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Flora Europaea check-list and chromosome index Flora Europaea check-list and chromosome index D.M. MOORE CAMBRIDGE UNIVERSITY PRESS Cambridge London New York New Rochelle Melbourne Sydney CAMBRIDGE UNIVERSITY PRESS Cambridge, New York, Melbourne, Madrid, Cape Town, Singapore, Sao Paulo, Delhi Cambridge University Press The Edinburgh Building, Cambridge CB2 8RU, UK Published in the United States of America by Cambridge University Press, New York www.cambridge.org Information on this title: www.cambridge.org/9780521105736 © Cambridge University Press 1982 This publication is in copyright. Subject to statutory exception and to the provisions of relevant collective licensing agreements, no reproduction of any part may take place without the written permission of Cambridge University Press. First published 1982 This digitally printed version 2009 A catalogue record for this publication is available from the British Library Library of Congress Catalogue Card Number: 82-1335 ISBN 978-0-521-23759-8 hardback ISBN 978-0-521-10573-6 paperback CONTENTS Introduction i-iv Check-list and Chromosome Index 1-365 References 366-408 Index to Families and Genera 409-423 Introduction This book has grown out of the requests by many users of Flora Europaea (Tutin et al., 1964, 1968, 1972, 1976, 1980) for information on the sources of the chromosome numbers cited in that work. During discussions about the publication of these data it became apparent that there is also a demand for a basic checklist of the taxa recognised in the Flora, and that these two requirements could readily be met in the same volume. The Checklist Apart from the fundamental value of Flora Europaea to all modern taxon- omic studies of European plants, the names and sequence of taxa it presents are widely used for arranging herbaria and libraries concerned with these plants. It is intended that the handy format of this checklist will facilitate these latter activities, as well as being useful in recording and assessing local floras within Europe. The sequence of taxa is exactly the same as that used in Flora Europaea except that the Rubiaceae, which for technical reasons had to be published in volume 4 rather than in volume 3, have been positioned correct- ly in the Checklist. All species and subspecies 'accepted' in Flora Europaea (i.e. those prefixed by a number or a letter) are included here. Those unnumbered taxa of doubtful status are only included if a chromosome number is given for them in Flora Europaea, as is also the case for the agamospecies ('microspecies') listed, but not described, within the 1species-groups' of apomictic or partially apomictic genera. It has not been possible to include any synonyms in the Checklist but access to these is provided by the Consolidated Index of Flora Europaea (Halliday and Beadle, 1982) as well as the volumes of the Flora itself. The Chromosome Index As noted in the Introductions to the five volumes of Flora Europaea, diploid (2n) chromosome numbers were given when they were derived from vii material of known wild origin. In the case of naturalized or cultivated taxa chromosome numbers were included when they were known to be based on material with a status which justified their inclusion in the Flora: aliens had to be common and to have been naturalized for at least 25 years, while crops had to be widely cultivated. As can be seen from the Index, between 98% and 99% of the chromosome numbers included in Flora Europaea fulfil these criteria, whether they were based on a single observation from one locality or on many counts from a number of areas. In many instances the Editors were provided with unpublished chromosome data; some of these were subsequently published, hence references to them often postdate the publication of the relevant volume of the Flora, others are apparently still unpublished. The provenance of the material is indicated by using the abbreviations for the territories recognised in Flora Europaea (see p. iii) , except where the data only permit a wider area to be cited. In several cases the chromosome numbers were published under names different from those adopted in the Flora Europaea, but these can readily be traced by reference to the synonymy given in the Flora. Ideally, of course, a definitive chromosome index would give due weight to the number of observations on which each number is based, but the practical- ities of Flora-writing preclude this. The Index, consequently, cites only one reference for each chromosome number given, even though in many instances far more data were available to the editors of Flora Europaea; this has been done to obviate any impression of completeness in the coverage, which would be illusory. Less than 2% of the chromosome numbers cited in the Flora cannot be substantiated or are shown to be derived from material which does not fulfil the criteria referred to above. The commonest reason for this is that pre- liminary accounts of genera included chromosome numbers taken uncritically from the various worldwide compilations and their credentials were mis- interpreted or inadequately checked during the various stages of the editorial process; in other cases, far fewer, the sources cannot now be traced in the records of the Editorial Committee; occasionally, the numbers included in the Flora are simple misprints, undetected during proof reading. The Chromosome Index reveals, therefore, the problems of fully documenting multi-author Floras of substantial areas produced within a realistic, or realisable, time-scale. It is felt that the level of accuracy achieved fully justifies the approach adopted by the editors of Flora Europaea and the publication of this documented list of chromosome numbers. The extra effort required to achieve complete accuracy (if such were possible) would not have been worthwhile. Abbreviations of geographical territories (For precise definitions of these territories, see maps in Flora Europaea) Al Albania Hs Spain Amer.* America Hu Hungary Au Austria Is Iceland Az Acores (Azores It Italy Be Belgium and Luxembc Ju Jugoslavia Bl Islas Baleares Lu Portugal (Balearic Islands) No Norway Br Britain Po Poland Bu Bulgaria Rm Romania Co Corse (Corsica) Rs U.S.S.R. (European part), Cr Kriti (Crete) subdivided thus: Cz Czechoslovakia (N) Northern region Da Denmark (B) Baltic region Fa Faerber (Faeroes) (C) Central region Fe Finland (W) South-western region Ga France (K) Krym (Crimea) Ge Germany (E) South-eastern region Gr Greece Sa Sardegna (Sardinia) Hb Ireland Sb Svalbard (Spitsbergen) He Switzerland Scand.* Scandinavia Ho Netherlands Si Sicilia (Sicily) Su Sweden Tu Turkey (European part) *not used in volumes of Flora Europaea Acknowledgements This book is founded on the endeavours of the Editorial Committee of Flora Europaea (T.G. Tutin, V.H. Heywood, N.A. Burges, D.M. Moore, D.H. Valentine, S.M. Walters and D.A. Webb), to which I am the most recent recruit. Professor D.H. Valentine and later Dr. S.M. Walters were primarily responsible for the chromosome numbers cited in volume 1 of Flora Europaea, and Dr. Walters for a significant part of those included in the second volume. They, and my other colleagues on the Editorial Committee, have been a constant source of support and information. Dr. P.W. Ball, Mr. A.O. Chater, Dr. R.A. de Fillips and Dr. I.B.K. Richardson, Research Associates during the various phases of the Flora Europaea project, checked hundreds of chromosome references, while Dr. Askell LOve freely gave of his wide cytotaxonomical knowledge. Above all, however, the Chromosome Index is a testament to the many authors of Flora Europaea who provided accurate documentation of their chromosomal data; the cooperation of them and other friends and colleagues who made available unpublished information is greatly appreciated. The onerous task of typing the camera-ready copy was undertaken by Miss Valerie Norris, whose efforts are grate- fully acknowledged, whilst I am also indebted to the Flora Europaea Trust Fund of the Linnean Society of London for a grant towards the costs of preparing the manuscript. I LYCOPODIACEAE 1 Huperzia Bernh. 1 selago (L.) Bernh. ex Schrank Mart. 90 Fe Sorsa, 1963 264 Br Manton, 1950 c.272 Is Lttve $ Lttve, 1961a (a) selago (b) arctica (Grossh.) A. § D. Love (c) dentata (Herter) Valentine 2 Lepidotis Beauv. 1 cernua (L.) Beauv. 2 inundata (L.) Bttrner 156 Br Fe V. Sorsa, 1958 3 Lycopodium L, 1 annotinum L. 68 Br He Su Manton, 1950 2 dubium Zoega 68 Is Lttve S Love, 1958 3 clavatum L. 68 Br He Manton, 1950 4 Diphasium C. Presl. 1 complanatum (L.) Rothm. 46 Scand. Lttve $ Lttve, 1961a 2 tristachyum (Pursh) Rothm. 46 Scand. Lttve $ Lttve, 1961a 3 issleri (Rouy) J. Holub 4 alpinum (L.) Rothm. 46 Su Wilce, 1961 48-50 Cz Manton, 1950 II SELAGINELLACEAE 1 Selaginella Beauv. 1 selaginoides (L.) Link 18 Br Manton, 1950 2 helvetica (L.) Spring 18 He Manton, 1950 3 denticulata (L.) Link 18 It Manton, 1950 4 apoda (L.) Fernald 5 kraussiana (G. Kunze) A. Braun III ISOETACEAE 1 Isoetes L. 1 lacustris L. c.110 Is Lbve $ Lttve, 1961d 2 setacea Lam. 22 Is Lttve § Lttve, 1961d 100 Hb Manton, 1950 3 brochonii Moteley 4 azorica Durieu ex Milde 5 boryana Durieu 6 delilei Rothm. 7 malinverniana Cesati § De Not. 8 heldreichii Wettst. 9 velata A. Braun 10 tenuissima Boreau 11 histrix Bory 20 Br Manton, 1950 12 durieui Bory IV EQUISETACEAE 1 Equisetum L. 1 hyemale L. 216 Is Lttve 5 Love, 1961c 2 ramosissimum Desf. c.216 It Manton, 1950 3 variegatum Schleicher ex Weber Mohr 216 Is Lttve $ LOve, 1961c 1