A new genus and new species of Colpurini (Heteroptera: Coreidae) from New Guinea

11 Pages·2003·3.3 MB·

by  Harry Brailovsky

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PROC. ENTOMOL.SOC.WASH.
105(2). 2003,pp.362-372
A NEW GENUS AND NEW SPECIES OF COLPURINI
(HETEROPTERA: COREIDAE) FROM NEW GUINEA
Harry Brailovsk^' and Ernesto Barrera
Departamento de Zoologia, Instituto de Biologia, UNAM, Apdo Postal No. 70153,
Mexico 04510, D.F., Mexico (e-mail: [email protected])
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Abstract. The new genus Ullrihygia and five new species Ullrihygia iriana.Heisshy-
gia tafa. Lothygia sordida. Lothygia coitsocia, and Sciopliyroides splendidiila collected
in New Guineaaredescribed inthe tribeColpurini (Coreidae). Dorsalhabitusillustrations
and drawings ofthe head, male genital capsule, and female genital plates are providedto
distinguish each species. A key to the Lothygia species is given.
Key Words: Insecta, Heteroptera, Coreidae, Colpurini, new genus, new species. New
Guinea
The island ofNew Guineacontains a re- many); ZSM (Zoologische Staatssa-
markable and highly endemic assemblage mmlung, Miinchen, Germany).
ofgeneraandspecies in thetribeColpurini, All measurements are given in millime-
many ofwhich are still undescribed. ters.
The tribe Colpurini is represented by a
numberofspeciesthatshowvariousbizarre Ullrihygia Brailovsky and Barrera,
moiphological specializations. The most new genus
striking features ofthese genera are the re- Diagnosis. Ullrihygia isrelatedtoBau-
markable reduction of the hemelytra, the mannhygia Brailovsky (1996), Nishihygia
developmentofthe head includingtheeyes Brailovsky(2000),and WygohygiaBrailov-
and tylus, and the shape of male genital sky (1993a). Similar to Baiinuinnhygia
capsule and female genital plates (Brailov- withoutocelli,headquadrangularandwider
sky 1996, 2000). than long, antenniferous tubercle armed
The current paper is a further contribu- withlongrobustspine,eyessmallandglob-
tion tothis study. Provided aredescriptions ular, buccula with sharp projection, femora
foronenewgenusandfivenew speciesand unarmed, micropterous, abdomen elliptical,
a key to the species belonging to the genus and connexivum higher than terga. Baii-
Lothygia Brailovsky. tnannhygia known only from the Philippine
The following abbreviations are uAseMdNfHor Republic, is recognized by having the tylus
the institutions cited in this paper: unarmed, the postocular tubercle protuber-
(TheAmerican MuseumofNatural History, ant, antennal segment 1 longer than 111 and
New York): BPBM (Bernice R BishopMu- shorter than maximum length of the head,
seum, Honolulu, Hawaii); BMNH (The humeral angles rounded and not raised,and
Natural History Museum, London. Eng- scutellum apically acute. In Ullrihygia. re-
land): UNAM (Instituto de Biologia. Univ- corded from Irian Jaya. the tylus isapically
ersidad Nacional Autonoma de Mexico): acute and upturned to form a median horn,
WUCO
(Wolfgang Ullrich collection, Ger- the postoculartubercle is absent, the length
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VOLUME lOS. NUMBER2 363
ofantennal segment I is equal than III. and Thorax: Pronotum wider than long, tra-
equal than maximum length of head, the peziform, bilobed. and non declivent; an-
humeral angles are raised, and the scutel- terior lobe slightly longer than posterior
lum is apically rounded. This new genus lobe, and each margin convexly rounded;
resembles Nishi/iygia in having the tylus collar weakly apparent; frontal angles pro-
apically acute, antenniferous tubercle duced forward asconical lobe; humeralan-
armed,ocelliabsent,eyesglobular,humeral gles rounded, raised, and directed upward;
angles slightly raised, femoraunarmed, mi- posterolateral borders and posterior border
cropterous, apex ofscutellum rounded, ab- straight; calli weakly convex. Anteriorlobe
domen elliptical, and connexivum higher of metathoracic peritreme reniform. poste-
than terga. In Nisliihygici. the frontal angles rior lobe sharp, small.
are rounded and not exposed, the posterior Legs: Unarmed; tibiae cylindrical and
lobe ofpronotum depressed, postocular tu- sulcate.
bercle moderately protuberant, and buccula Sciifelliiin: Wider than long. flat, mod-
rounded, without sharp acute projection. erately rectangular; apex rounded.
Wygohygia is like Ullrihygia with the eyes Heiiielytron: Micropterous. reaching an-
small and protrudent on short stalks, head terior third ofabdominal segment I; wings
quadrate,bucculawith sharpprojection,an- reduced to small pads, widely separated;
tenniferous tubercle armed, and frontal an- clavus and corium fused, and abdomen ex-
gles produced forward as rounded lobes. posed mesially; membrane absent.
Wygohygia is recognized by the following Abdomen: Conspicuously elliptical;con-
combination ofcharacters; headlongerthan nexivum higherthan terga;connexivalmar-
wide, tylus apically globose, ocelli present, gin entire, with posteriorangleofeachseg-
postocular tubercle protuberant, femora ment not extending on a short spine; terga
armed, apex of scutellum acute, and sub- II to VI barely convex.
macropterousreachingthe posteriorthirdof Integument: Body surface dull, almost
abdominal segment VI. glabrous; antennal segments and legs with
Nisliihygici and Wygohygia are recorded long erect setae; head with circular,grayish
from New Guinea. white farinose punctures; pronotum, scutel-
Generic description. Head: Widerthan lum, hemelytron, abdominal segments, tho-
long, quadrangular, and dorsally slightly rax, and abdominal sterna strongly punc-
convex; tylus apically upturned to form a tate;connexivumanduppermarginofpleu-
subacute median horn, extending anteriorly ral abdominal sterna smooth.
to and laterally higher than juga; juga un- Male genitidia: Posteroventral edge of
armed,thickened,andapicallybarelyacute; genital capsule strongly produced forward
antenniferous tubercle armed with long ro- as robust conical projection (Figs. 7-8).
bust spine; antennal segment I stout, thick- Female, unk—nown.
est, slightly curved outward and equal than Etymology. NamedafterWolfgangUll-
maximum length of head; segments II and rich, a distinguished German entomologist.
III cylindrical and slender; segment IV fu- Gender feminine.
siform; antennal segment II longest, IV Type species. Ullrihygia iriana Brai-
shortest, and I equal to III; ocelli absent; lovsky. new species.
preocellar pit deep, circular; eye small,
globular,andprotrudingon shortstalk;pos- Ullrihygia iriana
tocular tubercle absent; buccula rounded, Brailovsky and Barrera, new species
short, not projecting beyond antenniferous (Figs. 7-8. 18)
tubercle, with sharp middle projection; ros- Description. Measurements: Male:
trumreachinganteriormarginofabdominal Head length 1.48; width across eyes 1.84;
stemite III; mandibular plate absent. interocular space 1.32; preocular distance
PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON
Figs. 1-8. 1-6. Lolhygia spp.. male {icnital capsule in caudal view. 1. L. papiuiiui. 2. L. cordigera. 3. L.
enarotadi. 4, L. consocia. fi. L. surdidii. (n. L. saiidaracinc Brail(i\sky. 7-8, Ulliihxfiici iriaiHi, male genital
capsule. 7.Caudal view. 8. Lateral view.
1.34; length antennal segments: 1, 1.48; II, terolateral margin and upper margin ofhu-
2.02; III, 1.48; IV, 1.08. Pronotum: Total meral angles ochre orange; scutellum red-
length ofanterior lobe 0.94; total length of dish brown with wide ochre-orange longi-
posterior lobe 0.89; total width of anterior tudinal stripe at middle third; hemelytron
lobe2.02;totalwidthofposteriorlobe2.92. and abdominal segments reddish brown;
Sciitellar length 0.40; width 1.32. Maxi- connexivum reddish brown with posterior
mum width ofabdomen 4.80. Body length margin of connexival segments V and VI
8.95. dark orange. Ventral coloration: Head
Dorsal colonition: Head black; antennal black; rostral segmentsdarkyellow;thorax,
segments I to III reddish brown, with basal abdominal sterna, and genital capsule dark
joint of III ochre orange; antennal segment orange yellow with punctures reddish
IV dark chestnut orange with basal third brown to black; coxae black; trochanters
reddish brown; anterior lobe of pronotum yellow; femorayellow withapical thirdand
blackandposteriorlobereddishbrown; an- few and scattered discoidal spots reddish
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VOLUME 103. NUMBER2 365
brown; tibiae reddish brown with two yel- Genital capsule: Pt)steroventral edge
"V
low rings one subbasal. the other one near with open and deep concavity, en-
middle third; tars—i chestnut orange. closed by two broad rounded lobes (Fig.
Type material. Holotype: d, Indonesia: 14).
Irian Jaya, Paniai Province. N Mulia Trail Coloration: Ground color bright reddish
to Dowome. 2.200-2.500 m. 8 July 1994, orange with headdarker;antennalsegments
A. Riedel (Ent. Expl. New Guinea, 1990- ItoIII brightreddishorange,andIVorange
1994) (WUCO). Paratype: 1 6. same label yellow with basal third bright reddish or-
data as holoty—pe (UNAM). ange; apex ofscutellum dark yellow; upper
Etymology. The name is derived from marginofconnexivumandrostralsegments
the type locality. 1 to IV pale chestnut t)range; posteriormar-
gin of acetabulae yellow; coxae, trochan-
Heisshygia tafa Brailovsky and Barrera, ters, and fore and middle femora pale or-
new species ange yellow; hind femur dark chestnut or-
(Figs. 14. 16) ange with two yellow rings, one basal, the
Description. Measurements: Male: other one near middle third; tibiae pale
Head length 1.60; width across eyes 1.56; chestnutorange with two yellow rings, one
interocular space 0.94; interocellar space subbasal, the other one near middle third;
0.32; preoculardistance 1.04; length anten- tarsi chestnutorange;anteriorlobeofmeta-
nal segments: I, 1.20; II, 1.80; III, 1.24; IV. thoracic peritreme pale yellow, and poste-
0.88. Pronotum; Length 1.32; width across rit)r lobe reddish orange.
frontal angles 1.60; width across humeral Female: Headas longeraswide; rostrum
angles 2.76. Scutellar length 0.88; width reaching posterior third of abdominal ster-
0.96. Maximum width of abdomen 4.39. nite V. Coloration: similartomale.Connex-
Body length 8.85. Female: Head length ival segments VIII and IX. dorsal abdomi-
1.60; width across eyes 1.60; interocular nal segmentsVIIIand IX.andgenitalplates
space 0.96; interocellar space 0.34; preo- bright reddish or—ange.
cular distance 1.09; length antennal seg- Type material. Holotype 6. PapuaNew
ments: I. 1.28; II, 2.00; III, 1.40; IV. 0.92. Guinea: Mt.Tafa, 8,500', February 1934,L.
Pronotum; Length 1.60; width across fron- E. Cheesman (BMNH). Paratypes: I d. I
talangles 1.68;widthacrosshumeralangles 9, same data as for holotype (BMNH,
2.76. Scutellar length 0.84; width 1.06. UNAM). —
Maximum width of abdomen 5.30. Body Discussion. Among New Guinea Heis-
length 10.00. shygia species (Brailovsky 1995), H. tafa,
Male: Head: Pentagonal, longer than is superflcially most similar to H. novogui-
wide; antenniferous tubercle barely diverg- nensis Brailovsky (1993b), but may be sep-
ing anteriorly; ocelli obscure; rostrum arated by the following characters: In H.
reaching anterior border ofabdominal ster- tafa, the posteroventral edge of the male
nite VI (Fig. 16). genital capsule has an open and deep V-
Thorax: Pronotal disc flat, with posterior shaped concavity, enclosed by two broad
median depression. rounded lobes (Fig. 14), the rostrum reach
Scutelliim: Wider than long, with apex the posterior third ofabdominal sternite V
rounded. or anterior border of VI, the antenniferous
Hemelytron: Micropterous, reaching tubercle are unarmed, and the hemelytra
posteriorthirdofabdominal segment I;cla- reaching the posterior third of abdominal
vus and corium fused into coriaceous pad, segment I. In H. novoguinensis (Fig. 17),
and wings widely separated from each oth- theposteroventraledgeofmalegenitalcap-
er, leaving abdomen totally exposed mes- sule has two short lateral projections, sur-
ally; membrane represented by small flap. roundingabroadmiddleplate(Fig. 15),the
PROCEEDINCiSOI-TUBENTOMOLOGICALSOCIETYOFWASHINGTON
Figs. 9-17. ^)-\}. S,ioi)/ivri>iJcs spp. '> \\. M.ilc gL-niliil capsule in caudiil \ic\v. '), .V splcndidula. U). i".
tuumiualus. \\.S.piiiiiilns. 12-L^.Femalegenitalplatesin lateral \iew. 12.S. aviiinimiliis. \?.S. splcndiiliihi.
14-17,Heisshygiaspp. 14-15.Malegenitaleap.suleineaudalview. 14.H. uifa. \f>. H. iu>riii;iiiiun.sis, 16-17,
Headindorsalview. 16.H. tafa. 17,H. novoguineiisis.
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VOLUMI-; 105, NLIMBliR 367
Fig. IX. Dorsiil view cil lllhihyKiaiiiaiia. iiiiile
rostrum reach the anterior third of abdom- Lothyi^ia sordida
inal sternite V, the antenniferoustubercle is Brailovsky and Barrera, new species
armed, and each lobe raised and diverging (Figs, 3. 19)
anteriorly, and the hemelytra reaching the Description, McasnrcnwiUs: Male:
posterior third—ofabdominal segment II. Head length 1,96; width across eyes 1.84;
Etymology. This species is named for interocular space 1.04; interocellar space
its occurrence in the Tafa mountains. 0.48; preoculardistance 1.28; length anten-
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PROCEEDINGSOFTHEENTOMOLOGICALSOCIETYOFWASHINGTON
nal segmems: I, 1.40; II, 2.08; III, 1.36; IV. lobe black; coxae, trochanters and femora
1.42. Pronotum: Length 2.12; width across pale reddish brown; tibiae and tarsi pale
frontal angles 1.56; width across humeral chestnut orange.
angles 3.80. Scutellar length 1.48; width Female; Genitalia: Gonocoxae I en-
1.60. Body length 10.95. Female; Head larged dorso-ventrally, in caudal view
length 2.16; width across eyes 2.00; inter- closed, in lateral view barely convex, with
ocular space 1.16; interocellar space 0.52; upper border almost straight. Coloration;
preocular distance 1.48; length antennal similar to male. Rostral segments I and II
segments; I. 1.52; II, 2.16; III, 1.40; IV, dark chestnut orange and III and IV pale
1.44. Pronotum: Length 2.44; width across chestnut orange; legs bright pale reddish
frontal angles 1.68; width across humeral brown with subbasal yellow ring at tibiae;
angles 4.16. Scutellar length 1.64; width connexival segments VIII and IX black
1.96. Body length 12.30. with anterior third dark yellow; abdominal
Male; Head: Longer than wide; tylus segmentsVIIIand IX darkbrown withdark
apically truncated; antenniferous tubercle yellow reflections; gonocoxae I bright red-
with small lobelike projection; eyes spher- dish brown with uppermargin darkyellow;
ical, elevated, and protruding on short paratergite VIII and IX bright reddish
stalks; postocular tubercle protuberant, and brown. —
acutely expanded; rostrum reaching middle Type material. Holotype 6, Indonesia;
third ofabdominal sternite VII. Irian Jaya. Wamena Pass Valley, 1,900 m,
Thorax: Pronotumwiderthanlong;fron- 15-16 September 1990. A. Riedel (ZSM).
tal angles produced forward as conical Paratypes: 1 6. Indonesia: Irian Jaya, Wa-
teeth;humeral anglesrounded,notexposed. mena Llugwa Melanggama. 1.800-2.000
Heinelytron: Macropterous, reaching m. 9-12 September 1990, A. Riedel
apex oflast abdominal segment. (UNAM). 1 S. \ ?, Indonesia: Irian Jaya,
Genitalcapsule: Posteroventraledgelat- Wamena Pass Valley, 1,900 m, 15-16 Sep-
erally produced on a short blunt lobes, and tember 1990.—A. Riedel (ZSM).
between them a deep "U" concavity (Fig. Discussion. This species is diagnosed
5). mostly on basis ofthe shape ofthe poster-
Dorsal coloration: Head shiny black oventral edgeofmalegenital capsule(Figs.
with tylus bright reddish brown; antennal 1-6), the presence of an acutely produced
segment I bright reddish brown; segments postoculartubercle, andby the shapeofthe
II to IV dark chestnut orange; dorsal face eyes that are spherical, elevated, and pro-
ofpostoculartubercle dark yellow; anterior truding on a short stalks. In L. cordigera
lobe of pronotum black, and posterior lobe Brailovsky (1994), the postocular tubercle
dark reddish brown; scutellum dark reddish is not acutely produced, theeyes are spher-
brown; clavus and corium pale reddish ical and weakly protrudent, and the poster-
brown; hemelytral membrane dark brown; oventral edge of male genital capsule is
connexivum pale orange yellow with pos- heart-shape (F—ig. 2).
terior third or posterior borderofsegments Etymology. From the Latin sordidus,
III to VII black; abdominal segments III to meaning dirty and referring to the blackish
VI dark orange and VII dark orange with color.
extensive black marks. Ventral coloration:
Head black; rostral segments dark to pale Lothygia consocia
chestnut orange; thorax, abdominal sterna Brailovsky and Barrera, new species
and genital capsule bright reddish brown (Fig. 4)
with pleural abdominal segments III to VII Description. Measurements: Male:
pale orange yellow; anterior lobe of meta- Head length 1.80; width across eyes 1.64;
thoracic peritreme yellow and posterior interocular space 0.96; interocellar space
VOLLIME 105. NUMBER2
Fig. 19. DiirsalviewofLorhyaiasonlhla.
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.
PROCEEDINGSOHTHEENTOMOLOGICALSOCIETYOFWASHINGTON
0.48; preoculardistance 1.20: length anten- Female: Tylus apically with small spiny
nal segments: I, 1.36; II, 1.76; III. 1.16; IV, projection; hemelytra submacropterous,
1.36. Pronotum: Length 2.04; width across reaching posterior margin of abdominal
frontal angles 1.60; width across humeral segment VII. Genitalia. Gonocoxae I en-
angles 3.36. Scutellar length 1.36; width larged dorso-ventrally, in caudal view
1.64. Body length 10.00. Female: Head closed, in lateral view barely convex, with
length 1.92; width across eyes 1.72; inter- upper border almost straight. Coloration:
ocular space 0.98; interocellar space 0.47; similar to the male. Upper margin of me-
preocular distance 1.36; length antennal tapleuron with orange yellow rectangular
segments: I. 1.40; II, 1.88; III, 1.24; IV, spot close to middle third; connexival seg-
1.36. Pronotum: Length 2.24; width across ments VIII and IXorangeyellow,withpos-
frontal angles 1.60; width across humeral terior border black; abdominal segments
angles 3.52. Scutellar length 1.48; width VIII and IX dark orange; gonocoxae I
1.64. Body length 11.30. bright reddish brown with internal angle
Male: Head: Longer than wide; tylus and apical margin yellow; paratergite VIII
apicallytruncate;antenniferoustubercleun- and IX dark chestnut orange, with basal
armed, with apex truncate; eyes spherical, third black. —
and barely protrudent; rostrum reaching Type material. Holotype i. Indonesia:
middle thirdofabdominal sterniteVII;pos- Irian Jaya, Paniai Province, S Mulia Trail
tocular tubercle conically protuberant. to Wuyuneen, 1,900-2,000 m, 6-7 July
Thorax: Pronotumwiderthan long;fron- 1994, A. Riedel (Ent. Expl. New Guinea,
tal angles produced forward as conical 1990-1994) (WUCO). Paratypes: I 9,
teeth;humeralanglesrounded,notexposed. same data as for holotype (UNAM). 1 9,
Henielytron: Submacropterous, reaching Indonesia: Irian Jaya, Paniai Province, N
middle third ofabdominal sternite VII. Mulia trail to Dowome, 2,200-2,500 m. 8
Genitalcapsule: Posteroventraledge lat- July 1994, A. Riedel (Ent. Expl. NewGuin-
erally produced in a short blunt lobes, and ea, 1990-1994) (WUCO).
between them with large concave plate Discussion. Lotliygia consocia. is easi-
(Fig. 4). ly recognized by the shape of the poster-
Dorsal coloration: Head black; tylus, oventral edge ofmale genital capsule (Fig.
andjuga shiny dark reddish brown; anten- 4), and by the submacropterous condition
nal segments dark chestnut orange; dorsal of the hemelytra which reach the middle
surface ofpostocular tubercle dark yellow; third ofabdominal sternite VII. On the pre-
anterior lobe of pronotum black, and pos- vious known species ofLotliygia. the hem-
terior lobe and scutellum dark reddish elytra exhibit a macropterous condition,
brown; clavus and corium pale reddish reaching or extending beyond the apex of
bwirtohwn;veihnesmelayntdralbasmaelmbanrgalneeddaarrkkobrraonwgne,; lastEtaybmdoolmoignya.l—sFergmoemntthe(Blraatiinlocvosnksyoci19u9s4,).a
connexivum bright orange yellow, with companion: one of several species occur-
posteriorthirdofsegments III toVII black; ring sympatrically in Irian Jaya.
abdominal segments orange. Ventral col-
oration: Head black; rostral segments Key to Lothygia Species
bright chestnut; thorax, abdominal sterna, (Modified from Brailovsky 1994)
and genital capsule bright reddish brown
with pleural margin ofabdominal sternaIII 1 AbdominalsegmentsItoVIblack;gonocoxae
otof VmIeItatbhroigrhatciocrapnegreitryeemleloywe;llaontwerainodr ploosb-e - gAIobindnooccmaoiuxndaaaellIvsiieengwmceaonuptdeasnleIdviteowVcIloosreadnge yellow: 32
terior lobe black; legs reddish brown, with 2. Tylus apically with a short spiny projection:
dark chestnut orange reflections at femora. posteroventral edge of male genital capsule
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VOLUME 105, NUMBER2
with short iiicdiun phite (Fig. 1) shortest, and I longer than III; mandibular
L.iHipiiiinaBrailovsky plate armed with sharp and small tubercle,
- Tylusapicallyglobose,truncate;posteroventral
edgeofmalegenitalcapsulewithlargemedian visible in dorsal view; postocular tubercle
plate(Fig. 3) L. enarnladiBrailovsky protuberant: rostrum reaching middle third
3. Frontal angles poorly developed; malegenital ofabdominal sternite V.
capsule with posteroventral edge sinuate, and Thorax: Pronotum widerthanlong;fron-
laterally barelyexposed (Fig. 6) tal angles produced forward as small coni-
- Frontal angles well deveLl.opseadn,darparcoidnueceBdratiolr-ovsky cal teeth; humeral angles rounded, not ex-
wardasconicalexpansions 4 posed.
4. Submacropterous,reachingmiddlethirdofab- Legs: Femora unarmed, and ventrally
dominal sternite VII; posteroventral edge of with tiny denticles; tibiae cylindrical and
male genital capsule with large concave plate sulcate.
- (MFaicgr.op4)terous, reaching orL.exctoennsdoicniganbeeywosnpdecies Scutellum: Triangular, flat, and conspic-
theapexoflastabdominal segment;posterov- uously longer than wide; apex subacute.
entral edge of male genital capsule without Hemelytron: Macropterous, reaching
largeconcaveplate(Figs. 1.5) 5 posteriormarginoflastabdominalsegment.
5. Eyes spherical, elevated, and protruding on a Genitalcapsule: Posteroventral edge tri-
cshaoprstulsetalwki;thpodseteerpov"eUnt"racloneLcd.agvesiotoryiflim(dFaail.ge.ng5ee)wni.tsa.ple.cies fluorbceatteh,eelaocngheslto,bedirroebcutesdt uapnwdawriddea;ndcebnatcrka-l
- Eyes spherical, weakly protrudent; posterov- ward, and lateral lobes remarkably folded
entral edge of male genital capsule heart- downward (Fig. 9).
shaped(Fig. 1) I., cordiiiera Brailovsky Dorsal coloration: Head dark reddish
brt)wn with the space between ocelli and
Sciophyroides splendidula
eye, and dorsal view ofpostoculartubercle
Brailovsky an(dFigBsa.rr9.er1r3a), new .species dchaersktnyuetlloorw;anagnetewnintahlbsaesgamlejnotisntIotofIIIIIIdyaerlk-
Description. Measiiremeiils: Male: low; antennal segment IV dark chestnut to
Head length 2.79; width across eyes 2.17; darkyellow withbasal andapical thirddark
interocLilar space 1.14; interocellar space reddish brown; pronotum, scutellum, cla-
0.48; preoculardistance 1.88: length anten- vus, and corium pale reddish brown with
nal .segments: I, 3.28; II, 4.52; III. 2.74; IV. external marginofhumeral anglesandapex
1.98. Pronotum: Length 3.59; width across of scutellum yellow; middle third of calli
frontal angles 2.04; width across humeral dark reddish brown; hemelytral membrane
angles 5.64. Scutellar length 3.10; width dark ambarine; connexivum dark reddish
2.48. Body length 18.38. Female: Head brown with posteriormargin yellow;dorsal
length 2.82; width across eyes 2.23; inter- abdoininal segments bright orange with
ocular space 1.20; interocellar space 0.53; posteriormarginofVIIdarkreddishbrown.
preocular distance 1.82; length antennal Ventralcoloration: Ground colordarkyel-
segments: I. 3.03; II. 4.27; III. 2.66; IV. low with punctures dark to pale reddish
2.04. Pronotum: Length 3.53; width across brown; rostral segments I and II darkchest-
frontal angles 2.10: width across humeral nutorange. III palechestnutorange,andIV
angles 6.07. Scutellar length 3.34: width dark yellow; anterior lobe of metathoracic
2.66. Body length 19.43. peritreme yellow and posterior lobe dark
Male: Head: Elongate, longer than reddish brown; coxae and trochanters red-
wide, and dorsally flat; tylus projecting an- dish brown; femora reddish brown with ir-
teriad ofjuga, upturned to form sharp bifid regular yellow ring close to middle third;
horn; antenniferous tubercleunarmed;ocel- tibiae reddish brown withtwoyellowrings,
li with large diameter, and clearly tubercu- one subbasal, the otherone close to middle
late; antennal segment IIthe longest, IV the third; tarsi pale chestnutorange; pleuralab-