©EntomologicaFennica.3March2006 Finnish flies of the family Therevidae AnttiHaarto&KajWinqvist Haarto,A.&Winqvist,K.2006:FinnishfliesofthefamilyTherevidae.—Ento- mol.Fennica17:46–55. Anillustratedkeyforthe17FinnishspeciesofTherevidaeispresentedthatin- cludessixspeciesrecordedfromneighbouringareas.DistributionsoftheFinnish species are discussed and presented in maps; evaluations concerning the threatenednessstatusofFinnishspeciesareincluded. Antti Haarto, Zoological Museum, Section of Biodiversity and Environmental Science, University of Turku, FI-20014 Turku, Finland; E-mail: antti.haarto @turkuamk.fi Kaj Winqvist, Zoological Museum, University of Turku, FI-20014 Turku, Fin- land;E-mail:[email protected] Received31January2005,accepted13July2005 1.Introduction book on lower Brachycera (Stubbs & Drake 2001)lacksmanyFinnishspecies.Themostac- FliesofthefamilyTherevidaebelongtothesub- curate key for determining Finnish Therevidae orderBrachyceraandarerelativelylargeDiptera. was presented by Lyneborg and Spitzer (1974); UnlikeAsilidae,amorphologicallyandbiologi- thiskeycoverstheCzechRepublicfauna. cally close family, adults of Therevidae are not Thepurposeofthisarticleistoprovideanup- predatoryandlackthedeepgroovebetweenthe dated key for the Finnish Therevidae and some eyescharacteristicofAsilidae. speciesfoundintheneighbouringcountries.The InFinland,therehasbeenverylittleinforma- distributionandhabitatassociationsforeachspe- tionpublishedonTherevidae.Frey(1911)listed ciesarebrieflydiscussedbasedonthefewavail- some of the oldest records for the lower abledistributionalandecologicaldata.Aprelimi- Brachyceranflies.Intheirexchange-scorecata- nary understanding of the Finnish Therevidae logue,Frey etal.(1941)recorded13speciesof faunawasobtainedduringtheautumn2003,the TherevidaeasbeingFinnishbutusingnamesthat Finnish museum material of Therevidae was arecurrentlyinvalid.Hackman’s(1980)checklist checkedforaBrachycerachecklist(Kahanpää& included20speciesofTherevidae. Winqvist 2005). Three species listed by Hack- Aftertheturnofthenewmillennium,anew man(1980)appearednottobelongtotheFinnish waveofdipterologicalinteresthasbeguninFin- fauna: both European Cliorismia species had land.Aslargeandconspicuousflies,Therevidae been recorded In Russian Karelia, and Thereva canbeconsideredasuitablegroupforresearch, circumscripta (Loew) is a more southern Euro- buttherehasbeenalackofupdatedtaxonomicin- peanspecies.Nowthereare17speciesofThere- formation.Forexample,theTherevidaekey for vidaeknownfromFinland. theEuropeanpartofRussia(Zaitsev1988)leads Thereremainsalottobediscoveredaboutthis to unreliable or inadequate identifications of elusiveflyfamily.Newstudiesareneededtoob- FinnishTherevidae.Furthermore,arecentBritish tainamoreprofoundunderstandingofthedistri- ENTOMOL.FENNICAVol.17 (cid:127) FinnishTherevidae 47 Fig.2.Themiddle Fig.1.(A)TheantennaeofDialineuraanilisand(B)Acrosathe femorainanteriorview annulata. ofAcrosatheannulata male. Fig.3.Theheadof(A)Dichoglena nigripennisand(B)Psilocephalaimberbis males. Fig.4.Thetipofaedeagusinlat- eralview.(A)Cliorismiaardeaand (B)Cliorismiarusticamales. butions, ecological associations, and conserva- Female:Fronswithacontinuousordivided tionstatusofTherevidaespeciesinFinland. shinyblackcallus(frontalcallus) Thereva 4a Knobofhaltereslight 5 4b Knobofhalteresdark Spiriverpalunulata 2.KeytotheFinnishspecies 5a Male: Middle femora with usually 2 ofTherevidae anteroventralbristles(Fig.2) Female: Erect hairs on tergites 5–8 black The following key covers all the 17 Finnish Acrosatheannulata Therevidae species and six additional species 5b Male: Middle femora usually without (listedwithinbrackets)thathavebeenfoundfrom anteroventralbristles Scandinavia,RussiaorBalticcountries.Thiskey Female: Erect hairs on tergites 5–8 white isbasedonourstudiesandliterature(Lyneborg& [Acrosathebaltica] Spitzer 1974, Stubbs & Drake 2001, Zaitsev 6a Males 7 1988). 6b Females 11 7a Frons broader than anterior ocellus (Fig. 1a First segment of antennae thicker than the 3A). Abdomen silvery pollinose or black pedicelandflagellum(Fig.1A) withsilverypattern 8 Dialineuraanilis 7b Frons narrower than anterior ocellus (Fig. 1b Allsegmentsofantennaeequallythick(Fig. 3B).Abdomensilverypollinose 9 1B) 2 8a Halteres black. Abdomen silvery pollinose 2a Facehairy 3 Dichoglenanigripennis 2b Facebare 6 8b Halteres yellow. Abdomen black, tergites 3a Male:Tergitessilverypollinoseandthehairs withweaksilverytriangles onabdomensilverywhite Pandiviriliaeximia Female:Fronswithoutcallus 4 9a Femoracompletelyblackfrombasetoapex 3b Male:Tergiteswithoutpollentofullydusted Psilocephalaimberbis with dirty yellow or grey pollen (usually 9b Femorawithyellowapex 10 withadarkanteriorandlighterposteriorre- 10a Tip of aedeagus straight (Fig. 4A) gion),hairsonabdomenmixeddark(black [Cliorismiaardea] to brownish black) and light (yellow to 10b Tip of aedeagus S-shaped (Fig. 4B) white) [Cliorismiarustica] 48 Haarto&Winqvist (cid:127) ENTOMOL.FENNICAVol.17 Fig.6.Thehindfemora inposteriorviewof Therevainornatamale. Fig.5.Thetipofaedeagusinlateralview.(A)Therevacinifera,(B)T. fulva,(C)T.inornata,(D)T.plebejaand(E)T.strigatamales. Fig.8.Thehindfemorainanteriorviewof(A)Thereva inornataand(B)T.nobilitatamales. Fig.7.Themesonotumof Thereva. 11a Femoracompletelyblackfrombasetoapex 5b The basal part of hind femora without 12 posteroventralbristles 10 11b Femorawithyellowapex 6a 2pairsofdorsocentralbristles(Fig.6).The [C.ardeaandC.rustica] basal part of hind femora with 3–4 12a Tibiae and tarsi as dark as femora. Wing posteroventralbristles(Fig.7).Theantero- membranewithbrownishborderadjacentto ventral bristles of the hind femora usually wingveins Dichoglenanigripennis uniserial (Fig. 8A). Tip of aedeagus long 12b Tibiaeandtarsiyellowish,lighterincontrast curved(Fig.5C) T.inornata tofemora.Wingmembranewithoutbrown- 6b 1pairofdorsocentralbristles.Thebasalpart ishborderadjacenttowingveins 13 ofhindfemorawith1–3posteroventralbris- 13a Wingmembraneyellowbasally.Largespe- tles. Hind femora with additional bristles cies(12–18mm) Pandiviriliaeximia outsidethenormalanteroventralrow(asin 13b Wing membrane completely transparent. Fig.8B) 7 Smallspecies(8–10mm) 7a Anterior part of tergites 4–5 with narrow Psilocephalaimberbis darkband(morethanhalfthelateralpartof tergites4–5dustedwithgreypollen) 8 GenusThereva 7b Anteriorpartoftergites4–5withbroaddark band(onlyhalfthelateralpartoftergites4–5 1a Males 2 dustedwithgreypollen) 9 1b Females 14 8a Tergite 6 fully dusted with greyish pollen 2a All tergites light anteriorly, tergites com- (darkanteriortransversebandabsent).Wing pletelydustedwithdirtyyelloworgreypol- membrane greyish, veins light brown and len 3 crossveins weakly darkened. Hairs on the 2b Anteriorregionofatleasttergites2–5dark 4 lower part of the occiput and parafacial 3a Knob of halteres brownish black. Tip of mixedwithwhiteandblack T.unica aedeagusshort(Fig.5A) T.cinifera 8b Tergite 6 incompletely dusted with greyish 3b Knob of halteres yellowish brown. Tip of pollen (dark anterior transverse band pres- aedeaguslong(Fig.5B) [T.fulva] ent). Wing membrane whitish, veins dark 4a Alltergitesglossyblack [T.atripes] brownandcrossveinsintensivelydarkened. 4b Atleastpartofthetergitesmatt 5 Hairs on the lower part of the occiput and 5a At least one posteroventral bristle in the parafacialwhite [T.marginula] basalpartofthehindfemora(Fig.6) 6 9a Mesonotumgreyishbrownandusuallywith ENTOMOL.FENNICAVol.17 (cid:127) FinnishTherevidae 49 acleardarkbrowncentralline.Hairsonter- rowtransversebandordividedintotwosep- gite1completelyblackordarkbrown.Tipof aratecalli 15 aedeagusshort(Fig.5D) T.plebeja 14b Onewide,usuallylarge,shinyblackfrontal 9b Mesonotumbluishgray andatmostwitha callus 16 diffuse dark central line. Hairs on tergite 1 15a Frontalcallusanarrowtransversebandthat greyoryellowishmedially.Tipofaedeagus is sometimes divided medially (Fig. 9A). long(Fig.5E) T.strigata Anteroventral bristles of the hind femora 10a Hairs on scutellum sparse and completely uniserial(asinFig.8A) T.valida black. Hairs on abdomen predominantly 15b Frontalcallusclearlyseparatedontwoparts black. Usually 1 dorsocentral bristle. Front (Fig.9B).Anteroventralbristlesofthehind femoraanteroventrallyusuallywithoutbris- femora nearly uniserial, but irregularly ar- tles T.valida rangedtowardsbase(asinFig.8B) 10b Hairsonscutellummixedyellowandblack. T.unica Hairsonabdomenyellowlaterallyandblack 16a All tergites light anteriorly, tergites com- dorsally 11 pletelydustedwithdirtyyelloworgreypol- 10c Hairs on scutellum and abdomen almost len 17 completelywhite 12 16b Anteriorregionofatleasttergites2–5dark 11a Anteroventral bristles of the hind femora 18 multiserial(somebristlesoutsidethenormal 17a Knob ofhalteres blackish. Tergite7 dusted anteroventral row) (Fig. 8B). Hairs on with grey pollen. The frontal callus sepa- scutellum not exceptionally dense and ratedfromtheanteriorocellus(Fig.9C) mixed yellowish and black. Usually 1 T.cinifera dorsocentral bristle. Front femorawith 1–4 17b Knobofhalteresyellowish.Tergite7glossy anteroventralbristles T.nobilitata blackandonlylaterallygreypollinose.The 11b Anteroventral bristles of hind femora frontalcallusextendstotheanteriorocellus uniserial(asinFig.8A).Hairsonscutellum (Fig.9D) [T.fulva] dense and almost completely yellow. 2 18a The frontal callus extends dorsally around dorsocentralbristles.Frontfemorawith2–7 theocellartubercle(Fig.9E) 19 anteroventralbristles T.handlirschi 18b Thefrontalcallusextendsdorsallyatmostto 12a 2–3 pairs of dorsocentral bristles (Fig. 7). theanteriorocellus 21 Wing veins narrowly bordered by dark 19a Tibiaeblack.Wingmembranedark membrane. Front femora without [T.atripes] anteroventral bristles. Mesonotum with in- 19b Tibiaeyellowish.Wingmembranewithdark distinctlongitudinalstripes T.fuscinervis areasonlyadjacenttowingveins,predomi- 12b 1 pair of dorsocentral bristles. Wing veins nantlycleartotranslucentwhitishgrey 20 not bordered by dark membrane. Front 20a 2–3 dorsocentral bristles (Fig. 7). Wing femora with or without anteroventral bris- veinsnarrowlyborderedbydarkmembrane. tles. Mesonotum with distinct or indistinct FronsandfrontalcallusnarrowerthaninT. longitudinalstripes 13 lanata T.fuscinervis 13a Front tibia longer than front femora. Head 20b 1 dorsocentral bristle. Wing veins not bor- (in dorsal view) as broad as mesonotum. deredbydarkmembrane.Fronsandfrontal Front femora with anteroventral bristles. calluswiderthaninT.fuscinervis(Fig.9E) Mesonotum with distinct light longitudinal T.lanata stripes T.microcephala 21a Anteriorpartoftergites2–5withdarkband 13b Fronttibiaaslongasfrontfemora.Head(in extendingmediallytoposterioredge(poste- dorsalview)broaderthanmesonotum.Front rior of tergites 2–4 with broken or nearly femora without anteroventral bristles. brokenbandofgreypollen).Frontalcallusin Mesonotum with indistinct light longitudi- Fig.9F T.microcephala nalstripes T.lanata 21b Anteriorpartoftergites2–5withdarkband 14a Shinyblackfrontalcallusformingonenar- extending medially near to posterior edge 50 Haarto&Winqvist (cid:127) ENTOMOL.FENNICAVol.17 Fig.9.Thefrontalcal- lusof(A)Thereva valida,(B)T.unica,(C) T.cinifera,(D)T.fulva, (E)T.lanata,(F)T. microcephala,(G)T. plebeja,(H)T.strigata, (I)T.nobilitata,(J)T. inornataand(K)T. handlirschifemales. Fig.10.Thetipof ovipositorin posterolateral view(A)Thereva plebejaand(B)T. strigatafemales. (posterioroftergites2–4withcompleteband row)(asinFig.8B).Frontalcallusnarrowly ofgreypollen) 22 separatedfromanteriorocellus.Wingmem- 22a Frontalcallusextendslaterallytoeyemargin branewhitish,veinsdarkbrownandcross- 23 veinsborderedbydarkmembrane.Tergite3 22b Frontalcallusseparatedfromeyemarginby without contrasting line between anterior amattarea 24 darkbrownishpartandposteriorgreyishpart 23a Spines on the tip of ovipositor broad and ofthetergite [T.marginula] blunt(Fig.10A).Mesonotumdark,brown- 24c Knob of halteres blackish. Anteroventral ish grey with distinct light longitudinal bristles of hind femora uniserial (as in Fig. stripes.FrontalcallusinFig.9G T.plebeja 8A). Frontal callus separated from or ex- 23b Spinesonthetipofovipositorslender(Fig. tends to anterior ocellus. Wing membrane 10B). Mesonotum light, bluish grey with greyish brown, veins light brown and distinct light longitudinal stripes. Frontal crossveinsnotborderedbydarkmembrane. callusinFig.9H T.strigata Tergite3withcontrastinglinebetweenante- 24a Knobofhalteresyellowishbrown.Antero- riorbroadblackishpartandposteriorgreyish ventralbristlesofthehindfemoramultiserial partofthetergite 25 (some bristles outside the normal antero- 25a Hairsonscutellummixedblackandyellow- ventral row) (as in Fig. 8B). Frontal callus ish. Frontal callus separated from the ante- separated from anterior ocellus (Fig. 9I). riorocellus(Fig.9J) T.inornata Wingmembranegreyishbrown,veinslight 25b Hairsonscutellumyellow.Frontalcallusex- brownandcrossveinsnotborderedbydark tendstotheanteriorocellus(Fig.9K) membrane.Tergite3withoutcontrastingline T.handlirschi betweenanteriornarrowdarkbrownishpart andposteriorgreyishpartofthetergite 3.Faunisticcommentsand T.nobilitata theevaluationofthreatenedspecies 24b Knob of halteres blackish. Anteroventral bristlesofthehindfemoramultiserial(some The following faunistic comments are based on bristles outside the normal anteroventral checkingtheFinnishmuseumcollections(Finn- ENTOMOL.FENNICAVol.17 (cid:127) FinnishTherevidae 51 Fig.11.TheFinnishbiogeographical Fig.13.ThemapwithFinnishvegeta- provincesandtheircodes.Al=Alandia, tionzones.(1a)Hemiboreal,ÅlandIs- Ab=Regioaboensis,N=Nylandia,Ka lands,(1b)Hemiboreal,Southwestern =Kareliaaustralis,St=Satakunda,Ta coast,(2a)Southernboreal,South- =Tavastiaaustralis,Sa=Savoniaaus- westerndistrict,(2b)Southernboreal, tralis,Kl=Karelialadogensis,Oa= Finnishlakedistrict,(2c)Southernbo- Ostrobottniaaustralis,Tb=Tavastia real,CoastofSatakunda,(2d)Southern borealis,Sb=Savoniaborealis,Kb= boreal,CoastofOstrobothnia,(3a)Mid- Kareliaborealis,Om=Ostrobottniame- dleboreal,Ostrobothnia,(3b)Middle dia,Ok=Ostrobottniakajanensis,Oba boreal,Kainuudistrict,(3c)Middlebo- =Ostrobottniaborealisparsaustralis, real,TriangleofLapland,(4a)Northern Obb=Ostrobottniaborealisparsbore- boreal,Koillismaadistrict,(4b)Northern alis,Ks=Regiokuusamoensis,Lkoc= boreal,SouthernLapland,(4c)North- Lapponiakemensisparsoccidentalis, ernboreal,ForestLaplandand(4d) Lkor=Lapponiakemensispars Northernboreal,MountainLapland. orientalis,Le=Lapponiaenontekiensis, Li=Lapponiainarensis. Table1.Thecategoryintheevaluationofthreatenedspeciesandthecorrespondingnumberofknownlocations, followingRassietal.(2001). Category Meaning Numberofpopulation locations CR Criticallyendangered Extremelyhighriskofextinctioninthewild 1 EN Endangered Veryhighriskofextinctioninthewild 2 VU Vulnerable Highriskofextinctioninthewild 3–5 NT Nearthreatened Closetoqualifyingfororneartobecomevulnerable 6–25 LC Leastconcern Commonandabundantorhassteadypopulations >25 52 Haarto&Winqvist (cid:127) ENTOMOL.FENNICAVol.17 Fig.12.DistributionmapsoftheFinnishspecies(A)A.annulata,(B)D.anilis,(C)D.nigripennis,(D)P.eximia, (E)P.imberbis,(F)S.lunulata,(G)T.cinifera,(H)T.fuscinervis,(I)T.handlirschi,(J)T.inornata,(K)T.lanata,(L) T.microcephala,(M)T.nobilitata,(N)T.plebeja,(O)T.strigata,(P)T.unicaand(Q)T.valida. ish Museum of Natural History in Helsinki species,andmanyotherinsects,iscriterionD,or (FMNH), Zoological Museum of University of “populationrestricted”.CriterionDis,however, Turku (MZT), Natural History Museum of acommoncategorybecauselimitedresourcesare Forssa, Natural History Section of Museum of available in Finland to monitor populations of KuopioandZoologicalMuseumofUniversityof threatenedinsectspecies.Becausecollectingac- Oulu) as well as numerous private collections. tivitiesinFinlandwererelativelyinfrequentuntil Therevidae specimens collected during various theturnofthecentury,lowercategoriesarealso entomologicalprojectshavebeenidentifiedand preferred for species represented by few speci- usedintheseevaluations. mens in collections. Category A, “reduction of After the faunistic text for each species, we populationsize,”isusedforonespecies:Thereva categorizeeachspeciesbasedonevaluationcrite- strigata. The categories in the evaluation of riaforthreatenedspeciesapprovedbytheWorld threatenedspeciesareshowninTable1.Species ConservationUnionin1994(Rassiet.al.2001) distributions (Fig. 12) were estimated using lo- andlistgeographicalprovincesfromwhicheach calityrecordsanddistributionsofFinnishvegeta- specieshasbeenrecorded(Fig.11).Themostfre- tionzones(Fig.13). quently used category for Finnish Therevidae ENTOMOL.FENNICAVol.17 (cid:127) FinnishTherevidae 53 Acrosatheannulata(Fabricius,1805) mostnorthernlocalitybeingKuusamo.Females Aspeciescommonlyfoundondunesandinother aremorecommonlyfoundthanmales.Thisspe- sandybiotopesthroughoutthecountry.Thisspe- ciesisassessedtothecategoryLCanddoesnot ciesisassessedtothecategoryLCanddoesnot needmonitoring.Ab,N,Ka,St,Ta,Sa,Kl,Tb,Sb, needmonitoring.Al,Ab,N,Ka,St,Ta,Sa,Kl,Oa, Ks. Tb,Sb,Kb,Om,Ok,Oba,Obb,Ks,Lkoc,Li. Psilocephalaimberbis(Fallén,1814) [AcrosathebalticaAndersson,1994] Aspecies historically distributed evenly almost Thisrecentlydescribedspeciesisfoundfromthe throughoutthecountry,buttherecordsofwhich BalticIslandsinSweden.ItispossibletofindA. arepredominantlyold.Sincethereareonlythree balticaondunesfromthesouthcoastofFinland. recordsmadeinthelatest30years,Ta:Tammela ThemaleofA.balticacanbereliablydifferenti- (Mattila 1998), Ta: Ylöjärvi (Mattila 2001) and atedfromA.annulataonlybygenitalicfeatures. Ta: Lammi (Ahlroth 2002), the species is as- (Andersson1994). sessed to the category VU and the last known populations and the possible new findings will [Cliorismiaardea(Fabricius,1794)] needmonitoring.Ab,N,St,Ta,Sa,Kl,Oa,Tb,Sb, The only specimens have been recorded from Kb,Om,Ok,Oba,Li. Karelia,Russia.C.ardeaisthusdeletedfromthe Finnishchecklist.ItispossibletofindC.ardea Spiriverpalunulata(Zetterstedt,1838) from sandy riverbanks in southeastern Finland Atleastpredominantlyanorthernspeciesfound neartheRussianborder. in sandy biotopes. The few Finnish museum specimens of this species had been loaned by [Cliorismiarustica(Panzer,1803)] DonaldWebb,whoreportedS.lunulatafromthe The only specimens have been recorded from provinces Ks–Li in a letter. He also reported an Karelia, Russia. C. rustica is thus deleted from olderrecordfromN:Lappohja,whichcouldbea the Finnish checklist. Also this species can be labelling mistake. There are only two new re- foundfromsandyriverbanksinsoutheasternFin- cords of this species, Li: Utsjoki (Haarto 1994) landneartheRussianborder. andLkor:Savukoski(Bagge2003),butthecol- lectingactivityhasbeenlowinnorthernFinland. Dialineuraanilis(Linnaeus,1761) ThisspeciesisassessedtothecategoryVUand Aspeciescommonlyfoundondunesandinother does not need active monitoring. (N,) Ks, Lkor, sandybiotopesthroughoutthecountry.Thisspe- Li. ciesisassessedtothecategoryLCanddoesnot needmonitoring.Al,Ab,N,St,Ta,Sa,Kl,Oa,Tb, [Therevaatripes(Loew,1847)] Sb,Kb,Om,Ok,Obb,Ks,Lkoc,Lkor,Li. Aneasternspeciesdistributedfromthesouthern SiberiatotheSt.Petersburgarea(Zaitsev1988). Dichoglenanigripennis(Ruthe,1831) It might be possible to find T. atripes from the Ararespeciesofwhichonlysevenscatteredre- southeasternFinlandneartheRussianborder. cords from various parts of the country are known.Sincethereisonlyonerecordmadeinthe Therevacinifera(Meigen,1830) recent years, Ok: Sotkamo (Söderman & Lei- Inadditiontotheoldknownpopulation,thereare nonen2003),thespeciesisassessedtothecate- three new populations of this scarce species goryENandtheknownpopulationandthepossi- found,Ab:Dragsfjärd(Clayhillsetal.1996),N: blenewfindingswillneedactivemonitoring.Al, Hanko(Haarto1996)andN:Helsinki(Kahanpää Ab,N,St,Kb,Ok,Oba. 2003).Alltheselocalitiesarefromthesouthwest- ern coast. This species is probably connected Pandiviriliaeximia(Meigen,1820) withtheBalticsandyshoresinFinland.Itisas- Arathercommonspeciesfoundmostlyinmoist sessedtothecategoryVUandneedsmonitoring. meadowsinthesouthernandcentralFinland,the Ab,N,St. 54 Haarto&Winqvist (cid:127) ENTOMOL.FENNICAVol.17 [TherevacircumscriptaLoew,1847] Therevamicrocephala(Loew,1847) The specimens determined as T. circumscripta This species is distributed along the coast. turnedouttobelongtothespeciesT.validaLoew. T.microcephalaisfoundinforestsintheCentral T.circumscriptaisthusdeletedfromtheFinnish Europe(Keil2003)andalsoinFinlandthenew checklist. recordsofthisspecieshavebeenmadeinmoist forests.Despitethefewrecordsmade,thespecies [TherevafulvaMeigen,1804] might thus be less vulnerable than many other TherewasonespecimenmisidentifiedasT.fulva speciesinthegenus.Thisspeciesisassessedto inthecollectionofFMNH.ThisisaCentralEu- the category NTand does not need monitoring. ropeanspeciesdistributednorthtotheSt.Peters- Al,Ab,N,Om. burgarea(Zaitsev1988).Itmightbepossibleto findT.fulvafromthesouthcoastofFinland. Therevanobilitata(Fabricius,1775) Aspecies historically thought to be common in TherevafuscinervisZetterstedt,1838 Finland(Frey1911)butithasoftenbeenmixed Arathercommonlyfoundspeciespredominantly withT.handlirschi(Kroeber).Thereareonlyfour inthenorthernFinland,buttherearetwospeci- certain records from Finland, all from dry or mensfromAbintheMZT,whichwerecollected sandybiotopesnearthecoast:Al:Kökar(Hellén), by Lahtiperä in the 1950’s. These southern re- Ab: Korppoo (Panelius 1961), Ab: Dragsfjärd cordsareprobablylabellingmistakes.Thisspe- (Haarto1994),Om:Lohtaja(Ahola2004).This ciesisassessedtothecategoryLCanddoesnot speciesisassessedtothecategoryVU.Thelast needmonitoring.(Ab,)Obb,Ks,Lkoc,Le,Li. knownpopulationsandthepossiblenewfindings willneedmonitoring.Al,Ab,Om. TherevahandlirschiKroeber,1912 Oneofthecommonestspeciesofthegenusdis- Therevaplebeja(Linnaeus,1758) tributed in the southern Finland. This species is A dark Thereva species distributed evenly and assessed to the category LC and does not need commonly in the southern and central parts of monitoring.Al,Ab,N,Ka,St,Ta,Sa,Oa. Finland.Thisspeciesisassessedtothecategory LCanddoesnotneedmonitoring.Al,Ab,N,Ka, TherevainornataVerrall,1909 St,Ta,Sa,Kl,Oa,Tb,Om,Ok. Arathercommonspeciesinthesouthernandcen- tralFinland,butmay becommonestintheeast. Therevastrigata(Fabricius,1794) This species occurs mainly in sandy biotopes. A very rare species connected with sandy Thisspeciesisassessed to thecategory LCand biotopesandknownonly fromthesouth.There doesnotneedmonitoring.Al,Ab,N,St,Ta,Sa,Kl, arenorecordsofthisspeciesaftertheyear1943. Tb,Sb,Kb,Ok. It is possible that T. strigata is actually extinct fromFinland,butbecauseofthelimitedcollec- TherevalanataZetterstedt,1838 tionactivityitmightbeoverlooked.Atthisstage One of the commonest species of the genus in itisassessedtothecategoryCR.Ifapopulation Finland distributed especially abundant in the of T. strigata is found again fromFinland, then north. This species occurs mainly in sandy the population will need active monitoring. Al, biotopes.Thisspeciesisassessedtothecategory Ab,N. LCanddoesnotneedmonitoring.Ab,N,Ka,Ta, Sa,Kl,Tb,Sb,Kb,Om,Ok,Oba,Obb,Ks,Lkoc, Therevaunica(Harris,1780) Lkor,Le,Li. InadditiontotheoldknownpopulationinHanko area, N: Tvärminne, there are two new popula- [TherevamarginulaMeigen,1820] tions found, Ok: Sotkamo (Söderman & Lei- ASouthandCentralEuropeanspeciesdistributed nonen 2004) and Om: Raahe (Varpenius 2004). northtothesouthernSweden.Itmightbepossi- Thisspeciesmightbeconnectedwithdry heath bletofindT.marginulafromthesouthcoastof forestsandsandybiotopes.T.unicaisassessedto Finland. thecategoryVUandthelastknownpopulations ENTOMOL.FENNICAVol.17 (cid:127) FinnishTherevidae 55 andthepossiblenewfindingswillneedmonitor- torumFenniae,VI,Diptera.—HelsinginHyönteis- ing.N,Om,Ok. vaihtoyhdistys:1–63. Hackman,W.1980:AchecklistoftheFinnishDipteraI. Nematocera and Brachycera (s.str.). — Notulae TherevavalidaLoew,1847 Entomologicae60:17–48. Thisisarathercommonspeciesinthesouthern Kahanpää,J,&Winqvist,K.2005:Check-listofFinnish and central Finland. This species is assessed to flies:familiesXylophagidae–Microphoridae.—Sahl- the category LC and does not need monitoring. bergia10(1):10–27. Keil, P. 2003: Habitat and microhabitat preferences of Al,Ab,N,St,Ta,Sa,Kl,Kb,Om,Oba. mountainTherevidae(Diptera)andecologicaloutline of Czech Therevidae. — BSc thesis, University of Acknowledgements.WearegratefultoKevinHolstonfor SouthBohemia. adviceandhelpinformulatingthelanguageofthekeyand Lyneborg,L.&Spitzer,K.1974:TheCzechoslovakspe- assessingthebiologicalstatusofTherevidaespecies.We ciesofTherevaLatr.(Therevidae,Diptera)withthe alsothankTheFinnishEnvironmentInstituteforfinancing descriptionofanewspeciesfromHungaryandAus- thisstudyintheautumns2002–2004. tria.—ActaSci.Nat.Mus.Bohem.Merid.14:13–42. Rassi,P.,Alanen,A.,Kanerva,T.&Mannerkoski,I.(eds.) 2001:Suomenlajienuhanalaisuus2000.—TheFinn- ishMinistryofEnvironmentandtheFinnishEnviron- References mentInstitute,Helsinki. Stubbs,A.&Drake,M.2001:Britishsoldierfliesandtheir allies.—BritishEntomologicalandNaturalHistory Andersson,H.1994:AnewspeciesofAcrosathe(Diptera: Society.Pp.512. Therevidae)fromtheBalticIslands.—Stobaeana2: Zaitsev,V.F.&Bei-Bienko,G.Ya.(ed.)1988:Keystothe 1–6. InsectsoftheEuropeanpartoftheUSSR,VolumeV, Frey,R.1911:ZurkenntnisderdipterenfaunaFinlands.— DipteraandSiphonaptera,PartI.—SmithsonianIn- ActaSocietatisproFaunaetFloraFennica34:6. stitutionLibrariesandTheNationalScienceFounda- Frey,R.,Storå,R.&Tiensuu,L.1941:EnumeratioInsec- tion,Washington,D.C.