Entomological Review of Japan

76 Pages·1996·69.1 MB·English

Vol. 51 No. 1 June 1996
見 學評論
THE ENTOMOLOGICAL REVIEW OF JAPAN
日 本 甲 蟲 學 會
THE JAPAN COLEOPTEROLOGICAL SOCIETY
OSA KA
TheJapanColeopterologicalSociety
(Foundedin1945)
President: MASA0 HAYASHI
Managingdirectors:YAsUHIKoHAYASHI, KOZ0MIZUN0,HIDEY0NOMURA,TATE0ITO
Editorial Board
KATSURA MORIMOT0(Chairman), KOHEISAWADA、TAKASHIKISHll
NOBU0OHBAYASHI.YASUHIK0 HAYASHI
Councillor
HIROSHIKON0, YOSHIHIK0KUROSAWA, TAKEHIK0NAKANE,SHUN-ICHIUEN0
TIKA00HKAWA, SADANARIHISAMATSU, SHINSAKUKIMOT0
/_、 長 林 王l 夫
_、
運営委員 林 靖彦 水野弘造, 野村英世 伊藤建夫
編集委員 森本 桂 澤田高平, 岸井 尚 大林延夫, 林 靖彦
評 議 員 河野 洋 黒澤良彦, 中根猛彦 上野俊一, 大川親夫
久松定成 木元新作
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En†.Rev.Japan,51(l):l -8, June l5,1996
NotesontheSpeciesofStaphylinidae (Coleoptera) fromJapanX.
FourNewAdditionalSpeciesofLobratluum MULSANTetREY.
By TATE0 ITO
E7-303,0tokoyamaYutoku8,Yawata.Kyoto614Japan
Abstract Four new speciesof Loフ/・,(It/1111mare described from JapanunderthenamesofL.
)/t1,Ill-alspnov.、L.111二i1101spno、,..L.1.l1/it/tlispnov andL hokkaido'1、'sp nov.
InthepresentpaperofthisseriesIamgoingtodescribe「ournewadditionalspeciesof thegenus
Lobi-at/lift,11MULAsNATetREY.whichare recognizedbytheyellowisho「b「oWniSh「edpatch nea「apex
on eachelytron.underthenamesofL.o/11、-u1-(11sp nov from Kinki District, L. 'n、l,imoI sP nov from
chubu District、L.1s/11da1sp nov from Kanto District andL hokk(fide/1se sp nov from Hokkaido
District.
BeforegojngfurtherIwouldliketoexpressmydeepcordialthanksIoMr.T.SHIBATAforhiscritical
readjngofthemanuscriptandtoallthegentlemenwhosenamesareprintedinthesectionsoftypesand
specimensexamined.
l rar/1li//77oil・1f1'a s nov. (Figs.1-4)
o
Bodyrelativelylarge,robust,subcylindrica1,alittleshiny,black;elytraeachwithabrown
spotaboutonapicalthird,71habdominalsegmentbrownishapicad,and8thand9thsegments
almostwhollybrownish;mandibles, labrum,antennae, femoraandtibiaereddishbrown,
max1j1aryandlabialpalpi,farsiyellowishbrown.ventralsideofbodygenerallyreddishbrown
toblack;pubescenceonbodydarkbrowntoblack,thoseonantennae,mouthpartsandle9sPale
yellowtodarkbrown. Length:6.3-7.2mm.
Headsubquadrateralherthansuborbiculate,aboutaswideaslong,coarsely.closelyand
deeplypunctateexcept impunctateclypeus.puncturesonfronsandvertexmode「atelySpa「So
alongmjddle,andbecomingrelativelycloser lateradaswellasposteriad,eyesslightly
promjnent,thelongitudinaldiameteralittlelessthanahalfthelengthofpost9ena,
postgenaenearlyparallel-sidedandclearlyroundedtowardneck,antennaeslenderand
somewhatrobust,slightlyincrassatedistad,extendingbarelytobaseofpronotum,allSo9mentS
apparentlylongerthanwide,1stsegmentlargestbutlessthantwicethefen9thof2ndwhichiS
slightlyshorterthan3rd,4thIo6thsubequa1inlengthtoeachother 7thshorterthanthe
preceding,and1010thalsosubequa1inlengthtoeachother.11thconicalandaston9as3「d.
ventralsurfaceof headalmostuni「ormly,coarselyandcloselypunctate、puncturesnea「eyes
slightlysparser mentumclearly depressedonsides,submentumcoarsened,9ularPlatesmooth,
gularsuturessubpara11e1andfairlyseparatedfromeachother.
pronotumoblong_oval,slightlynarrowedbehind(length/width=1.27). considerably
longer(122: 1)andalittlenarrower(0.96:1)thanhead.coarsely,moreo「lessClosely
punctateexceptmedianline, thepuncturesbecomjng finerandobsoletenearapicalangles.
TATE0 ITO
medianlinerather wide,impunctatefrombasetoapex
butslightlynarrowedinmiddlebydisca1 punctures;
,
一 一 lateralmarginsinvisiblewhenviewed fromabove bul
thickthroughoutaswellasbothapical andbasal
margins.
Elytralongitudinallyoblong,subpara11e1atsides,
hardlydilatedapicad,alittlelongeralshoulders(1.08:
1)andwiderthanpronotum(1.23:1),ratiooflength al
shoulderstowidthat thewidestpointnearapexabout
1.12;surfacewithpuncturesdistinctlycoarser thanon
pronotum, not arranged in any rowsand becoming
slightly finer laterad:pleuralmarginsdistinctly
thickenedandpleural keelsreducednearalshoulders.
Elytra1marksill-defined notcircularinshape,touching
1 atapices.Scutellumpunctate.Prosternum smooth
apicadandslightlycoarsenedonitsprocess
m eso_
sternumdistinctlycoarsenedandmetasternumfinely
Fi9.1 oフ/ 'e'1/111- oi人l ll'ell sp nov : punctate
habitus.
Abdomenslightlyexpandedlaterad,divergent
graduallytowardbaseof7thsegment, thenconvergent1-apidlytowardapexoftheapjcalmosl
Se9ment;allsegmentswithanobservablemic1-osculpture;eachbaseoftergiteswithpunctures
Coarse,obsolete,becomingsparserandweakerapicad,thoseoneachsternitesimilartothoseon
theCon'espondingtergite.Inmale,6thto8thstemiteseachdepressedalongmiddle,6th_slema1
dep「eSsionhardlyornotperceptible 7th-sternaloneweak,widenedapicad,andwjthanarrow
andimpunctatemedianspacealongmiddlebefo1-emedianwideemarginationofapicalmargjn,
8th-stomaldepressiondeepfromnearbasetoapex,bearingfineblackgranulesexceptforboth
mediannalTowspaceandapico-margina]ratherwidespace,theapicalmarginnearlycircularly
-- -_、、-
一_一ーーー一
-
-
-
、
-
2 、
4
3
Fi9S.2-4. LoI''(It/ittt'11o/1kll'al spnov:2,aedeagusinlateralview;3,sameinventralview;4.oulljneofthe8th
stemiteofmale.
NotesonStaphylinidaefromJapanX
excisedinmiddle,andslightlyproducedasaprocessateachapicalangleof theexcision
(Fig4)
Legswithprofemoraveryrobustandprotarsimoderatelydilatedinbothsexes.
Aedeagus(Fjgs2-3)wellsclerotizedexceptdorsalside.withaventralprojectionlong,
thick,whollys1l-onglysclerotized.slightlycurvedventradandclearlytruncateatapex theapex
distinctlyhookedlateradandextendingwhollybeyondmedianlobe.
Aa.,... Holotype:ou Dorogawa Yoshino Nar Prof..3.V.1976 T.IToleg (T.SHIBAT coll) ParatyPes:1df.1早.
Ml.Kohjin、NaraPrer.,21.VIand15.VIII.1975,T.IToleg.11of, Mt.Shakagatake.Yoshino.NaraProf..l-i.X.1984,K.
HARUsAw,、leg.,;1一 M1.Daifugen,Yoshino,NaraPref..26.VI.199-1.K.HARLSIAWAleg.
,
specjmensexamined:3 早(tenera1).Mt.Kohjin,NaraProf..17.VIII.1965.8.VIII.1?)66.an(」1、'.Vm.1975.
T.lT0lcg.
ThepresentspeciesiseasilydistinguishedfromLobi・athiu,nntldtl1'1 (SHARP)inhavingthebody
largerand robusler, theaedeagusquitedifferntinshapeandthemaleabdominalstemitesdifferently
modifiedbythesecondarysexual features.
ThepresentspeciesisnamedtothememoryofthelateMr.MASAFUMIOHKuRAwhowasafOunde「
ofoursociety,theJapanColeoptero1ogica1Society.
rar/1lum mizzlf1ol sp nov. (Figs 5-7)
o
Body slender,subcylindrical,shiny,black:elytrawithapical thirdyellowishbrown,
someposteriorsegmentsofabdomen,greaterpartofventralsideofbody,mouth
apjcesof
paris,antennaeandlegsreddishb・lown apicesofappendicessomewhatyellowishbrown;
pubescenceonbodydarkbrowntoblack thoseonappendicespaleyellowtodarkbrown.
Length:5.6mm.
Headsubquadrate,justonlyalittlelongerthanoraboutaslongaswide,coa1-selyand
closelypunctateexcept forsparselypunctatefl'onsandvertexandimpunCtateClypeuS、eyes
moderatelysized,slightlyprominent,eachlongitudinaldiameterlessthanahalf thefen9thof
poslgena,postgenaeparallel-sidedandwidelyangulatetoneck,antennae1-elativelylong,
reachjngbaseofpronotum,allsegmentslongerthanwide,1stsegmentlarge,butshe「to「than
thefo11owjngtwosegmentstogether,3rdlongerthaneither2ndor4th,5thto7thsubequa1in
lengthIoeachother 8thalittleshorterthantheprecedingandaboutaslongasthefo11oWin9.
111hconjca1andlonger than10th.Ventralsurfaceof headmoreuniformlyand lessclosely
punctatethanonthedorsum,butweaklymicrosculptured.
pronotumova1.1ongerthanwide(1.23:1),longer(1.14:l)andalittlenarrower(0.95:1)
thanhead,slightlynarrowedbehind,coarselyandsparselypunctate, thepuncturessomewhat
jrregularinarrangementandbecomingslightlycoarseranddeepertowardlateralsides,median
linequitesmoothfrombasetoapex.
Elytrarathershort,slightlywidenedapicad.widestatapical third、wider(1.18:1)and
sljghlly1onger(1.04:1)thanpronotum;surfaceslightlyundulate,coarselyandobsoletely
punctate,thepuncturesmuchcoarserthanonpronotumandnotseriatelyarranged,elyt「alma「kS
similartothoseontheprecedingspecies.
Abdomenslightlyexpandedlaterad.whollywithanobservablemicrosculpture, ter9a1
puncturescoarseatbaseandfineatapex. Inmale、4thto8thsternitesmoreor lessdep「eSSed
alongthemiddle, thedepressiononeachsternitewideranddeeper thanonthepreceding
TATE0 ITO
-
/
,
-
-,
/
7
Fi9S・5-7. Lob''athl1l'11nli1lno1spnov:5、aedeagusinlateralview:6、sameinventralview:7,outlineofthe81h
stemiteofmale
Ste「nife,4thand5thsternitesveryfaintlydepressed,6thto8thsternitesdistinctlydepressed,6th
Ste「nifewithsmallandroundeddept'ession,7thsternitewithratherwideandU_shaped
deP「eSSOin,boththedepressionssmoothandimpunctate,apicalmarginof7thsterniteslightly
andwidelyemarginateinmiddle,8th-stomaldepressionshallowalbase,horseshoe_shapedand
furnishedwith fineblackgranulesexceptwidelysmooth, flatandtriangularspacebefore
medianexcisionofapicalmargin,theexcisionwide,deepandsemicircular(Fjg7).
Legsusuallywithprofemoravery1-obustandprotarsidilatedinbothsexes.
Aedeagus(Figs5-6)consistingofi11-scle1-otizeddorsalpartandwe11-sclerotizedventral
P「ejection, theventralprojectionalmostsymmetrical,unevenlycurvedventrad, fajrly
Constrictedatbase,fromtherethickendapicad,bisinuateonapico-1alera1sides.widestalapjca1
fifth,theapexwithaspoon-likedepressionondorsalsurface,andnotpointedattjp
Holotype:oア.GOZaishiSpa、YamanashiProf..13.V.1989、K.HosoDAleg.
(T.SHIBATAcol1) para1ype :1
samelocalityastheholotype,27.VI.1990.K.HosoD、、leg.
TheP「eSentspeciesisalsorelatedtoLobi'at/tiumnudtml ,buttheaedeagusisquitedifferentinshape,
theVent「alp「ejectionof theaedeagusismuchwiderandlarger,themaleabdomjnalslernjtesare
diffe「entIymodified the7thstemitehasnotanytuberclesthe8thsternitebearssomefjneblackg]-anules
theantennalSo9mentSaremoreeton9ate、theheadismorecloselypunctateandtheelytraarenotseriately
punctateonsulface.
ThespecificnameisdedicatedtoMr.K.MIzuNo,whohaveofferedmemanystaphylinidspecimens
capturedbyMr.K.HosoDAfromGozaishiSpa.
oフt rathzMm z'shldazsp
nov
(Figs8-10)
Bodysubcylindrica1,colorationandpubescencesimilartotheprecedingspecjes.Length
Nolo・conStaphylinidaefromJapan X
5.9-6.3mm.
Headsuborbiculate aslongaswide.coarsely,closelyanddeeplypunctateexcept for
sparselypunctatevertexandfronsandimpunctateclypeus thelongitudinaldiameterofeye
nearlyequal Ioahalfthelengthofpostgena.postgenaemoderatelyroundedbehind antennae
ratherlong,robust,slightlyincrassatetowardapicalsegment reachingbarelybaseofpronotum,
allsegmentsapparentlylongerthanwide、1stsegmentlargest,veryrobustandmorethant5
tjmesaslongas2ndwhichisalittleshorterthan3rd,4thto6thslightlyshorteneddistad,7thto
lothsubequal inlengthtoeachother,11thconicalanddistinctly longer than10th. Ventral
surfaceofheadcoarselyandrathersparselypunctate,thepuncturesbecomingsparserlaterad,
mentumclearlydepressedonbothsides.submentumcoarsened gularplatesmooth,9ular
suturesfairlyseparatedandsubpara11e1toeachother.
pronotumoblong-ovate(length/width=1.27),moderately longer(1.22:1)andalittle
nan-ower(0.96:1) thanhead, faintlyna1Towedbehind,disca1puncturescoarse, ratherclose,
somewhatirregularlyarrangedandfairlysparserthanonpostgenaeofhead,medianlinewholly
smoothbulslightlynarrowedinmiddle,lateralmarginsinvisiblewhenviewedfromabove.
Elytraoblong,scarcely widenedapicad about aslongasandwider (1.14 : 1) than
pronotum,ratiooflengthatshoulderstowidthatthewidestpointnearapexabout1.12;surface
wjlhpuncturesmuchcoarserthanonpronotum,andnotseriateina1Tangemen、teIytra1markings
placedalapicalthird.ill-definedandtouchingtomarginsofapices.Wingswelldevelopedand
functional.
Abdomenslightlyexpandedlaterad allsegmentsweakly microsculptured;eachbasal
tergjtewithpuncturescoarseand obsoleteatbase,becomingsparserandweakerapicad,those
onsomeapjca1tergilesthroughout fine. Inmale,5th sternitescarcelydepressed,6th sternite
weaklyand71hslernilerather deeplydepressed thedepressionof7thsterniteU-shaped.
wjdenedapicadandwithanarrowsmoothspacealongmiddle apicalmarginof7thsternite
feeblyemarginateinmiddle,8thsternitedepressedfrombasetoapex, thedepressiondivided
l 1/
l ノ
/
、、 1
8
\ /:
、
ノ
10
9
Fjgs 8_10 L1/)1・al/1j″l11is1/idtli spnov.,8.aedeagusinlateralview:9. sameinventral view: 10,outlineofthe
Sthsterniteofmale.
TATE0 IT(1
intotwopartsbytheconstrictionatbasalthird,ofwhichthebasalpari smallandshallow.the
apicalonedeepandhorseshoe-likeinshape.clearlyedgedalongborderline,withfinegranules
except thataverynan'owmedianspaceandmarginalareasmoothalongapicalexcjsjon the
excisiondeep,oblonginoutlineandslightlyprotuberantatitsapicalangles(Fig.l0).
Aedeagus(Figs8-9)comparativelysimpler inconstructionthanthatof thepreceding
species. ventralprojectionevenlycurved notsinuateonapico-1atera1sidesandproportionally
widerinmiddle,theapexwithapairedhooksondorsalside.
Holotype:d?.Mt.Nasu、TochigiProf.,16.Vm.l959.T.SHIBATAlcg.(T.SlllB、TAcoli).Paratype:1、..?amedata
a、theholotype.
SPecin、en、.examined:)d?くi、(tenera1).samedataastheholotype.
ThepresentspeciesissimilaringeneralappearancetoL.o/1kl1l-c11sp nov.withexceptjonof the
followingdistinctionsbetweenthetwospecies:inthepresentspeciesbodylessrobustandsmallerinsize
elytrarelativelyshorter not longerthanpronotum、 male8thstemiteofabdomenmoredeeplyexcjsed.
ventralprojectionofaedeagusmuchnarrowerinwidthanditsapexquitediffernl inshape,inL.o1lku1-a1'
body largerand robuster,elytraslightly longerthanpronotum male8thslernilesemicircularlyexcjsed
aedeaga1projectionlongandthickanditsapexhooked.Thepresentspeciesisalsoa11jedIotheL.,11t-u11()1'
sp nov. inthesimilarshapeofaedeagus,but the excisionof themale81hslernile has asmall
p「otube「anceateachapicalangle,theelytraarenotlongerthanpronotuman(」theheadissuborbjcular
ratherthansub(luadrateinprofile.
ThespecificnameisdedicatedtothelateMr.HIROSHIIsHIDAwhowasamanagingdirectorofour
society.
Lobi・at/1iun111okkaidense sp.nov. (Figs.l1-14)
Body slender,rathersubdepressedabove,alittleshiny,black:elytraeachwithayellow
markinginapicalareaapicesofposteriorsegmentsofabdomenslightlyreddish,ventralsideof
bodygenerallyb1-ownishtoreddishblack,mandibles.
1ab1'Im,basal segmentsofantennaefemoraand tibiae
- - ・
、 , reddish brown maxillaryandlabial palpi apical
segmentsofantennae tibiaeandtarsireddishyellowto
brown. Length:5.6-6.6mm.
Headnearlyorbiculate,alittlelonger thanwide,
slightlyunevenbetweeneyes,coarsely, ratherclosely
anddeeplypunctate,butfronssparselyandmoreorless
irregularlypunctateinart'angement,clypeusandlopof
vertex impunctate, eyes moderately sized, the
longitudinaldiameteralittlelessthanahalf thelength
ofpostgena,postgenaesubpara11e1-sidedandclearly
roundedbehind,antennaelongandslender.slightly
incrassatedistad,extendinghardlytobaseofpronotum.
11 allsegmentsclearlylongerthanwide.1stsegmentvery
robustandtwiceaslongas2nd,3rdclearlylongerthan
thepreceding 4th to10thsubequa1inlengthtoeach
Fig. ll. ot),-t11/111l/11 1/0人人c11t/e11.le SP other. 11thconical and longer than 10th. Ventral
no、,.: habitu、
NI)lc、)nStaphylinidaefromJapanX
一、、、、、、
/
ノ 、
、
, 、.
, 、、
ー、
.、
12 、
-
/
ノ /
14
13
Figs12_l o/)1-rtf1/1'l川l /10 ~(/- so 、、p no、,.: 12. acdeagusinlateral、'low; 13, amein、'entra1、low; 1
outlineofthe8thstemiteinmale.
surfaceof headwjthfaintandaciculatemicrosculpture,moreregularlyandlessCOa「Sely
punctatethanonthedorsum,mentumandgularplatesmooth、submentumuneven,gularsutures
fairlyseparatefromeachother.subparalle1butslightlynarrowedtowardneck.
pronotumoblong_oval,longerthanwide(1.28:1).longer(1.17: l)andalittlenarrower
(0g5: l)thanhead,slightlynarrowedposteriad.coarsely、sparselyandl'ather irregula「ly
punctate, thepuncturessparser thanonpostgenae medianlineratherwide, impunCtatef「om
basetoapexbulslightlyinterruptedbydisca1punctures;apicalanglesmoregentlya「Cuatethan
basalangles.1atera1marginsinvisiblewhenviewedfromabovebutthickthroughoutasWellas
bothapicalandbasalmargins.
scutellumdislincllypunctate.Prosternumsmoothapicadandweaklyre9ula「basad.
mesoslemumcoarsenedandmetasternumfinelyanduniformlypunctate.
Elytralongitudinallyoblong,slightlyenlargedapicad,subpara11e1atsides.slightlyIon9e「
(1.09:1)andfairlywiderthanpronotum(1.17:1),ratioofhumeral lengthtothemaximum
wjdlhnearapexabout1.20;puncturesslightlyobsoleteandalittlecoarserthanonP「onOtum,
an-angedinregularrows,yellowmarkingssituatedatapicalthird,joiningtoapicalmarginsbut
separatedfromlateralmargins;pleuralmarginsdistinctlythickenedandpleuralkeelsolea「but
evanescentnearIoshoulders.Wingswelldevelopedandfunctional.
Abdomenwithmicrosculplurewhollyobservable,punctures ratherfine,closeand
becomjngsparsertowardapicalsegments.Inmale、4thslerniteveryfeebly,5thand6thfairly
depressedalongthemiddle,71hclearlydep・tessedfl-ombasetoapex andapicalmar9inof7th
sternjteweaklyemarginateinmiddle,8thsternitealsoclearlydepressedthrOu9hOut, the
depressionlongitudinallyelongate horseshoe-likeinshape,bearingfineblack9ranuleSexcept
formoderatelywidesmoothareaalongexcisedapicalmargin,theexcisionrelativelyShallow・
semicircularandprovidedwithasmallprocessateachapicalangle(Fig.14).
Aedeagus(Fjgs.12-13)withventralprojectionheavilysclerotized,alittleasymmet「leafin
profjle, ratherslender,slightlythikenedapicad,stronglycurvedventradespeciallynea「base,
TATE0 ITO
theapexclearlyrounded.
Holotype: .JyOZankei.Hokkaido23.VIII.1964,M.YAsulleg(T.SHIBATAcoll).paratypes:1 '.samedata
asholotype:1 Chienbetsu,Hokkaido,25.VII i962.K.UEDAleg:1早same1ocalily.1 vm.lg81.N YAsUDAleg:1
,
diSouunkyo,Hokkaido.18.VII.I980,N.YAsUDAleg.
TheP「eSentspeciesisdistinguishablefromtheprecedingspeciesbyhavingtheaedeaguswjlhthe
Venta「alP「ejectiondifferentinshape,thesubapicalpartofventralprojectionnotemargjnatelalerad,the
maleSte「na1deP「eSSiOnSColTesPondinglyweaker thedepressionofthe81h slemitenotangularlyedged
alongborderline, andthebodyslightlyslendererandlessconvexabove
Additional reference
ITO,T・1995. NotesontheSpeciesofStaphylinidaefromJapan IX.Thedescrjpljonsofthreenewspeciesof
Lob''at11iu'11MuLSANTetREY(Coleoptera).Ent.Re1,.Japan、50(2):10g_1I8
(ReceivedDec. l8. l995;AcceptedFeb29,1996)