Four New Species of the Group of Trichotichnus leptopus (Coleoptera, Carabidae) from Central Japan and Shikoku, with Some New Records

2005·7.3 MB·

by  Ito, N

E、l'Ira,T()k、・().33(2):629-640、No、,cmbcr19.200・,
FourNewSpeciesoftheGroupofTrichotlchnusleptl)pus(Coleoptera,
Carabidae) fromCentralJapanandShikoku,withSomeNewRecords
Noboru ITO
1-7-18Higashiuneno,KawanishiCity,HyogoProf.666-0117Japan
Abstract Fournewspeciesofthc/ep1optlsgroupo「thegenusT;・lc/fofl(・/1,1usare
describedunderthenamesof「1・lc/toric1/1111.i-(「1・lc1/J(rlc1/11ll.l,)a1,1clglsc,,111sfromMI.Amagi_
San.「.(7.)/1aktlsan1lsfromMt.Haku-san,T.(T)111as(,taka、・os/1ltla1fromMt.Tsurugi-san
and「.(「.)1,0.1'1/111'll、/'11fromMt.Mitsuji-yama.Also newlocalitiesof「.(「.)1.l,a,川Ila,1c1/,,c11'
N. ITO, 「.(T)s/lit'oktlellslsKAS・、ll・、RActY.ITOandT.(7.)1le11o1-t1l11KAsAHARAclY IT-()
are recorded.
Speciesof thelept)pusgroup(1973)of thegenusT1・1chotlchnusMoRAwlTzare
welldiversifiedineachnarrowregionduetoapterism.Someworkershavestudiedthe
speciesofthegroupandrecentlydescribedmanynewspeciesasmentionedinmy for_
merpaper(2005).Ihavebeencontinuingtoexaminemanyspecimensfrommany10_
calities.AmongthemI foundonenewspecies fromMt.Amagi-saninShizuoka,one
newspecies from Mt. Haku-sanin Ishikawaandnearby area,and twonewspecies
fromShikoku.
Inthispaper Iamgoingtodescribethe fournewspeciesas follows:Trichotlch-
nl!s ( 'lc/7oflc/m's)emaglsa?' fis from Mt. Amagi-san, 「. (「.) /Ia llsa1711s from Mt.
Haku-sanandIkegahara,T. (T)111asat(1kayoshldai fromMt. Tsurugi-sanandT. (T)
1'os/11j'u/al fromMt.Mitsuji-yama.Further Inewlyrecord「. (「.)1samlra11a/fa N. ITO
fromKyoto,T. (T)shikokuenslsKAsAHARAetY.ITOfromTokushimaandEhime,and
T.(T)shlkokuenslsKAsAHARAetY. lTofromEhime.
Beforegoingfurther,Iwishtoexpressmyspecialthankstothefollowingtwoex-
cellent loversofCoIeopterainShikoku, Mr. Yoshiyuki ITO, Kochi andMr. Masataka
YosHIDA.Tokushima.Mr.Y. ITOhasbeenvigorouslycollecting forrespondingtomy
requestandkindlyofferingmemany importantspecimensformystudy,andMr.M.
YosHIDAkindlyguidedinfieldworkonMt.Tsurugi-sanandhasbeenofferingnumer-
ous importantmaterial.AlsoIheartilythankMr.MasaharuTAKABA, Kanazawa.Dr.
HidetoHosHINAofFukui University, Fukui,andMr.Seiji MORITA, Tokyo for their
kindofferofinvaluablematerial,andtoDrs.Shin-ichiYosHIMATsuoftheNational In-
stituteofAgro-Environmenta1Sciences,TsukubaandShuheiNoMURAoftheNational
ScienceMuseum(Nat.Hist),Tokyofortheirkindlyloaningmaterial.Iemploytheab-
breviationsofdepositoryasfollows:theOsakaMuseumofNaturalHistoryasOMNH,
theNationalScienceMuseum(Nat.Hist).TokyoasNSMT,andtheauthor'scollection
630 NoboruITO
asNIc.Concerningmeasurementofbodyparts,seetheauthor'sformerpaper
nric1lotic11mls(Trichotichnus)anlagisa'ulsN.ITO,sp nov.
(Figs.1&5)
Bodystout,oblong,black,shiny,weaklyiridescentonelytra;maxillaryandlabial
palpi,antennaeandtarsilightreddishbrown, legsbrown,labrumandmandiblesdark
brown
Headlarge,0.70 timesaswideasthepronota1width, fairlyconvexonvertex,
partlywithsparseandvaguepunctures; labrumtransverse,withratherwellconcave
apex;clypeusalmostsmooth,shallowlyemarginateat apex;clypealsutureshallow,
clearlycarved; frontalimpressionslinear,moderatelyengravedinapicalhalves,thence
shallowedbehind,reachingsupraorbitalgrooves;interocularspace0.68timesaswide
aswidthofheadincludingeyes;eyesratherwell prominent, thoughsmall; temples
long,two-fifthstheeyelength;spacebetweengenuineventralmarginsofeyesand
buccalfissurewide;labialpalpislender,3rdsegmentweaklydilated,approximatelyas
longasthe2nd;ligulawedge-shaped;parag1ossaenarrow,adnatewithligulatobehind
itsapex;mentumwithepiIobesweaklywidenedapicad,bearingmediantoothrounded
atapex;microsculptureweaklyvisible,composedofisodiametricmeshesonclypeus
andofobscuretransverseonespartlyontheotherportions.
Pronotumcordate,widestatapicaltwo-fifths,approximatelytwo-fifthswiderthan
long,wellreflectedatsides,ratherstronglyelevatedondisc,theelevationdivideddue
todeepmedianline;sides wellarcuateapicad,andlinearly convergent basad from
middle,ratherdeeplysinuatebeforebase;apexshallowlyemarginate,unborderedme-
dially; basesublinear,withborderthininmiddle; lateral furrowsnarrow, gradually
widenedbackwards, fallenintobasal foveae;basal foveaeeachdeeplyandlongitudi-
nally groovedat innersidesandhumpedoutsidethegrooves; both front andhind
transverseimpressions deep;surfacewhollycoveredwithpunctures,whicharefine
andsparseondisc,rathercoarseandmoderateinapicalandlateralareas,andstrongly
coarseandpartlyconfluent inbasal foveae;microsculptureveryobscure,partly ob-
servedas transversemeshes.
Elytraoblong-oval,widestalittlebehindmiddle,1.30timesaswideastheprone-
tatwidth, ahalf longer thanwide,gentlyconvex, without punctures,sidesweakly
curvedinhumeri, veryshallowlysinuatepreapica11y;apiceseachproducedbehind,
narrowlyroundedatdistalmargin;basesmoreorlessemarginate,withsmall toothat
eachend;striaedeep,narrow,finelyandclearlycrenulate,scutellarstrictomoderatein
length; intervalsraisedevenondisc,morestronglysobasadandapica 3rd interval
withsetiferousporebetweenmiddleandapicaltwo-fifths;marginalseriescontinuous,
Figs. 1 Habitusof「,-lc/folic/1Mlsspp 1,「1・lc/toric1/川Is(「1・lc/forfeit,7,Is)e,n1ag1sa,IMsN. ITO,sp.
.
nov;2, T.(T)1/aktlsaml.,,N. ITO,sp 3, T. (T) ,11asatakal・os/tidalN. ITO, sp nov:4, T. (T)
、'os1/1、,tlkiiN.ITO,sp nov.
NewSpeciesoftheTlrichotichntlsleptopusGroup 631
l 2
4
3
632 NoboruITO
composedof25-27umbilicatepores;microsculpturecomposedofveryfinetransverse
lines.Hindwingsvestigial.
Ventralsurfacemoreorlesscoarselyandmoderatelypunctateonpro-,meso-and
metepisterna,and lateralareasofmetasternum;metepisternumone-fifthwider than
long;apicalmarginof6thabdominalsternitetruncateattipandunisetoseateachside
in andwidelyroundedatthetipandbisetoseatthesidein
.
Legslong; foretibiaeclearlysulcate;1stsegmentofmidtarsusin (5bearingad-
hesivehairsventrallyinapicalhalf,hindtarsusone-seventhin andone-eleventhin
longerthanthewidthofhead,1stsegmentalittlelongerthanthree-fourthsaslongas
the2ndand3rdtakentogetherandtwicethe2ncし3rdahalflongerthanthe4th,claw
segmentquadrisetosealongeachventralmargin.
Aedeagus (Fig 5) robust, weakly arcuate preapica11y and thence gradually
thinneddista(i, feeblydirectedobliquelyventradattip;apicalorificewide,innersac
armedwithrather longpeg-shapedsclerite;apical loberoundedandobscurely bor-
deredatdistalmargin;ventralsurfaceshallowlyconcave,thinlyborderedatsides.
Length:12.1-12.8mm.Width:4.9-5.2mm.
Holotype: Amagi Pass,Shizuoka,S. KAsAHARA leg. (preserved inNSMT).
,
Paratypes:2(3(3,samedataastheholotype;1?,Mt.Amagi-san,Shizuoka,8-V-1993,
S. MORITA leg; 1 1 Kawazu-cho, alt 400m,Shizuoka, 22-IX-2002, H.WATA-
, ,
NABEle9.
ema1fo'. Thisnewspeciesisallied to 'lc/1oflc/1mls(「1,lc/1oflc/1,7tfs) /lf,-a川siltr
MORITA,but isdistinguished fromthelatterbythepronotuminbasal foveaedeeper
andwithclearhumps.
Etymology. Thespeciesisnamedafterthetypelocality,Mt.Amagi-san
nri'c1lotichnus(Tric1lotic/1nus)/takusa,MsN.ITO,spnov
(Figs 2&6)
Body rathernarrow,oblong,moderatelyconvex,slightlybrownishblack,shiny,
with very slightly iridescent lustreonelytra; labial andmaxillary palpi yellowish
brown,antennaeandlegslightbrown,labrumandmandiblesdarkreddishbrown.
Headwide,0.73-0.56timesaswideasthepronota1width,notpunctate;labrum
mediuminshape;clypeuswithseveralobscurerugositiesnearsetiferousporeateach
side, subtrapezoidally emarginateat apex; clypea1suture thin, clear and straight;
f「ental impressionsweaklyarcuatelydivergentbehin(i,moderateindepth,obljterated
nearSupraorbitalgrooves;interocularspacewide,three-fourthsaswideasthewjdthof
head;eyesalittlemoreprominent thaninusualspeciesofthegroup;genujneventral
marginofeyeswidelyisolatedfrombuccal fissure;labialpalpislender;mentumwjth
epi1obeSSlightlywidened forwards,mediantoothofmentumnarrow;mjcrosculpture
almostinvisibleonfrons,detectedasisodiametricmeshesinapicalhalfofclypeus.
P「onOtumcordate,notsowide, 1.30-1.39timesaswideaslong,wellre?ectedat
Sides,fairlyconvex;sidesmorestronglyarcuateforwardsthanbackwards,deeplysjnu_
NewSpeciesoftheTrichotlc1/ntls. lptop11sGroup
633
Figs.5-6. MalegenitaliaofTli・ic1/ottc1/1,usspp. - 5,T.(T)amaglsanusN. ITO,sp nov;6,T.(T)
hakusanusN.ITO,spnov;d,dorsalaspect.Scale:1mm
atebeforebase;apexratherdeeplyemarginate,borderedthroughout;baseone-seventh
wider thanapex,bisinuatelyemarginate, thickly bordered lengthwise; apical angles
wellprotruding forwards,narrowlyrounded;basalanglessmallerthanrightangle,tri-
angularlyprominent;lateral furrowsnarrowinapical fourth, relativelywideinthere-
634 NoboruITO
mainingportions; basal foveaewide, longitudinallygrooved at inner sides, gently
humpedalongoutersidesofthegrooves; fronttransverseimpressionsratherdeep,the
hindoneshallow;medianlinefine,lyingbetweenboththeimpressions;dorsalpunc-
tureslackingorverysparseandfineondisc,moreorlesscoarseandsparseinthefur-
rowsandcoarseanddenseinthefoveaewherethepuncturesarepartlyconfluent;mi-
crosculpturecentrallyvisibleastransversemeshesandsointhesurroundingareasas
finelyisodiametricmeshes.
Elytraoblong-oval, weakly convex,ahalf longer thanwide, 1.29-1.31 timesas
wideasthepronota1width,withoutpunctures;sidesgently divergent fromapex to
middle, veryshallowlyconcavepreapica11y;apicesnarrowly roundedat tips;bases
eachshallowlyemarginate,withaminutehumeraltooth;striaedeep,wide、andclearly
crenulate,scutellarstrictomoderateinlength;intervalsgentlyconvexondiscandbe-
comingmoreconvexbasadandapicad,asetiferousporeof3rdintervalsituatednear
middle;marginalseriescontinuous,consistingof24-28umbilicatepores;microsculp-
turedetectedasvaguetransverselines.Hindwingsatrophied.
Ventral surfaceobscurelypunctateonprosternumandprepisterna,coarselyand
sparsely so onmetepisterna and laterally onmetasternum; metepisternumone-fifth
wider thanlong;6thabdominalsternitein truncateatapexandunisetoseateach
sideandin widelyroundedattheapexandbisetoseattheside.
Hind femurbisetosealonghindmargin; fore tibiadorsallysulcate, with threeor
fourspinesalongapico-externa1margin;hindtarsus1.08timesin and 1.02times in
as longasthewidthofhead.1stsegmentone-sixthshorter thanthe2ndand3rd
combine 2ndone-thirdlonger thanthe3rdand2.70timesas longasthe4th,claw
segmenttri-orquadrisetosealongeachventralmargin.
Aedeagus(Fig 6)robust,thickinmiddle,thenceabruptlytapereddistadLthinand
weaklydirectedventradatapex,slightlythickenedat tip;dorsalsurfacefeeblycurved
totherightnearapex;apicalorificewidelyopen,innersacarmedwithspinemoreor
lesslong,peg-shapedandweaklycurved;apical lobesubquadrate,widelyarcuateand
borderedatdistalmargin.
Length: ll.4-12.3mm.Width:4.1-4.9mm.
Holotype: Shaka-rindou, Mt.Haku-san, Ishikawa, 2-VIII-1997, M. MATsul
,
leg.(preservedinOMNH).Paratypes:1!,samelocalityastheholotype,22~27_viii_
2002,H.HOSHINAleg;1 Nakahanba,Mt.Haku-san,Ishikawa;1 3 Ikegahara,
Fukui,13-VII-1986,M.SA,IToleg;1d,Mt.Haku-san,Fukui,26_vi,l_1g6g, TMlzu_
NUMAleg.
e'71a''、/'S・ ThisnewSpecies isalliedto ,lc/7ofoc/mls(n,!c/7of1c/mls) late,77a,._
91natiSl N・ITO、buttheP「onOtumismorestronglyreflectedatthesjdesandnotormore
Spa「SelyPunctateonthedisc,andtheelytraarealittlemorewjdenedbackwards
Etymo1o9y・TheSpecificnameisderivedfromthetype1ocaljtyoftheholotype
andsomeparatypes,Mt.Haku-san.
NC、vSpeciesOfthe「''lc1/011・t1/川1、./e1リ10/)ll、.Group 635
nfC/lOfic/m‘s(nfc/1of''c/lM's)'nasafa ayes/Ii'dal'N.ITO,sp
nov
(Figs 3&7)
Bodyoblong,darkreddishbrowntoblack,withweaklyiridescentlustreonelytra;
labialandmaxillarypalpi,andantennaeyellowishbrown, tarsiandlegsexcept for
da「k brown portions ofjoints light reddishbrown, labrum and basal portjons of
mandiblesdarkbrown.
Head far9e,0.76 timesaswideasthepronota1width, fairlyconvexonvertex,
Withinterocularspaceratherwideand0.67-0.72timesaswideasthewjdthofhead;
labrumsubtrapezoida1,ratherdeeplyandtriangularlyconcave;clypeusvaguelyrugose,
Weakly protrudent at apical angles; cIypeal suture fine,shallow tosomewhat deep;
f「ental impressions moderate in depth, attaining to supraorbital grooves, though
St「on9lyShallowednearthegrooves;eyesnotprominent; templesgentlyconvergent
behind,two-fifthstheeyelength;spacebetweengenuineventralmarginsofeyesand
buccalfissuresimilarinwidthtothatofthepreviousnewspecies1antennaelong,sIen_
de「,andreachingbasal fifthofelytra;labialpalpislim;mediantoothofmentumwjde,
a「Cuateatapex;microsculptureobscurelyvisible,composedofisodiametricmesheson
clypeusandofsubsquaremesheshereandthere.
P「onotum subcordate,widest nearapical two-fifths,approximately14 times as
Wideaslong,rathersteeplydeclivousapico-laterad;sidesgentlyarcuatelyconvergent
forwardsandweaklysinuatelysobackwardsfromthemiddle;apexmoderatelyemar_
ginate,thinlybordered,theborderinterrLlptedmedially;basealmoststraightorbarely
emarginate, thinlyandentirelybordered;basalanglesrectangular,withsmall teethat
eachtip; lateral furrowseachmoreorlesswideinapicalhalf,graduallyandweakly
expandedbasadfromthere;basal foveaewide,almostflattened,shallowlygroovedat
innersides;both frontandhindtransverseimpressionsshallow;medianlinefinebut
clear, reducednearapex andbase;surfaceminutelyandsparsely punctateondisc,
somewhatcoarsely anddenselysoinapicalareas,coarselyanddenselysoinlateral
furrowsandbasal foveae, where thepuncturesarepartlyconfluent;microscuIpture
partlyvisibleasveryvagueandqubsquaremeshes.
Elytra oblong, 1.51-1.55 times as longas wide, gently convex; sides gently
rounded in humeri, subarcuate inmiddle, somewhat deeply sinuate before apices,
whichisnarrowlyroundedateachdistal margin; bases eachshallowly emarginate,
roundedathumeralangle;striaerelativelywideandfinelycrenulate;intervalsflattened
ondisc,slightlyconvexapicadandbasad,asetiferousporeof3rdintervalsituatednear
middle; marginal series uninterrupted, consisting of 26-28 umbilicate pores1 mi-
crosculptureinvisible.Hindwingsreduced.
Ventralsurfacevaguelyandsparselypunctateonpro-,meso-,andmetepisterna;
metepisternumnearlyone-tenthwiderthanlong;6thabdominalsternitein truncate
or feeblyemarginateatapexandunisetoseateachside, in widelyroundedatthe
apexandbisetoseattheside.
Legslong; fore tibiaeobscurelysulcate; hindtarsus1.02-1.08 times in (S and
636 NoborulTo
0.97_0.99 times in aslongasthewidthofhead,1stsegmentone-seventhshorter
thanthe2ndand3rdtakentogethernearly twicethe3rdwhichistwo-thirdslOnge「
thanthe4th,eachventralsideofclawsegmentwithfoursetae.
Aedeagus(Fig7)stout,wellthickinmiddle,thencesteeplynarrowedbackwards,
thinnedatapex,slightlyreflectedat tip;apicalorificewide, innersacbearingshort
peg_shapedsclerite;apical lobesubquadrate,withdistalmarginborderedandweakly
rounded.
Length:11.5-12.4mm.Width:4.5-4.9mm.
Holotype: Minokoshi, Mt. Tsurugi-san,Higashi-iyayama-mura,Tokushima,
,
24_VII-2004.N. ITOleg.(preservedinOMNH).Paratypes:2 7 .samedataas
,
theholotype; 11 ,samelocalityandcollectorastheholotype,25-VII-2004; 1
,
samelocalityastheholotype,26-VII-2004,H.NoMuRAleg;6 ,samelocalityas
theholotype, 10~11-VI-2000,N. ITOleg;、(3, 1?,samelocalityastheholotype,
16~23-IX-2005,Y. ITOleg;1(S,samelocalityastheholotype,10-VI-2000,Y. ITO
leg;2(S(S,4 ,same localityas theholotype,5~12-IX-2002, Y. ITOleg; 1(3, 1?,
Va1. 0tsurugi-dani,Mt.Tsurugi-san,Tokushima,14~15-IX-1996, Y. ITO leg; 1
,
Mt.Tsurugi,Awa,29-VII-1960,T.SHIBATAleg;13,ditto,30-VII-1960; ld,19,
Val.Nikubuchi-dani,Kisawa-mura,Tokushima,16~17-VII-1994,Y.ITOleg;19,Mt.
Kotsu-san, alt 950-1,100m, Yamakawa-cho, Tokushima, 14-VII-2001,Y. ITOleg;
3 Mt.Kumosou-yama,Kamiyama-cho,Tokushima,17~18-VII-1988,Y.ITOleg;
,
1 2 ditto,2~3-VIII-1997,M.YosHIDAleg.
, ,
emc'lf,s. Thisnewspeciesissimilar tO「rio/70「lc/7'71's(「'''c/lotic/7mfs)tfeno''im
KAsAHARA et Y. ITO but is discriminated from the latter by thepronotum sinuately
convergentbasadinsteadofbeinglinearlysoandnarrowerinlateral furrows.
ThisnewspeciesiscompletelysympatricwithT. (T)sill/,,01、、uensls.As farasI
knew infieldworks, thisspecies is foundinthesame valleynearMinokoshiofMt.
Tsurugi.EcologyofthesespeciesonMt.Kumosou-yamaandMt.Kotsu-sanmaybe
thesame.Fromthisphenomenon,itcanbeestimatedthatthetwospeciesarewidely
isolated fromeachother inphylogeneticandgeneticaspects,inspiteofbeingsimilar
inapparentexternalcharacteristics.
Etyn1o1ogy. ThespeciesisnamedafterMr.MasatakaYosHIDA
nfC/lOflC/1M‘s(nfc/loflc/1Mls)yes/Ii;vMl'l'N.ITO,spnov.
(Figs 4&8)
This new species iscloselyallied to thepreviousnewspeciesT. (T')masa_
「aA'f0'oS/1fC/a!,but thepronotumismoretransverse a littlemorearcuate behjnd from
theWidestPoint,notsinuatebeforebase, lesscoarselypunctateinbasal foveae,and
WithSmalIe「toothatthetipofeachbasalangle,theelytraareshorterandtheother
Characteristicsaredifferentasmentionedinthefo11owjngdescrjptjon
BodyWidelyoblong,black,shiny,withiridescentlustreonelytra:maxj11aryand
labialPalPi li9htbrown,antennae,tarsiandtibiaelightreddishbrown, femoradark
NewSpeciesofthe'「,,-io/1()tichtltlslep1()ptlsGroup 637
Figs 7_8 MalegenjtaljaOf r,・1'c/7o1ic加Illsspp. - 7, 「.(「.)'nasa「a人C'り'oS加(fatN・ITO'SP nov;8' 「
(r)los110,t,/a・・,NITO,spnov;d,dorsalaspect.Scale:1mm・
638 NoboruITO
reddishbrown, labrum,mandiblesandlateralmarginsofpronotumslightlybrownish
black.Headlarge,0.78timesaswideasthepronota1width、almostsmooth.withinter-
ocularspacemorethantwo-thirdsofthewidthofhead;clypea1suturefineandclear;
frontalimpressionsmoderateindepth,weaklyarcuatelydivergent;eyesrelativelylarge
andmoreor lessprominent;microsculpturemostlyinvisible,observableasisodiamet-
ricmeshes inapical halfofclypeus.Pronotumsubquadrate,1.45timesaswideas
long,weaklyelevated;base1.09timesaswideasapex,clearlyborderedlengthwise;
lateral furrowsnarrowinapicalhalf,thenceweaklywidenedbehind;basal foveaeeach
wide,alittlemoredistinctlyridgedinoutersideofinnergroovethaninTnlasataka-
、・os111da1;dorsalpunctureslackingondisc,minuteandsparsenearapex.sparselyand
moderatein lateral furrows, andsomewhatcoarse inbasal foveae: microsculpture
partlyandvaguelyvisibleastransversemeshesinmostareasandasfneisodiametric
meshesinbasal foveae.Elytraovate,1.44-1.48timesas longaswide, flattenedon
disc,impunctate;preapica1sinusveryshallow;apicesnotproduce widelyarcuateat
tips; intervalsweaklyconvexondisc,graduallyraisedapicadandbasad;marginalse-
riesof26-27umbilicatepores;surfacevaguelyandfinelymicrolined.Hindwingsre-
duced.Metepisternumone-ninthwiderthanlong.Foretibiaesulcatedorsally:hindtar-
sus 1.07 times in and 1.02 times in aslongasthewidthofhead.1stsegment
three-fourthsas longasthe2ndand3rdtakentogetherandtwicethe3rcLwhichis
three-fifthslonger thanthe4th,clawsegmentquadrisetose(rarelytrisetose)ventrally
alongeachmargin. Aedeagus (Fig 8) thick near basal bulb, gradually thinned for-
wards,alittlethickeratapicalportionthaninTmasalakay(:),,/tidal,thickenedat tip;
dorsal surfaceslightly asymmetrical;apical orificewide, innersacarmedwithpeg-
shapedscleriteshorter andslenderer than that of T masatakayos111da1;apical lobe
widelytrapezoidal.
Length:11.7-12.3mm.Width:4.8-5.2mm.
Holotype: . Mt.Mitsuji-yama,alt 950-970m,Tosa-cho,Kochi,Japan,5~8-
VIII-1995,Y. ITOleg. (preservedinOMNH).Paratypes:1 3 ,samedataas the
,
holotype;1e,samelocalityandcollectorastheholotype,27-VIII-2005; l(5、,Mt.Ina-
mura-yama, Tosa-cho,Kochi,Japan,29-VIII~1-IX-1984,Y. ITO leg. (preservedjn
NIc).
Etymology. Thespecificnameisdedicated toMr.Yoshiyuki ITO, Kochj,who
collectedthespecimensofthetypeseries.
Tlric1lotic/lnus(Tlric/1otichmls)isa,nuta,lakaiN.ITo
「''IC/lotic/1'filsIsa'ntitanakal N. ITO,1996.Ent.Rev.Japan,51:127(typelocality:Kibunc.Kyoto)
MORITA.1998.Elytra.Tokyo,25:579.
Speel7'7e'7Se、a7'1'neCf. 3 ,Mt. Hiei-zan,Kyoto, 20-X-1965,K. lsHlzUKAleg;
1(5,Hatcho,PassDanno-toge,alt750m,Keihoku-cho,Kyoto,Japan,12_vI_1985,s
KASAHARAle9.;1e,19,Ashiu,10~11-VI-1999,Kyoto,Japan,N. IT01eg
Remalk. Thisspeciesiswidelydistributed fromtheeasternareaofHyogoto