A Systematic Revision of the Clavigerine Genus Diartiger Sharp from East Asia (Coleoptera, Staphylinidae, Pselaphinae) PDF
Preview A Systematic Revision of the Clavigerine Genus Diartiger Sharp from East Asia (Coleoptera, Staphylinidae, Pselaphinae)
九州大学学術情報リポジトリ Kyushu University Institutional Repository A Systematic Revision of the Clavigerine Genus Diartiger Sharp from East Asia (Coleoptera, Staphylinidae, Pselaphinae) Nomura, Shuhei http://hdl.handle.net/2324/2610 出版情報：ESAKIA. 37, pp.77-110, 1997-09-30. 九州大学農学部昆虫学教室 バージョン： 権利関係： ESAKIA, (37): 77-l 10. September 30, 1997 A Systematic Revision of the Clavigerine Genus Ear-tiger Sharp from East Asia (Coleoptera, Staphylinidae, ) N ShOhei OMURA tnemtrapeD fo ,ygolooZ lanoitaN ecneicS muesuM .taN(,).tsiH -32-3 1 ,6hc-ninukayH ,ukujnihS ,oykoT 961napaJ Abstract. ehT eniregivalC suneg kubotai morf napaJ dna .D k1fnnzil~gerl.si.s from Yunnan, Southwest China are described. D. izuinsulicola ,.von ispartae (Karaman), morimotoi ,.von imasakai ,.von hirashimai ,.von dentipes arumoN te eeL dna eht lacipytonimon .seicepsbus owT nwonk-elttil.seiceps D. spinipes prahS dna D. juponicus .K( )adawaS era .debircsedersdroceR fo eht tsoh stna era.deton yeK :sdrow ,ymonoxaT ,aretpoeloC ,eadinilyhpatS Diartiger, revision. tsaEaisA Introduction The genus Diartiger described by Sharp (1883) has been well known as a evitatneserper fo eht suolihpocemrym selteeb ni napaJ ,ihcadA( :0591 ,ikarihS.)45.91 ehT epyt ,seiceps D. (19X), though its taxonomical treatment has never been discussed. Modified structure of its mouthparts was observed and noted by Kubota (1956). Kistner (1982) and Ohishi (1992). On the other hand, D. spinipes prahS debircsed ta ehtemas emit sa eht remrof seiceps sah ton neeb dedrocer niaga ecnis sti lanigiro noitpircsedrof revo eno .yrutnec arumoN dna eeL )2991( yltnecer debircsed D. dentipes morf uj_ehC ,.sI fo htuoS ,aeroK hcihw saw osla dedrocer morf eht naeroK alusnineP yb arumoNdna Lee (1993). Besuchet (1986) revised the genera Microdiartiger .K adawaS dna Co<ffLitirr.v ,namaraK dezimynonys meht htiw Diartiger, dna detcerroc eht suoenorre epytytilacol ”yek rfuoT“ eh tseepiyct efpos e rh,etstutna el.gC ispartae namaraK ot.”napaJ“ 1) sihT yduts si detroppus ni trap yb a dia-ni-tnarG .oN 3114070 1 rofcifitneicS hcraeseR fo na lanoitanretnI cifitneicS hcraeseR margorP morf eht yrtsiniMfo ,noitacudE ,ecneicS stropS dna ,erutluC.napaJ 78 .SARUMON nI siht ,yduts eht yllacyhpargoeg elbairav ,seiceps .D sutalussof si deifissalcotni neves .seicepsbus .D eutrapsi )namaraK( dna .D sepitned arumoN te eeL era dedragersa sti .seicepsbus Two little-known species, .D jqonicus .K( )adawaS dna .D spirlipes prahS era debircseder dna rieht wen lanoitubirtsid sdrocer era .dedda owT wenseiceps are described, namely, D. kubotai morf napaJ dna D. kunmingerzsis from Yunnan. .anihC dnA sdrocer fo tsoh stna era deweiver rof lla eht.seiceps suneG Dim-tiger Sharp esenapaJ[ ih s:uemmaaknuzira-otubegih]-uoktoaZmaY Dim-tiger Sharp, 1883, Trans ent. .ceS ,nodnoL :3881 ;923 ,yarffaR ,0981 euveR..tnE :9 ;561 ,yarffaR ,4091 .slnnA .toS .tne ,.rF :37 ;464 ,yarffaR ,8091 .neG,.snI :46 ;734 ,yarffaR ,1191 .tpoeloC ,.taC :)72( ;671 ,kraP ,8491 .lluB ogakihC .dacA ,.icS :8 ;812 ,lennaeJ ,8591 .m&M .suM .tsiH ,.tan ,siraP ,)A( :81 ;521 notweN & ,reldnahC ,9891 anaidleiF ,.looZ .N( ,).S :)35( .46 epyT :seiceps Dial-tiger fossulutus ,prahS detangised yb ,sacuL.0291 Microdiartiger .K ,adawaS ,4691 .tnE .veR ,napaJ :71 ;21 ,tehcuseB ,6891 euveRessius ,.looZ :39 362 ;)dezimynonys( notweN & ,reldnahC ,9891 anaidleiF ..looZ.N( ,).S :)35( .46 epyT :seiceps Microdiurtiger japonicus .K ,adawaS yblanigiro .noitangised Coiffaitius ,namaraK ,9691 .loiB ,acinelleH-ollaG :2 ;05 ,tehcuseB ,6891 euveRessius ,.looZ :39 362 ;)dezimynonys( notweN & ,reldnahC .9891 anaidleiF ..looZ.N( ,).S :)35( .46 epyT :seiceps CoiYuitius ispurtue ,namaraK yb.ypytonom ydoB -llams ot ,dezis-muidem 3.2-3.1 mm ni ,htgnel denedaorb ,ylroiretsopylgnorts desserped ta esab fo.nemodba Head longer than wide, nearly subcylindrical, weakly broadened anteriad. Eyes ,depoleved detacol raen eht dim level fo .daeh straphtuoM erom ro ssel ,decudertsomla elbisivni ni lasrod .weiv eannetnA trohs dna ,kciht denekciht ,yllatsid .detnemges-4 1st dna dn2 stnemges yrev ,trohs dr3 egral dna ,etagnole ht4 eht.tsegral mutonorP ,esobolgbus htiw a naidem-isab dna a riap fo laretal-isab .eaevof artylE suoirav ni erutcurts fo las r,oedcafru shcae nortyle htiw a egr aelmo hlcair)rtetymloeh(cirt ta laretal-oretsop .trap sgeL trohs dna ,tsubor sretnahcort yrev egral dna .etagnoledim sretnahcort hcae htiw a egral enips raen xepa ni elam fo ynam ,seiceps dimaromef denekci hrtaen eht elddim ni ,larene ghtiw a egral enips no roirets oepdis ni elam niynam ,seiceps dim eaibi t,r eedtnaelluscitned no renni edis ni.elam nemodbA yrev ,egral dednuor no htob laretal dna roiretsop ,sedis dr3 ot ht6setigret yletelpmoc desuf ot hcae rehto ot mrof a egral etisopmoc ,mugret htiw a egral dnapeed naid eymt-iivsaacbnoc dna a riap f olar e,tsaelm-oihscaibr th claaeret aelm-oihscaibrtkciht ta roiretsop ,trap neht gnidnetxe yllaidem-oretna ot mrof a esrevsnart egnirf ni eht basi- naidem ytivacnoc ni emos ,seiceps dr3 ot ht6 setigretarap desuf ekil setigret otmrof etisopmoc ,agretarap hcae etisopmoc mugretarap ,etagnole deworran .ylroiretsop htiwa basal trichome (paratergal trichome) and an elongate to semilunar depressed part lagretarap( )csid tsuj dniheb eht lagretarap ,emohcirt ht7 ot ht8 setigret ,trohssemitemos NOISIVER FO DZARTZGER MO RTFSAAEISA 79 elbisivni ni lasrod ,weiv dr3 ot ht4 setinrets desuf ot hcae ,rehto detcirtsnoc dnaevacnoc ni lasab ,trap ht5 ot ht8 setinret shcae ll.edwetakramed sugaedeA llew ,dezitorelcs naidem ebol dedivid otni lasab blub dna lacipa .ebollasab blub htiw a lasab eludon gniniatnoc lasab nemarof dna na diovo suonarbmem trapno lasrod ,edis lacipa ebol deworran ,yllatsid htiw egral lacipa eciffiro no lasrod.edis sullahpod nyelka edwezitorelcs dna yrev rednels ni ynam.seiceps Remarks. sihT suneg si deziretcarahc yb detnemges-r ueoafnnetna htiw gnol drihtdna htruof .stnemges tI si tsom ylesolc deilla ot eht suneg vegirtylerciM arumoNyltnecer debircsed morf eht suykuyR dna nawiaT ni weiv fo eht ytiralimis fo eannetna dnaeht serutcur tf sosemohcir tno eht artyle dna eht.nemodba Diartiger fossulatus Sharp esenapaJ[ :eman]ihsumakuzira-otubegih-otamay-oK Diartiger fossulatus ,prahS ,3881 .snarT .tne .toS ,nodnoL :3881 ;033 ,esuohretaW 1882-90, diA .tnedI ,.snI :2 ;641 ,yarffaR ,4091 .slnnA .toS .tne ,.rF:37 464: Raffray, 1908, Gen. Ins., 64: 438; Raffray, 1911, .tpoeloC ,.taC :)72( 176: ,itugakaS ,3491 ,op-nakuhcnoK-akuzarakaT :)73( ;2 ,kraP .8491 .lluB ogakihC .dacA ,.icS :8 ;812 ,lennaeJ ,8591 .meM .suM .tsiH ,.tan ,siraP ,)A( :81;621 Nomura, ,9891 kcehC tsiL .npJ ,.snI ,akoukuF :1.392 Male. htgneL 42.2-88.1 .mm htdiW 58.0-07.0.mm ydoB de n,eydlarooribre ttssoepdiw ta eht elddim fo.nemodba Head longer than wide, subcylindrical, gently thickened anteriad, covered with ralugerri noitaluciter no lasrod ;ecafrus suepylc yrev ,trohs ylthgils denedaorb.dairetna ylraen etaucra no roiretna ,nigram snorf ylgnorts ,xevnoc ralugnairt ni roiretna,trap xetre vylthgi l,sxevnoc htiw a riap fo tcnitsi dlnaisr oldairotnet stip tsuj edisni ,seyeeaneg ylkaew deworran ni tnorf fo ,seye eanegtsop etalugna .yllaretal-oretsop seyE llamsdna ,diovo hcae desopmoc fo tuoba 51 .stecaf eannetnA ,trohs gnihcaer roiretna nigramfo ,mutonorp 45.0-54-O mm ni ,htgnel denekciht ,yllatsid ts1 tnemges yrev trohs dna elbisivni ni lasrod ,weiv dn2 ,trohs ylthgils rediw naht ,gnol ,lacirdnilycbus dr3etagnole dna denekciht ,yllatsid ht4 eht ,tsegral regnol naht ts1 + dn2 + 3rd, .lacinocbusderevoc htiw gnol dna tcerebus sriah no laretal ,ecafrus ylwollahs evacnoc dna ylesnedderevoc htiw trohs eates ta .xepa straphtuoM ylgnorts ;decuder murbal yrev trohs dna.esrevsnart yltneg etaunisib no roiretna ,nigram htiw a riap fo raludnalg secifiro raen eht:esab selbidnam trohs dna ,kciht ylesutbo etalugna ta ,xepa hteet ,tcnitsidni hcae htiw aegral raludna legcifiro ta laretal ,trap eallixa mdecuder dna yl k,adeewzitorelcs aelag dnaainical yllamron esotes ta ,xepa yrallixam palpi hcae decuder ot a trohs dna elpmis.etirelcs nellows ta,.xepa muibal gnisirpmoc dezis-lamron ,mutnem tnerapsnartbus dnalamron assolgara pdna a riap fo decude rlaibal palpi. mutonorP tuoba sa gnol sa ,daeh rediw naht ,daeh ,esobolgbus sa gnol sa.ediw derevoc htiw ralug enroriitaluciter dna trohs dna dlob sriah no lasrod .ecafrus htiw aegral naidem-isab dna a riap fo kaew laretal-isab .eaevof munretsateM esrevsnart dnaylgnorts xevnoc tsuj dniheb ,eaxocose mylesned derevoc htiw lanidutign orlaenil erutplucs nohtob laretal ,sedis htiw a lanidutignol emohcirt gnola naidem .enil artylE deworran.ylroiretna .SARUMON ylraen ,ladiozepart ylkaew ,xevnoc yletaucra etanigrame no roiretna ,nigramylwollahs etanigrame no roiretsop ,nigram hcae nortyle htiw 2 tcnitsidni lasab eaevof dna 2 tcnitsidni lanidutignol iclus ta lasab ,trap lartyle emohcirt ,egral deneprahs.ylroiretsop sgeL trohs dna ,kciht sretnahcort etagnole dna denekciht ,yllatsid dim sretnahcorthcae htiw a egral enips no lacipa-oretsop ;edis aromef ,kciht hcae rumef htiw 2lanidutignol eanirac no renni ,edis dim arome fyltneg ,tneb hcae htiw a egr aellcitnedla snrood-oretsop edis ni emos ,seicepsbus eaibit ,rednels ylesraps derevoc htiw tcerebus ,sriah dimeaibit ,ydlegnne othrtctaaslef htiw a l lealmcsit nreaden eht.elddim Abdomen widest and very large, slightly wider than long, rounded posteriorly. etisopmoc mugret ylpeed evacnoc ni lasab ,trap xevnoc ni lacipa ,trap ylesrapsderevoc htiw gnol tcerebus ,sriah laretal-isab emohcirt suoirav ni ,epahs erom ro sseldelruc ,yllanretni htiw a egral esrevsnart egnirf ni sti naidem-oretna ,trap lagretarapemohcirt egral dna ,kciht lagretarap csid ylthgils rellams naht lagretarap ,emohcirt regnolnaht ,ediw ht7 etigret trohs dna ,esrevsnart ladiozepart ni roiretsop ,weiv ht8 yrev llamsdna ylraen ralucricimes ni roiretsop ,weiv ht4 etinrets tsegral dna .esrevsnart htiw awollahs dna tcnitsidni esrevsnart evoorg ta ,esab ht5 ot ht7 ylevisseccus denetrohs ,dairetsophcae trohs dna ,esrevsnart derevoc htiw tcnitsidn inoitaluciter no htob laretal ,sedis ht8 small dna ,ralucricimes ylkaew.xevnoc sugaedeA egral dna ylg n,odretzsit osreerlec ms,atrcanpitsid nniaidem ebol -lellarapbus dedis dna dednuor ta ,esab htiw a egral dna llew detcejor plasab .eludon a egral dnadiovo suonarbmem trap no lasrod edis dna a egral dna peed lartnev ,noisserped lacipa ebolyrev llams dna ladiozepart ni lasrod ,we isvullahpodn eyrev rednels dna ,niht elbisivni niemos cases. .elameF htgneL 52.2-88.1 .mm htdiW 58.0-17.0 .mm ralimiS ot ,elam tubeannetna ylthgils retrohs naht ni elam ni ,larene g5.0-44.0 1 mm ni ,htgnel dim sretnahcotrutohtiw ,enips dim aromef erom rednels dna ylrae n,thgiar ttsuoht i,welcitned dim eaibityllamron ,rednels tuohtiw.elcitned Distribution of the species. htuoS aeroK dna napaJ ,odiakkoH( ,uhsnoH ukokihSdna Kyushu). .skrameR sihT seiceps si ralimis ot D. spinipes prahS dna D. kuhotai sp. ,.vontub y lsrraeefl fcmiodrf meht ni gniv ayhre vll almasci peabol fo .eshutgaedea A gnikirts lacihpargoeg noitairav si dezingocer ni siht .seiceps tI si deredisnoc tonot be a enilc tub a de tnaouittcaniurpav no eht sisab fnooitanima xfeo yn asmnemicepmsorf lla revo napaJ dna htuoS .aeroK tI si erofereht dedulcnoc taht siht seiceps nac eb deifissalc otni neves seicepsbus sa debircsed ni siht.yduts Host ants for subspecies indetermined: Paratrechinajluvipes :htimS ,akasuK etawI onakaN(.ferP & ,adihsoY ;)9391 otoyKytiC ,itugakaS( ;)3491 seitilacol nwonknu ,ihcukonaT(.)b9791 Lasius niger :sueanniL ,awakasA oykoT ;.ferP ,ekatiM oykoT ;.ferP ,enokaHawaganaK Pref. (Nakano & Yoshida, 1939); nijGyR Spa, Wakayama Pref.; Kyoto City (Sakaguti, 1943); Minamizawa, Tsukui-gun, Kanagawa .OyriK(.ferP 1978): seitilacol nwonknu ,ihcukonaT(.;)b9791 Lasius umbratus (Nylander): ,emhgog-nihS Mt. Fuji, Shizuoka Pref.(Tanokuchi, 1979a);. Formica juponica Motschulsky: ,emagog-nihS .tM ,ijuF akouzihS.ihcukonaT(.ferP ;)a9791 NOISIVE RFO DZARTZGER MORF TSAEAISA 18 Diartiger fossulatus fossulatus Sharp Male. htgneL 9.1 l-2.09 .mm htdiW 08.0-57.0 .mm ydoB .dezis-muidemeannetnA ylevitaler ,trohs llew denedaorb ,yllatsid 84.0-54.0 mm ni .htgnel .o-11.0 31 mm ni ,htdiw ht4 tnemges yrev ,kciht 3.2-2.2 semit sa gnol sa .ediw diM sretnahcort,etagnole hcae htiw a egral dna etuca enips no lacipa-oretsop ,edis dim aromef .tsubor hcae htiwa egral dna ralugnairt elcitned raen eht elddim no roiretsop .edis laretal-isaB emohcirtfo etisopmoc mugret yrev daorb ta ,esab thgiarts dna ylecracs deneprahs ta .xepalagretarap emohcirt egral dna ,et algangorleetara pcsid etagnole dna ylrae.nmrofisuf .elameF htgneL 11.2-40.2 .mm htdiW 38.0-67.0 .mm eannetnA 94.0-44.0 mm ni ,htgnel 31.0 mm ni ,htdiw ht4 tnemges 3.2-0.2 semit sa gnol sa.ediw snemicepS examined. ]odiakkoH[ 1 ,elamef ,ibiB esotihC ,.C .irakihsI .02 .iv .4-.iiv ,2991 .S iroH ;.gel 1 ,elam uotnomiK ,ekaL ,8hc-ikiaT ,ihcakoT .62 .iiiv .3991 .SiroH ;.gel 1 ,elamef ,nagiak-atikoyoT ,hhc-oroharU ,ihcakoT .62 .iv .7- .iiv .3991 .SiroH .gel ]uhsnoH[ 1 ,elam 1 ,elamef ,ikasim-ippaT ,arum-ayamniM iromoA ..ferP .03 .xi ,2991 .S arumoN;.gel 1 ,elamef ,onusaY ikamanaH ,.C etawI ..ferP .2 .x .2991.S arumoN ;.gel 1 ,elamef ,otasiiN onhoT ,.C etawI ,.ferP .1 .x ,2991 .S arumoN :.gel 1 ,elamef ,emonihcO atakaS ,.C atagamaY ,.ferP .3 .x ,2991 .S arumoN ;.gel 1.elamef ,uykA igayiM ,.ferP .41 .iv ,6891 .Y onariH;.gel 1 ,elamef .tM ,amayirotoKamihsukuF C., Fukushima Pref., 24. ix. 1987, K. Tazoe leg.; 2 males, 2 females, Natsui. ,arum-awakoruK atagiiN ,.ferP .3 .iiv ,5891 .S arumoN ;.gel 1 ,elamef.ariad-ianiaT Kurokawa-mura, Niigata Pref., 2. ix. 1991, H. Koike leg.; 1 male, Higashizawa, ,awazubotO ,arum-oneU amnuG ,.ferP .5 .v ,9791 .Y awakihsiN ;.gel 1 ,elam 1,elamef ,okkiN igihcoT ,.ferP .02 .v ,8791 .Y onariH ;.gel 1 ,elam .tM .nasozimaYikarabI ,.ferP .92 .v ,8891 .Y onariH ;.gel 4 ,selam 2 ,selamef ,ajnij-ikasonuS amayetaT..C abihC ,.ferP .21 .v ,6991 .S arumoN ;.gel 1 ,elam 1 ,elamef ,amayukaK .ihcam-awagO amatiaS ,.ferP .03 .iii ,3991 .T ubmaN;.gel 1 ,elamef emas ,ytilacol .31 .ix .3991.T ubmaN ;.gel 2 ,selamef ,ariadaK-ihsagiH ,nonnaK ,ihcam-amadoK amatiaS ..ferP.71 .iiiv ,9891 .T ubmaN;.gel 1 ,elam ,ihcam-oakatarU ijhihcaH ,.C oykoT ..ferP 2 .I .v ,4991 .S adihsoY ;.gel 1 ,elamef .tM ,nasoakaT oykoT ,.ferP .12 .vi ,0691 .YatabihS leg.; 1 female, Mt. Mitakesan, Tokyo Pref., 5. v. 1956, Y. Shibata leg.; 1 ,elam 1 ,elamef ,awiM adihcaM ,.C oykoT ,.ferP .01 .v ,0891 orihuzaK adawaS ;.gel 1,elamef ,igasiH ihsuZ ,.C awaganaK ,.ferP .21 .vi ,7591 .K adihsI ;.gel 1 ,elam.ekakaduF ,awaznaT awaganaK ,.ferP .61 .iv ,7891 .Y onariH ;.gel 1 ,elamef emas ytilacolsa above, 17. v. 1994, Y. Hirano leg.; 4 males, 4 females, Mt. ,amayQ Tanzawa. awaganaK ,.ferP .5 .v ,0691 .Y atabihS ;.gel 1 ,elamef emas ytilacol sa ,evoba .3.iiv ,7791 .Y onariH ;.gel 2 females, Utsukushigahara, Nagano Pref., 5. v. 1959. K. awasuziM ;.gel 6 ,selam 2 ,selamef emas ytilacol sa ,evoba .1 .v ,0691 .Y atabihS.gel Japan: Hokkaido, Honshu ,ukohhT( ,8tnaK ubithC Distribution of the subspecies. stcirtsid dna atagiiN.).ferP Remarks. ehT lacipytonimon seicepsbus si deziretcarahc yb eht yllacipa denedaorb eannetna htiw yrev kciht htruof ,tnemges dna yldrah deneprahs laretal-isab emohcirtfo eht etisopmoc .mugret tI si detubirtsid ni eht egral aera fo eht nrehtron dna nretsaestrap fo .napaJ A melborp si no eht noitalupop fo enokaH hcihw si detangised sa ehtepyt ytilacol fo siht ,seiceps sa lliw eb deton ni eht noitces fo D. fossdatzts subspecies 82 .SARUMON indetermined under the part of .D if ispartae (Karaman). In spite of this. the characteristics of this nominotypical subspecies are unquestionably described and detratsulli yb prahS )3881( dna esuohretaW.)09-2881( Host ants: Lasius (Chthonolasius) sp. (L. umbratus )rednalyN( ro L. rnericliom~1i.s :))tiordnoB( ,ikasim-ippaT iromoA .ferP tneserp(.)yduts Diartiger fossulatus izuinsulicola subsp. nov. .eluM Unknown, .elameF htgneL 52.2 .mm htdiW 58.0 .mm ydoB .dezis-egral eannetnA ,rednels 5.0 mml ni ,htgnel 11.0 mm ni ,htdiw ht4 tnemges ylthgils denedaorb .yllatsideerht semit sa gnol sa.ediw Basi-lateral trichome of composite tergum small, strongly deneprahs dna delruc ,ylroiretsop lagretarap emohcirt ,trohs lagretarap csid egraldna ,etagnole ylraen.mrofisuf ,epytoloH ,elamef uzoK ,.sI uzI ,.slsI oykoT ,.ferP .71 .iiv ,9891 .M awagesaH.gel Distribution of the subspecies. :napaJ uhsnoH uzI( .).s1~1 Remarks. sihT seicepsbus sreffid morf eht lacipytonimon seicepsbus ni ehtrednels eannetna dna eht deneprahs dna d ellarruect aelm-oihscaibrt fo eht etisopm.omcugret Host ants: Unknown. Diartiger fossulatus ispartae Karaman, stat. nov. CoifSaitius ispartue,namaraK ,9691.loiB ,acinelleH-ollaG 2: 50. Dim-tiger ispartae: ,tehcuseB ,6891 euveR essius ,.looZ :39 ;362 ,arumoN 1989, kcehC List .npJ,.snI Fukuoka, :1.392 .elaM htgneL 81.2-40.2 .mm htdiW 0.78-0.85mm. Body medium-sized. Antennae etagnole dna ,rednels 45.0-05.0 mm ni ,htgnel 1.0 31.0-l mm ni .htdiw ht4tnemges ylthgils denedaorb ,datsid 0.3-5.2 semit sa gnol sa .ediw diM sretnahcort ,etagnolehcae htiw a egral enips no lacipa-oretsop ,edis dim aromef ,tsubor hcae htiw owt yrevllams selcitned raen eht elddim fo lasrod anirac dna ta lasab 8/l fo lartnev anirac no roiretsop side. Trichomes on basi-lateral part of composite tergum similar to those in the lacipytonimon.seicepsbus .elameF htgneL 2.04-2.18mm. htdiW 0.76-0.85mm. eannetnA retrohs naht ni,elam 05.0-64.0 mm ni ,htgnel 1.0 31.0-l mm ni ,htdiw ht4 tnemges 7.2-3.2 semit sa gnolsa wide. Specimens examined. ]uhsnoH[ 2 ,selam 1 ,elamef ,ihcugawasoruK .tM.nasekatnO ,osiK onagaN ,.ferP .02 .iiv ,2591 .S oneU ;.gel 1 ,elamef ,inatebaN -ihcukonustaT ,ihcam awakihsI ,.ferP .5 .iv ,4991 .Y eiguS ;.gel 1 ,elam .tM ,amayakatirihS -igurusT ,ihcam awakihsI ,.ferP .61 .ix ,1991 .K atakaN ;.gel 1 ,elam ,ihcam-igurusT awakihsI ,.ferP .8 .v ,3991 .I ihsagoT ;.gel 1 ,elamef ,imakawaK ,ihcam-akouraM iukuF Pref., 25. v. 1984, M. 8tiaS ;.gel 2 ,selam 2 ,selamef ikunanaH ,ssaP .tM .arahagiadOeiM ,.ferP .21 .iv ,2591 .H adihsI ;.gel 2 ,selam 1 ,elamef ,emaS ,ohc-agaT agihS ..ferP.42 v. 1952, S. o&U ;.gel 1 ,elam ,arahagomihS ,8hc-agaT agihS ,.ferP .42 .v .2591.S NOISIVER FO DIARTIGER MORF TSAEAISA 83 oneU ;.gel 1 ,elamef ,arukawI otoyK ,.C otoyK ,.ferP .02 .vi ,1891 .T atagO ;.gel1 female, Kibune, Kyoto C., otoyK ,.ferP .92 .iiiv ,3491 .K itugakaS ;.gel 2 .selam 1 female, same locality, 6. v. 19.52, S. oneU ;.gel 1 ,elamef ,ijuy-neS Imakumano. ,uk-amayihsagiH otoyK ,.C otoyK ,.ferP .3 .v ,3491 .K itugakaS ;.gel 3 .selam.tM Kuramayama, Kyoto ,.C Kyoto Pref., 26. v. 1979, Y. Hirano leg.; 1 female, Totsukawa, ,8hc-ohuziM Kyoto Pref., 10. vi. 1986, S. Nomura leg.; 3 males, 6 females, Mt. Mikusayama, ,8hc-esoN akasO ,.ferP .71 .iix ,2991 .Y adawaS :.gel4 ,selam 5 ,selamef emas ,ytilacol .32 .I ,3991 .Y adawaS ;.gel 10 ,selam 31,selamef emas ,ytilacol .91 .iii ,3991 .Y adawaS ;.gel 1 ,elam 2 ,selamef emas ,ytilacol .02.vi 1993, Y. adawaS ;.gel 1 ,elam emas ,ytilacol .02 .xi ,3991 .Y adawaS ;.gel 1.elam ,ijustnamuJ Takarazuka C., ogoyH Pref., 25. vi. 1989, H. Ohishi leg.; 1 female. ,akazaraN araN ,.C araN ,.ferP .2 .iiv ,6991 .Y aruihsagiH ;.gel 3 ,selam 1.elamef Kashiwadani, Kii-Ohshima Is., ,8hc-otomihsuK Wakayama Pref., 5. v. 1994. S. Nomura leg.; 1 female, Mt. ,nazohsQyK irottoT ,.C irottoT ,.ferP .12 .v 1989. N. ikasurusT ;.gel 3 ,selam 3 ,selamef emas ,ytilacol .02 .iix ,1991 .N ikasurusT.gel Distribution of the subspecies. :napaJ uhsnoH ,ubuhC( ,ukirukoH ikniK stcirtsiddna irottoT .).ferP Remarks. This subspecies is easily distinguished in the male from the other seicepsbus yb yrev llams owt selcitned fo eht dim ,rumef hguoht eht elamef tonnaceb ylida edretarapes morf eht.srehto tsoH ants: Unknown. Diartigerfossulatus seicepsbusdenimretedni Specimens examined. ]uhsnoH[ 1 ,elam ,arawadO awaganaK ,.ferP .2 .i ,6791.Y onariH ;.gel 1 ,elam emas ytilacol sa ,evoba .31 .ii ,7791 .Y onariH ;.gel 1 .elam1 ,elamef emas ytilacol sa ,evoba .61 .iix ,8791 .Y onariH ;.gel 1 ,elam emas ytilacolsa ,evoba .6 .v ,4891 .Y onariH ;.gel 8 selam 2 ,selamef .tM ,nazQyiaD ,enokaHawaganaK ,.ferP .92 .vi ,5791 .Y onariH;.gel 2 ,selam 2 ,selamef emas ytilacol sa ,evoba .8.v ,6791 .Y onariH ;.gel 1 ,elam 3 ,selamef emas ytilacol sa .evoba .6 .v ,9791 .YonariH ;.gel 1 ,elam emas ytilacol sa ,evoba .9 .iiiv ,1891 .Y onariH ;.gel 1 ,elam emasytilacol sa ,evoba .72 .iv ,2891 .Y onariH ;.gel 1 ,elam emas ytilacol sa ,evoba .9 .vi ,3891.Y onariH ;.gel 1 ,elam 1 ,elamef emas ytilacol sa ,evoba .1 .v ,3891 .Y onariH ;.gel 1,elam .tM ,amayarihb .rn okakanamaY ,ekaL ihsanamaY ,.ferP .41 .iv ,6891 .Y onariH ;.gel2 ,elamef ,ikasuS uzI ,alusnineP akouzihS ,.ferP .2 .i ,4791 .Y onariH ;.gel 1 female, ,ariadnohiN uzimihS ,.C akouzihS ,.ferP .7 .v ,6991 .T akanoN.gel Remarks. nI na aera gnidnuorrus eht enokaH ,.stM nretsae trap fo .tM ijuF dnaeht uzI ,alusnineP htob smrof fo eht seicepsbus ispartae )namaraK( dna ehtlacipytonimon seicepsbus rucco ni a .noitalupop roF ,elpmaxe lacipyt smrof fo eht owt seicepsbusdna emos etaidemretni smrof ni suoirav seerged era dnuof no .tM ,nazuyiaD ,enokaHhcihw si esolc ot eht epyt ytilacol fo siht.seiceps esuaceB fo eht ,msihpromylopseicepsbus dluoc ton eb yl rdaeezlicngocer ni siht ,aera neve hguoht a wef selam er edweifitnedhitiw eno .mrof sihT aera si nwohs yb nepo selcric ni .giF.31 84 .SARUMON Diartiger fossulatus morimotoi subsp. nov. h4aEe. htgneL 2.08-2.15 .mm htdiW 0.78-0.83 mm. Body medium-sized. Antennae etagnole dna ,rednels 5.0 45.O-l mm ni ,htgnel 11.0 mm ni ,htdiw ht4 tnemgesylthgils denedaorb ,datsid 8.2-7.2 semit sa gnol sa .ediw diM sretnahcort ,etagnole hcae htiwa egral dna yleuqilbo etacnurt enips no lacipa-oretsop ,edis dim aromef ylkaewnellows ,dacipa hcae htiw a ,egral etuca dna yllacipa devruc elcitned raen eht elddim noroiretsop .edis laretal-isaB emohcirt fo etisopmoc mugret daorb ta ,esab deneprahs drawot.xepa ylkaew delruc ,dairetsop lagretarap emohcirt ,egral suoirav ni .epahs lagretarap disc y.ldriaoevno Female. Length 2.09-2.24 mm. Width 0.79-0.84 mm. Antennae shorter than in ,elam 15.0-94.0 mm ni ,htgnel 1.0 31.0-l mm ni ,htdiw ht4 tnemges 7.2-3.2 semitsa gnol sa.ediw ,epytoloH ,elam .tM ,naziK ,ohc-amayiK agaS ,.ferP .21 .vi ,2991 .S arumoN.gel ,sepytopotaraP 2 ,selam 2 ,selamef emas atad sa ;epytoloh 1 .elam 2 ,selamefemas ,ytilacol .11 .vi ,3991 .S arumoN.gel ,sepytaraP 1 ,elam 3 ,selamef.arahakustanaN arabohS C., Hiroshima Pref., 10. x. 1987, I. Okamoto leg.: 1 male, 1 female. ,arabatawaY ,8hc-ukohieG amihsoriH ,.ferP .72 .iv ,7891 .S arumoN ;.gel 9 .selam 1 1 females, ,8hc-esoruK Kamo-gun, Hiroshima Pref., 2. v. 1988, I. Okamoto leg.; 1 ,elamef emas ,ytilacol .92 .v ,8891 .I otomakO ;.gel 2 ,selam 1 ,elamef.awag-esoruK eruK ,.C amihsoriH ,.ferP .6 .ii ,8891 .I otomakO ;.gel 1 ,elamef .tM .nasebnaSathO ,.C enamihS ,.ferP .2 .iv ,9891 .K atagO ;.gel 1 ,elamef imikih-arU ,.llaV ,ohc-imikiH enamihS ,.ferP .6 .iv ,8891 .K orihaY ;.gel 1 ,elamef .tM .nasijnamiaD .ohc-ogiaS ikO ,.sI emamihS ,.ferP .51 .iv ,8891 .S iukuF ;.gel 2 ,selam ,odihsoyikA ihcugamaY,.ferP .82 .v ,7791 .S akanaT ;.gel 5 ,selam 5 ,selamef ,oguod-ukO amayustaM ..CemihE ,.ferP .12 .iiiv ,6991 .I otomakO ;.gel 1 ,elamef ,ijoygnE ih&K ,.C ihcoK ..ferP .92.iii ,3591 .K otomiroM ;.gel 1 ,elamef ,iadoariH ihsahukuY ,.C akoukuF ,.ferP .81 .x ,2991 .S arumoN;.gel 1 ,elam ,inatoriH ,iadoariH uhsuykatiK ,.C akoukuF ,.ferP.52 .vi ,9891 .S arumoN;.gel 1 ,elamef .tM ,amayikuzakiM akoukuF ,.C akoukuF,.ferP .22 .i ,3991 .S arumoN;.gel 1 ,elam ,neek-imaniM akoukuF ,.C akoukuF ..ferP.91 .iiiv ,3891 .H adaraH ;.gel 1 ,elam ,ikazenaK ,ihcam-iakneG akoukuF ..ferP .6.xi 1983, H. Harada leg.; 2 males, 2 females, Hikosan skiing ground, ,ihcam-adeoS akoukuF ,.ferP .7 .iv ,3991 .S arumoN ;.gel 2 ,selam 1 ,elamef ,ekI .ihcam-ukatihsiN ukaT ,.C agaS ,.ferP .12 .iii ,9891 .S arumoN;.gel 1 ,elamef .tM ,amayenufiMoekaT ,.C .72 .v ,4891 .S arumoN ;.gel 1 ,elam ,urabihsaK ,arum-amayanaN agaS ..ferP .9.iv ,5891 .S arumoN ;.gel 2 ,selam 5 ,selamef emas ,ytilacol .41 .v ,9891 .S arumoN ;.gel2 ,selamef ,urabekI ,arum-amayanaN agaS ,.ferP .01 .iv ,0991 .S arumoN :.gel 1.elam Mt. Taradake, ,8hc-araT agaS ,.ferP .41 .vi ,5891 .S arumoN ;.gel 1 ,elam,o-onaT arumhO ,.C ikasagaN ,.ferP .51 .iii ,4991 .S akasamI ;.gel 1 ,elamef ,ikatanaN -enufiM ,8hc otomamuK ,.ferP .52 .iv ,4991 .S arumoN ;.gel 1 ,elam ,nijiadiaN Yabe-ch8, Kumamoto Pref., 28. vii. 1985, R. adoN leg.; 1 male, Mt. Oidake. ,ohc-ekairA Amakusa, Kumamoto Pref., 3. vii. 1994, S. Nomura leg.; 1 female. ,o-onasaS ,8hc-asukamA ,asukamA otomamuK ,.ferP .32 .iix ,1991 .S arumoN :.gel 3 ,selam6 females, ,urustagbB itjuK Mts., atinA ,.ferP .72 .v ,8891 .K atagO ;.gel 1 :elamef ,urabajohC ifjuK ,.stM at& ,.ferP .11 .iiiv ,2991 .S arumoN ;.gel 1 ,elam .tM.eukhO ,ahc-awagatiK Miyazaki Pref., 8. x. 1984, S. Nomura leg.; 1 male, 2 females. NOISIVER FO DIARTZGER MORF TSAEAISA 8.5 ,ohihcakaT ikazayiM ,.ferP ,5 .iv ,5891 .Y iakaT ;.gel 1 ,elamef emas ytilacol sa.evoba .72 .ix ,9791 .S akanaT.gel Distribution of the subspecies. :napaJ uhsnoH ukoguhC( tcirtsid gnidulcxe irottoT Pref.) including ikO Isls., Shikoku, northern Kyushu excluding the Shimabara .alusnineP Remarks. The new subspecies morimotoi is different from the nomynotypical seicepsbus ni eht rednels ,eannetna eht yleuqilb oetacnurt enips fo eht dim .retnahcortdna eht yllacipa de nleaprreatha sle-mioshacbirt fo eht etisopmo.cmugret Etymology. This subspecies is dedicated to Prof. Emer. Katsura Morimoto of Kyushu University for his great contribution to the Japanese coleopterology and s ueocunnaidtin uorgcof ym ydut secnis I saw a tneduts fo.sih Host ants: Lasius (L&us) productus :nosliW ,arabatawaY amihsoriH .ferP tneserp(.)yduts Lcrsius (Lasius) niger :)sueanniL( nasokiH gniiks ,dnuorg akoukuF .ferP tneserp(.)yduts Lasius (L&us) hayashi ihcuamaY te :adihsayaH ,nijiadiaN otomamuK .ferPtneserp( study). Diartiger fossulatus imasakai subsp. .von Male. Length 1.93-2.00 mm. Width 0.73-0.76 mm. Body small-sized. Antennae etagnole dna ,rednels 0.50-0.5 1 mm ni ,htg n.eol-01.0 31 mm ni ,htdiw ht4 tnemgesyrev ylthgils denedaorb ,datsid 1.3-8.2 semit sa gnol sa .ediw diM sretnahcort .etagnolehcae htiw a egral dna etacnurt enips no lacipa-oretsop ,edis dim aromef ylthgilsnellows ,dacipa hcae htiw a egral dna etuca elcitned ta lasab 3/l no roiretsop .edislaretal-isaB emohcirt fo etisopmoc mugret ylgnorts deworran dna deneprahs drawot ,xepaylgnorts delruc ,daire tlsaogpretar aepmohcirt egral ta ,esab lagretarap csid ,llams ylraen.ralugnairt Female. htgneL 69.1-19.1 .mm htdiW 57.0-47.0 .mm eannetnA 84.0-64.0 mm ni ,htgnel .o-01.0 31 mm ni ,htdiw ht4 tnemges 1.3-9.2 semit sa gnol sa.ediw Holotype, male, amay-nekoT ,kraP ,6hc-amabO ikasagaN ,.ferP .91 .xi ,3991.S akasamI .gel ,sepytopotaraP 1 ,elam emas atad sa ;epytoloh 1 .elam emas .ytilacol .5.vi ,4991 .S akasamI .gel ,sepytaraP 1 ,elam arabamihS ,.C ikasagaN ..ferP .52 .x.5891 .T ebanaT ;.gel 2 ,selamef ,ukogijoK ,neznU ,ohc-amabO ikasagaN ,.ferP .71 .iiiv ,3891 .H adaraH.gel Distribution of the subspecies. :napaJ nrehtroN uhsuyK arabamihS(.)alusnineP Remarks. sihT wen seicepsbus si deziretcarahc yb eht llams ydob dna ehtdeworran dna ylgnorts delruc laretal-isa bemohcirt fo eht etisopmoc mugret ni .elam .revewoHeht elamef si tluciffid ot eb dehsiugnitsid morf eht rehto ,seicepsbus esuaceb ehtretcarahc etats fo elhatre teamlo-hicsiarbt si se.mtictneimtossidni Etymology. sihT seicepsbus eman si detaicossa htiw .rM ihciohS akasam Iohw sina ttsn iergluolele tocdamxnmoeaatne si erhottcell ofco e.hetpytoloh Host ants: Unknown.
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