here on Earth we are surrounded by a remarkable range of living things, from the bacteria that inhabit the soil to the great whales that migrate through the oceans, from the giant redwood trees of the Pacific forests to the mosses that grow on urban side- walks. In a desolate universe, Earth teems with life in a bewil- dering variety of forms. One of the most exciting things about the Earth is the rich pattern of plant and animal communities that exists over its surface. The hot, wet conditions of the equatorial regions support dense rain forests with tall canopies occupied by a wealth of animals, some of which may never touch the ground. The cold, bleak conditions of the polar regions, on the other hand, sustain a much lower variety of species of plants and animals, but those that do survive under such harsh conditions have remarkable adaptations to their test- ing environment. Between these two extremes lie many other types of complex communities, each well suited to the particular conditions of climate prevailing in its region. Scientists call these communities biomes. The different biomes of the world have much in common with one another. Each has a plant component, which is responsible for trapping the energy of the Sun and making it available to the other members of the community. Each has grazing animals, both large and small, that take advantage of the store of energy found within the bodies of plants. Then come the predators, ranging from tiny spiders that feed upon even smaller insects to tigers, eagles, and polar bears that sur- vive by preying upon large animals. All of these living things IX