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© ZoolIongsitciSattlu. tPee,t2 e0r0s2b urg, A ngeewnu ansd n eswp eocfCi oecsc friodmCa een Amtrearilc a (HomoCpotcecr ian:e a) M.WillCihaH.mo sd,g &s oEn. DMa.n zig Wi�lmlsiM,. H,o dgsCoh_&n. D,a nzEi.gM2,.0 0A2n .e gwe naunsnd e swp ecoifes Coccfr1odmCa een tArmaeln( cHao mopCtoecrcaiZo:no esay)s.t eRmoasts, i 1ic1ca(a 1): 111-126. An egwe noufss o sftc ale, PrWiiolnloiHcaoomdcsgc&,su D osan n ziisg ,i nttrooduced tatkweno e swp ecPia egasWv,ie l liams&,D aHnoazdnigPdga s moenr iWcainlulsi ams, Hodg&sD oann zfriogmC enAtmrearlTi hcaead .ufe lmta loefbs o atrhde e scraisab reed thles t�t-aairnn2 dn d-imnaslatenfa der m aanlpdeu poatf h lea tstpeercP ia emse;.i canus habse einn tpetrescdee vetriamilent s hU en itedo fAS mteartoienics ma p orted plants. T?en egwe niucsso nsitdobe erletodotn hgCe o cctirdiCaboeec cainnadki e,iy ps r o­ vidtesode pairfrtao tmoe t hgeern ientr hati rs i be. MikWiel liaDmesp,a rtofmE enntto moalnPodlgy a Pnatt hoAluobgyu,Ur nni ve3r0Is ity, FunchHeaslAsll ,a ba3m6a8 49-USA5.4 13, ChrHiosdgs onD,e partomfBe inotd ivaenrBdsi iotyl oSgyisctaelm ThaetNa itciso,n al Museuofm WalCeasrd,if Cf,F /3 NP,Wa leU.sK,. EvelMy.Dn aan zZiogo,l oIngsitciaRtluu stseAi,ca and eofm Syc ienUncievse,r sitetskaya nab1.S, t .Pet1e9r9s0bR3uu4rs,gs ia. Introduction matcilces fth alalnoacwll, ed fti stmianrcg ti,n s adprebsunstoe futds eDdo.r sduemrm:me mbra­ Anu ndescsrpiebcoeifsde o ssf cta (lHeo mon­ouosny ouandgu blutwt,i tanh a rrsocwl ero­ pteCroac:c iCnoecac:ifr doamCe e)n tArmaelr ictaisaatrioouannn dt emrairogorif an n apll ates; hsab eeinn terocnes petveerda l boytc hcea bseicoonmssi cnlge rmoetdiisoaeuldtl wyaa rtmd as­ quarantiinnteh Uesn eirtveidcT ehSsi(t sP.a tteusr.wi ittayhr; e olaltairongneesasm,rta rgbien­, ameriacnaadnn ao)tr heelras tpeedc( iP.ae gsa ­comisnmga lolrae brs meendti aDlolryss.ea tl a e vea)l,sfr oo mC entArmaelr iacrhaee, r e dset­oustpliyna onstdea pesrpiantrghs;re o ughout. scriabsne edaw n pdl aicnaen de gwe nPursi,o D­orpsoarloe fts h rteyep( eils)a: rp gree ope rcular nocoWciclulsi Haomdsg,s& o Dna nzig . poTrheeeascp,ho rheea vsicllye rwoittaibh sr eo da,d formsepre caipepset aobr ehs i gvhalryi bau btlbea sdailsi cand: i stlionncgti tbuadneidxn taeln d­ noc onstant cdoiuffbleefodr u entndocs eesp ai­nagn terfrioomar nlpayll a ttaoeb sop urto thorax; rattheve a riloouotsfms a te rial. (idio)rmsiaclr odeuacctwhui ltaefih ,n ien ner DepositBoMNrHiT:eh seN: a tuHriaslt odruyctule:t hfrreoquugaehnno(dtui atis ,im )a ll MuseLuomn,d UoKnU;,S NMU:SN atiMoun­alo pesni mplep-rpeosrteehn:rt o ugDhoorusta.l seuomfN atuHriaslt Woarsyh,i nDg.tC,o. nt,u budluacartn sdd o rtsuable racblseeAnsnt a.l U�A;Z IAZSo:o loIgnisctaAilct audteeom, fy p laatpepsr oxiqmuaatderilanytn meea,rr gins ScienSctPe.es t,e rsRbuusrsCgiS,aD ;AC :a lim­oroerl epsasr allel, possltiegrhtiloyr margins forniSat aDteep artomfAe gnrti cuSlatcurrace­o,n veeaxcp;hl awtiet3 sh t rsoentgda oer sally mentCoa,l ifoUrnSiAa., neaarp e(x n 1e aera cihn nmearr gnienaa,rp ex annde paors tmearrigoairnn wd)i ,ta sh h osrett a Prionogceonnc ..c us ventnreaaalrpl eyAx n.o -gefonliwdti ataplh a ir ofl onsge otnaa ntemrairogarin n3d p aiorfs Type species: Pargiaosvnpeno. .c occus shorteorn l asteetmaraera glpi lnutssh at near DiagnAodsuilst: Bfeomdaoylv aetlo.a l mosatp esxu;p pobratdrii nsgte ixntcetna,dn itnegr ior­ rounsdy,mm etorrai scyammle trpircoab la,bl lyty oa ntemrairogorif an n apll aAtneasrl.i ng ratflhaeitrnl iwfei,d eacsrtao bsdso msetni;gw -itfihv pea iorfss e taanewd i tahni ncomplete 11 2 M.Willeitaa lmA.sn : e gwe noufCs o ccidaZeOOSYSRTO.S SIVCoA1l 1. doubrlooewf p orMeasr.g mianr:g sientaaalle l i nH odgs(1o 9n)9 ,P4 r ionockoecyocsuua tst sligdhitslpyl oancteod doofrt swumoma ,i nM esolecMaonriruitmsow o hni,ic ithsc learly typ(eiass) :t osupti,ns oestleao ,n gpeosstt ericolrolsryee llyaL tiekMedes. o lecaPnriiuomn,o coc­ bustl igvhatrliiyans b hlaeap pei;hc aaollfef a chcuasp petaobr ers e strtiotc hNteee wdWo rl d. spiwnietm hi nudtiem pelaecswh;i tabh ro adP,rionoccaobnces c eupsa rfraotmoe tdhC eorc ­ we-ldleelvopbeads- aslocukseuta:rl alty h ecribnyir eplaccoiunp1gl2i e ntt h Ceo cckienyi abundaanndt a;s( eitiso)es teeaa ,cw hi tawh e llofH odg(s1o9nw9 i4t)th hfo el lowcionugp lets: develboapsseaodlc ksepta:rd siesltyr bieb­uted twesepni nsocsteSa tei.g msaptiiuncne dsi ffe1r2P­.reopeprocrupelrsae rsi enan b tr olaodn gitudinal entifraotmme adr gsipnianEley ess.mp aortg inalba.n adn tetroai noparll a tweistfe;hw s etparee sent aloanngo -gfoelndid;to artlsu able arbcsleeensat c;h Vendteerrmm:e mbraPnroeugse.dn iistca-lp ores postbearniodofd r i sc-npoeotxr teesn frdoimen agc h eacwhi t5h-l 7o cpurleis:oe npn rte gesneigt­alp osteriotrom esspoaifder aacmchel seo c.o.1x.3a menotn lmya,i nllayt etroaa ndt eerniodofa r n al Preopeprocraueblssa ewrni tnt;uh m ersoeutasal eo ng cleafnatdn palla Stpeisr.a dciuslca-wrpi o trhesa no-gfoelndid;to artlsu able prrcelseeesan cptho; s ­ main5ll oyc iunaln ia:r rboawne dx tenfrdoimn g terbiaonordfs piradciuslca-prpa oesratec psho ste- risopri rtaomc elseoa fed a cmhe soc.o.x...a.. .. eacshp irtaomc alregV ienn.tm riaclr ofrdeu­cts ..............S.u.a.r.e.zM.ia.em.le.lt.a . ... queinnat b romaadr gibnaanoldn ,h eaadn d 13M.argsientaoalf2e o r3t ypoensfie,n esleyt aonsde aroulnadb ibuupmto ssaibbslmeyen dti oanl ly thoet hsetrosus tpliyns otsieg;mc alteeiftacscw hi th abdomen. Pproeraaebnsst eeVnnetnn.att lru a­l spinnoectsl edairffleyr efrnotmmi aartgesidpn ianle s buldaurca tbss eVnetn.ts reatalan ela:ol bs eet ae ................P.r.i.o.n.og.ce.onnc...c. u s - Margsientaoalfoe nt ey psep,i nboeswneit,at b hl unt undifferefrnotmsi uabtmeadrs geitnaael ; with apesxt;i gcmlaeteftiasccw hi tthh rsetei gsmpaitniecs lopnrge gesneitotan3aes l e gmoennltbysue, ta ch cleadrilffye refrnotmmi aartgesidpn ianelea smc,he ­ segmweintatsh p abrasneod fs hosretta acer ossd iasnp idnies tilnocnttghleaylrn a tseprianl.e . s. . eacahb domsiengamlue snuta;l l2oy rm owriet h ............M.e.s.o.l.e.cM.ao.nr.ir.ui.ms on setnaeeaa rrt icoufel aaccthoi xoiann; t era ntennal settawepo:a iwrists;he veral small PsruibomnaorcsgohicanwcraiueltssMe h s olecatnh ieum setbaeet wseteing maarteiwacis t;1h p ai r foolflowcihnagr a(cipt)re erosp:e procrqueulsia tre hypopysgeitaalc ; othearn qdus ie ttleaaer igsnaemb ,ar lolla odn,g itbuadnaidnn taeltr oi or sparAsnet.e n6n-asee gmsecnatwpeiedt3 ;sh e ­anpalla (tiediso);r sseatslal ei gshptiln(yoi siei;) tapee;d i2cs eelt+:ac ea mpanpioforrseme; g ­dorswuimta hr eola(tiipvor)ne sg;ed niistca­l menItI 3Is :e tIaVe1:;fl essheyt oanelV y:I, porreess ttrmiooc stpteo ds taebrdioomrsi enga­l flesshey+t as1 e tsoestaean V,dI 3:fl essheyt amee n(tvp;)r egedniistca-wlpi omtrahei ssn elvy en + 5o r6 s teose/sspeitnMaooesu.et hpuasrut­lso c(uvlaiib);s eonfvc een ttruablud luacartn sd, ally apslyamcmeed trliacbaigluelmny e;r a(lvlityii)bios-etgamresnuatsba stoeirnpo ton o rly twiswtiet4dph ,a isrmsas lelt Sapei.r naocrl­edse veloped. maLle.g uss uaqluliwyte edl elv elbouhptie gdh lyEtymolPorgyi:o noicfosor cmcfreuodsm prionos varisaobmleet,wi imtehsdo iusttt irnoccth an(tGmor)­.m ean'isnagrw e'fe,rr tiotn hgme a rngail femuarn tdi biosteagrmseanlts aettciaooexn:a; s etael wohorkia ctlhhi etkrhet e e eotnah c hna­i 5-t6r;o ch1al notnsegeor tn av entmraarlga innd saawn,-d c ocacftuetsrhfa emi lnya m e. onseh osretot nad ormsaarlg feimnu;2r ;t i b2i-a Prionoccuorcrcceuonsnt tl2ays i pnescP. i es, 4a ntda r2s-u4ts;a rcsaamlp anpioforraemb ­agaavneP.d a mericbaontduhes s,c raisnb eewd senctl;aw wi tadh i stinctta rddseianglti itcublleeel;. so w extenwdeiblnelgy otniodpfc lacwl;ad wi gitules subeiqlnue anltg tota hr dsiagli atnuedlx etse nPdr­ionoacgoacsvcpenu.. s inwge lble yotniodpf c laown;ce l adwi gitule generbarlolatydh eaornt hcelrad wi gitule.Holotyp<;>e,,M exiScaoLn,u iPso teoxsl ieo,aAf fg a ve CommePnrti.o noicosec acsruieslc yo gnifalscea(dtA ag avaceae)R,. HL9o..d VgId.o1n9, 8 0(,U SNM: Lare1d3o9 6y1o)u:un ngs cleardoutfelimtsa elodena by( it)hl ei onfel a rpgree opeprocruaelnsat re ­ sliwdiet5a h <;>d ,h olofetmyapclele e amralryko enmd a p ritootr h aen apll a(tietiseh,)r estroiftc htei oonrn i ghtl-ahbaenld. pregedniistca-ltp oto hrmeeo ssp to stperrei­orP artay pMeesx.i acsfoo r,h olot4sy cplee,r aod9t ised gensietgaml(e intithi;ae) b seonftc ueb udluacrto sn s liwdieth ho lot<;>y p(eU SNSMl)id.de ep osi inted onb otthhd eo rsaunmvd e nt(eirtv;h)r e o b usUtSNM. Othmeart erMieaxlii.cn ot,e racteB prtoewdn sville, spinmoasreg sineatelaa eco,hf w hiacphp ears offl eoaffA gasvter iBcutragR,ee sis(s UtS,N 6M8 97), toh avase l igdhitmlpayln esddp insuulrafatce1e /,a1 d <;> i ng oocdo ndiStliidodenep. o sZtIeAdS .i n an(dv t)h peo sseosfts witooy np oefms a rg inal setsapei:ns oestaean efld a gelsleattaee . Aduflte ma(lFei1 g().D escfrroim6bs epde ci­ DiscusTshigeoe nnP.ur si onoacpopcecaumrses n 2si ,ne xcelcloenndti4 tm ioorhnee, a vily tob elotnotg ht er iCboec cFianlial sed ne fin esdcleraontddio sressdup mbl uisttt qiuliglto eo d). byH odg(s1o9n9 I4nt) h.ke e gyi vteogn e neraU nmoumnatteednr oistae le:n . ZOOSYRSOTS.S IVoClA1. 1 M.Willeitaa lmA.sn : e gwe noufCs o ccidae 11 3 K m .\ [ f>-..; '. �-,- p··. ._, ·\�V�,�- C - �u '/ ;/ � . .. . ' --<' , .J ,•-•.�...- •----.... -.,, ....,........•,_ 1......1,1,.--. ·'l·1·• ··.A .-,.-1 I I N ® G Fi1gP..r ionoacgoacsvcpeun. s.a ,d ufelmta lIent. ha insid nF i2g-sA6 = :d orsseatlBa= ed ;o rssiamlpp olreCe ;= dormsiaclr odDu =c tpurleeo;pp eorrGce =u;a l naparll a Ht= ea sn;o -gfoelndKi; =t m aalr gsienta(alibe :u lb ous spisneoitsiale:;a ncesoplianstoeesti eaifl ;ia :g esleltaLat a)e=n; t eMn =ns ap;i radciuslca-Nrp p o=rr eeg;ed niistca­l poPr ev=;e nmtircarlo Qd um=ce tt;a thloeorgrca lc aiwc. 114 M.Willeitaa lmA.sn : e gwe noufsC occidZaOeOSYSRT.O SSIVoClA.1 1 Mounmtaetde rbioadslyh: o wvienrsgyl i ghltenogft"hl ofinnsgee" t soesotenta e rmsienga­l sigonfas s ymm(emtorosybt v iionpu oss itimoenna tb o3u8µt m M.o uthupsauratslshl oyw ing ofm outhplaerntg1st.)h5; 7 -m2m.,w6 i3d tho nlsyl isgihgotnfa s s ymmeltenrogyfct; lh y pe­ 1.3-m2m. .0 olabsrhail1e 5l3d- µ1m6l;0a bisuomm etimes Dorsduomr:ss eatlsa pei neoisteth,ae pre ritnwgi sStpeidr.na ocrmlaeelsa: ca hn tepreiroirt reme toas hapropi( nBtr ownmsavtielorlrire aa lt )h e3r3-µ4m2e, a cpho stpeerriiotr3r 8e-mµ4em3 . bluanntpd a rall(eSla-Lnsu iiPdsoe tdmo astie ­Legpse rhraaptssh mearwl ilt,h aonudyti stinct riaela)c7;h- µ9m l onDgo.r spaolr( eisp):r e­segmenbteattwiteorenon c haannfedtm euror r opercpuolrearea scb,hr obaads dailss ucp porbte­twteiebanin tada rlsounsg;ce osxstae let aac h inagr atflhaectrl opsoerdle a;r( geea5 c-hµ9 m 33-µ4m3l onlgo;nt gr ochsaenettaaec arhb out widien)a d: i stinct blaonondfg3 i1t-u6d32i 6n-µa3ml4l onlge;n g(tµhmos)fm etathloerga:cic por(eisdi;o) r sal miacnr(doi daisu imc)at lucllo ex,8a 3 -96; t+rf oecm1hu0ar7n -tt1ei1rb6 i;a + opesni mples-ipmoiirsnlei a,tzr moe i crod ucttaurl1se3u.2s- c1l4a02w;1 -24. Anapll aetaec1sh3 4-µ1m4c,5o mbiwnieddt hE tymolTohgysi.ps e cinisae msae fdtt ehhreo st 15158-µ0m e;a cphl awtiet3 sh e teaaec2 h1 -32plagnetnA ugsa (vAeg ava ceae). µml onAgn.o -gefonlidlt:oa nslge o tnaa nteriCoormmeP.n atg.a ivsve e rsyi mitlota hforel ­ margeianca hb o4u0t- µ4m5A. n arli nwgi:t3 h lowisnpge cbiudetis ff (ec rhsa ractoenPr -states paiorfls o nsge taanetd w roa ther shoarmteerri.icnab nraa ckienht asv)i( nidg)i s tinct Margimna:r gsienta(alise )p: i nsoestleaa en ­grooufsp psi intne hsse t igcmlaset( fingtcor oups ceollaotnegp,eo sstt ebruistol rilgvyha trliya bolfse t igmati(cio ins)lop ynit eny eopsfe)s ,p inose isnh aipsnei :n rgolawer omuanrdg hiingv,ha lryi m­argisneatla an(ttehsrepi ionrso estaear ee ablfree qinu elnecny1g;1t -hµ2 m0l onagn;(d i im)a ibnullyb woiurtsoh u nadpeidwc heesr,te haes setsoesteea aec1 h0 -µ1m7l ondgi;s troi fbumtoispoton s taerrleia onrc eaonl(dai tieic))l, a twhe setoanBe r ownsmvaitlelr2ei0s apli:n+ o s8e d igivteurldyei sff eirnes nit(zo eft ewni tohn e setboestew eeyeens 1p5o-ts1sp8,i n+o s2e s ebtroosaetdheatrnho et hbeunrte vaesmr a rkseod ly betweeyeensa pnoadtn tesrtiiogrm at1i6c- acsol nP.e a ftg,a vOet)h.e rtwhietssweeso p eacriee s 17s pin+o ss1ee tboestew seteing mcalte ifcct lseacrlloysr eellya ted. late5r7a-ls5lp9yi ,n+ o ss6ee tboestew peoe­sn tersitoirg mcalteaifnctad sn acll eafnt7d,- 9 Prionocaomcecruiscs apnn. .u s spinoonslaeyl omnagr goifan n calle Sfta;Ln u is Potomsait er1i2a-ls2:p0 i n+o 4s-es8 te ose Holotayd9p ,Me e:x iCcooz,u meexCl o,c coslpo.b a anterbieotrweleyyee ns p7o-t1ss0p, i n+o1 s- e2 (Polygon2a0c.VeIaIeIM)..,1L W9.i8 l8l,(i UaSm NsM , setose betwaenaednn t eesrytieiosgrp m oatt ic AL-028S-l8id8de)ep. o siinUt SeNdM . cle5ft-,1s 3p in+o sslee tboestew seteing matPiacratMyepxeisac.sofo ,rh olot2y9sp lei,wd ie8ts h clelfattse 2r3a-ls4lp5yi ,n+ o ss6ee t boestew eeynou andg9 ( 2o fd orsounmls1y 3)o ,la dd 9 ( 3p oor), postsetriigomcral teaiftncsad n calle aftn,1d - 62n9d, 22 nc!d, 1 4c raw(l8pe oros+r )eIm br(yDoe.­ posit2oa rd9i B eMsN:H 2,a d9 Z IAISa ,d9 NMW; spin+o0 s-se1e taolsoemn agr goifan n calle ft. reisnUt S NM.) Stigmsaptiiuncne dsi fferfreonmtm iaartgeidn al Othmeart eMreixailCc.uo a:m aMvoarceael xYo usc,c a lancesopliabntueefots r miand gi stgirnocutps ,p3 . .,lI II.E1.S9 h9u6v,a (cZhIi7An3Sa- 926/a)5d9, i n with9 -1p0ec rl (eftB rownsvilaln4ed- 9m agotoecdor nidaialtn)isd oe nv eurnamlo uantd9e dl+ a rvae; (SaLnu is PotoAsniac llm aeabftto e1uri/ta1 l0)tC.hh iav,Oe alxaocnaC ,o ccoslpo".bC aa me(rPoo"l ygo­ bodlye ngEtyhe.s epaoct1hs2 -µ1m6w id e. nac)e1,a9 e2 G6F,F. e r(rCiSsD- AX,- 64-remount-.X.1964 RJW3)/,a3 d9 ( +embirngyo ooc)do ndiitnitoenr;c epted Venteprr:e genitalw idt1ih1s -co1-n 4p ores: Browns(vUiSlAel)xAe, n thurium (cAlraarcienaeer)i,u m eitheSrp isriaddceiu.sl ca-rwp iot1rh8e -si2: n5 9.VIIIB.u1r9g(7eU7sS,sN 7 M5 915/)a4,d9 e+m bryos; eacahn tedriisocr-b paonardne 1d 8 -i3ne7 a chi nterBcreopwtnesdv ilelxcA nt(hUuShrAoi)ou,km e ri, postbearniVedon.rt rmailc roadsiu ngc etnse r2i6c.V.G.1 B9u7r8g,(e UsSsN8 M3 613/)a2,d9 ( poor)+2 diagnVeonstirssae.lt a aneal:lo bsee tgaeen er2­nd9 ;i ntercBerpotwends( vUiSlAel)xeA, n thurium robus\t0u.mV,I IBIu.r1g9Vae7ns7vs,a lke(nUbSuNrMg ally undifrffoemsr uebnmtairasgteietbndaua etl 75981)/a,2d 9 ,2 2 nd9 ;V eracrduizvi. n,t ercepted occasiuoptn o2a 2lµ lmly o nlgo:n gpersetg eni­ Browns(vUiSlAel)xAe, n thusrp2i.3u,.m lll.1973, Stevens .asle teaaeca hb o7u4t- 1µ0ml1 o nogt;hs eert aeeta l(.U SN3M5 725/)a3,d9 ,22 nc!d, 2 2 n9d p+u pa; presmeendti aalclryao sbsd omsiengamle nitnst:e rcaetMp itaemodinP , h ilodesnp(d.Ar roanc eae), VI1I:5o r1 6V,I8 -1V1:9, - 11,e aIcV6h-- I8I;2:7 .11A..1SM9.i5 l1(l,Us S N2M0,0 561,. 812/4a3)d9 , + 2nc!d. G uatemianltae:r LcoeAspn tgeed(l UeSsoA n) , setaases ocwiiatethae cdch o xmae:t ac2o,x a leoaffu nknopwlna2 n0t.,l .K1.S9 h8i6m,(a UdSaN M, mesoc2oa xnpadr oc4oa,x lsalh oirntt;e ranten­ 05858764--5624/a14d9) ;,i nteracNte epOwtr elde ans, nasle ttawepo:a ilrosn,ga ebso2tu5 µt m l on go;nC occoulnoifbe1ar7 Va.I, II9 I 8.H2 .,F inge(rmUaSnNM wit6h-s 8m aslulb marsgeit(naeaeal3 c -hµ4 m 014852/0a9)d9 : ;i nteracHteo pbtoekoden An n,t hurium lonbge)t wseteing maarteoiatcsh s;ee rts amea lslal1vl i.iV,I IL..S1 i9r4(o9Ut,So N 1M3 6520;- 217/13), and qsupiatrAesn et.e ntnoatea:l 1l7e5n-agd9t ,5h2 n9d.H onduirnatse:r Mcieapmetixee, pd i phytic cact2u5s.,V IlR.E1.C9 o7t5(t,Ue SrN 1M1 7430/ad)5 , 20µ8m t;e rmsienqtauali l toenl gne,g t3h3µ m ; 9,22 nc!d 2+n 9d. ZOOSYRSOTS.S IVCoA1l 1. M.Willieatam lsA. n :e gwe nofu Cso ccidae 11 5 Descrfriobm3e 0sd p ecimmoesnitsnlfa, yi rOaxaicnMa e xihcaomd o satn)(d;i ais )e t ose tgoo ocdo ndiNtoittoehtn ah.tii assh igvhalryi s­e1t6a-,µ4 m0l o nwgi:4t -hb9 e tweeyeens pots; abslpee c-iicetos u rledp raec soemnptol fe x1 2-b etweeaecenhy esapnaodnt t esrtiiogrm atic spebcuinteoc s o nscthaanrtca ocutbleefodr u n dcle1ftl ;a tebreatlwsletyei ng cmlaetafint6cd-s tsoe patrhaeatmnei d it hs e arses utmhetadht i s7a loenagcs hio dfea bdomaenwndi, 0t h-o 2n variraetpiroehnso esntt-sio nerdn uvcierdo nemaelcnah­t mearraoglfia n nc alle Sftti.g cmlaetfitcs tally-(iobnrod tudhci)eff de r ences.sligshctlleyr aoltoimnsager dgs itni:g matic spiunnedsi fferfreonmmt airagtbieundla blo us Adufelmta le( Fi2g,3s ;Ta bl e). spiEnyeess.ap obou1tt1 -µ1m3w ide. VenteDre.rm me mbraPnroeugse.dn iistca­l Unmoumnatteedrd iraislep:de ciomve anlsp, orweis4t:-h 2 o0ne itshie(dr1se p echiam den ratflhatetsor l igchotnlvdyeo xr,ss ucml er oats iisnedgdilsec -opnso ergem VeIn)St.p ir acular anbdr owwinat t hh ciont tcoonveyex rc meep­t disc-wpiot9rh-e 3is4ne: a cahn tebrainaodn r d diallelcayof;v eirnfle udffw yh ictoet twoan xy,1 33-9i ena pcohs tbearnVideo.nr t mriaclr oducts whiicashs sutmohe advb ee esne crbeytt h eeda si ng enedriiacg nVeonstirssae.lt l aoen:g est adufelmta l es. prgee nsietteaaaleac bho 7u9t-3 µ1 m3l onlgo,n g Mountmeadt ebroidsayol m:e toivm(aeFls i gs.eatlars aor perleyso ensn etg mIeVon;tt h seer­ 2b)ugte nesrhaolwlsiyln igtg och otn sidetraapebr leesmeendtia aclrlaoybs dso msiengamle nts asymmweittorhnys e,i sdieg nifilcoanngte(lrtyo tVaI:l1I s1)-V:1 I98 ,- 1V5:4, - 1I8V5,,- 14, thoatnha enwrdi atnhla olob noe n sei sdter onIgIl5Iy-:1a n4Id I2 :-O 1s; e taases ocwiiaettahec dh con(vFei3xg) a.;n callae bfot1u /t9 -1b/oldOyctohx mae:t a1c-omx5ea,s oc2o-ax6na pd r ocoxa lenlgetnh1g;.t 3h-m 3m.,w1 i d0t.h8 5m-m2..51 7-,u suaallsllhy o orctc,a siloonnaglelsyt Dorsudmo:r sseattlaa ep etraosi hnagpr opi ntpr:ocsoexutapta 1o 8 µ ml onega;cp hr othoracic perhmaopscsto mmmoend ioanla lbyd o mesne;gmweinsttoh m see tmaeed iianltleyr;a ntennal ea7c-hµ1 m0l onDgo.rp soarl(e ips)r: e op ersceutl2apa ear:i( rrsa -r5eG,lu ya teamLtao lAsan ­ porfaeisrl layr egaec5,h- 1µ0mw idwei,ta h gelleosn)g,2e 5s-µt6m 5l onwgi;t6 h- 1s3m all brobaads als-udpipsocarc toinnvcgel xo sesdubmarsgeit(nalaeel n3 g-µt8mh l) a tebrea­lly poriaend : i sltoinngcitbt aunod9dfi- np5ao9lr est;wesetni gamraetoaitschs; ee rts ameab lufrlte ­ (iasi m)amlilc rodpuecrthmuaolpscesto: mm onquepnatr,t iicanub lraomraaldry g biannAadnl.­ nemaarr gainundp m argoifan nsca lle afntd,t ennIaIeIs:re dg mreanrtwe iltayph s eudo-ar­ (iiais )maolples ni mpleA-npapollr aept.oe ss­:t iculeaatc1ih6o 0n-µ;2m 7l 4o nlge;no gftt ehr ­ termiaorrgo iftnewsni tadh i stiinndcetn tamtiinsoaenl2t; 8a- µ5m2l; e nogflt ohns ge tsoestea len1g1t1h- µ1m6c,2o mbiwnied1dt4 h0 -2o1n0t ermsiengam3le3 n-µt9m 1M. o uthupsaur­ts µma;n apll asteeta alselt rosnegtlaoyns dea ldliys taisnycmtmleyrt artichcleaortlso o,en re subeiqnlu eanleg atch2h,8 -µ5m0l ondgo;r saplrocloexnaog;fct lhy peoslhai1be2rl5ad-l 1 90 surfafrceeq uewnittplhaya l ree (aasr eolaµtmiS.op nisr)eaacaclnhet se:pr eiroir2t 8r-eµ4mm7e, neaanrt emrairogArin no.- gfoelndil:to anslge ­eacpho stpeerriiot3rr3 e-µm6me5L. e gpsr:e s­ taoena ntemrairoegrai 2cn2h - µ5ml8 o nAgn.a lencoeft ibios-etgamresnavtlaa rtiiuaosbnul ­e, riwnig2t o hr3 p aiorfls o ntgh,is cektp ale2u s alwliyst lhi sgihgotnfa ss m aplsle udo-segmen­ o3rp aiortfsh insnheosrre,tt eaarne ta;ula b beo uttatoinso mna llleegbssutw,t i tadh i stsiengc­t 1.5-l2e.n0ogxfat nhpa lla tes. mentaotnli aorng leesgltse ;n g(tµhmos)f MargiMna.r gsienta(alise )p: i nsoestoea f em etathloercgao:cx7 ia4c - 1t3r3o;c +h afen­ter twtoy paenst:e rbiulobrwoliuyatms ho roerl e ssmu9r7 -2t2i8+b; it aar1s0u7s- c2l5a12w;2 - rounadpeedax n wdi tahd impslufaercd e 35µ ml;o ngceosxstae let aac5 h1- µ1m1l 2o ng; apiceaalcalhby o,8u -t1µ 7ml onagtp; o st erlioonrgtersotc hsaenettaaec arhb o4u2t- µ9m5 enodaf b dommaerng,si entmaaolerl ea nce olloantcgel;,ad wi giatlumlsoeissmt io lns amra llest butthv iasr isaobmslepe e,c ihmaevnsisln ig g hltelgdysi,s tdiinscstiolmnlyi a lrlageresg slsti, g htly poinstpeiednv eeosnn h eaeda;ca hb o1u3t- 2b5roatdhetaran r dsiaglia tnuedlx etse nwdeilnlg µml onagl oanngca lle iftna;s intgodl oeu blbeeyotniodpfc law. roawr omuanrdgw iin1t;5h - b5e0t weeyeens poDtisscutshsmiiasot ne:wr aiesax lt revmaerliy­ onh ea6d-;4b 3e tweeaecenhy esapnoadtn tea­blTehm.ea ivna riwaatisin (o:ibn )o dsyh a pe: risotri gmcalte1ift1c;- l4a3t ebreatlwl eyte hnfraotmM oreMleoxsi,(c Foi2 g)w.,a bsa r ely stigcmlaet(ftirscom wosro erl edsosu wbhle en asymmewthreirmceoaaslost t hmeart e(er .i ga.l spianbeusn d3a6n-t1a)3l,8oe nagcs hio dfae b ­fromG uateimnatlear,ic nMe ipat(meFidi3 g) .) dom(eo nfw hipcohs tIe0 r-i2ao0rlr ea nce oslhaotwece)odn sidaesryambml(eietni ru)ym ;bo efr anwdi 1t-h1o 2ne itshieodrfae n calle aftl,w aymsargsipnianlso estteah ee:s em owrteehr ae n lonagnemdso tsp to in(tmeadtfr eormHi oanld ut­wiacaseb unodnma antte frroimMa olr ealn ods raansdo fPhfi lodefrnodroMmne xihcafeodw e stOaxaMceax,i acsoo nt hmea tewriittahhl e spinmoasregs ientatalhe ofr,so emM orealn odsfe wemsatr gsientafraloe mH onduarntadhs a t r e : t y n - r ;: P. ble. Ranges in data for 8 collections of adult female americanus from various localities. Where: No. = number of specimens studied; column 1 = length of anal plates (µm); 2 = no. of eopercular pores; 3 = no. of marginal spines along anal cleft; 4 = no. of marginal spines laterally between stigmatic clefts; 5 = no. of marginal spines anteriorly between eyespots; 6 = no. of length of coxa (µm); 10 = length oftrochaegenital disc-pores; 7 = no. of spiracular disc-pores between anterior spiracles and margin; 8 = width of anterior spiracular peritreme (µm); 9 = mur (µm); 11 = length of tibia+ tarsus (µm); 12 � claw digitules: where O indicates digitules similar ranging to 5 very dissimilar; 13 = length of antennae (µm); 14 = degree of asymmetere i indicates barely asymmetrical, only mouthparts displaced; 2: intermediate; 3: margin of body showing high degree of asymmetry, particularly posteriorly (as in Fig. 3); 15 = tibio� rsal articulation: where I indicates pseudo-segmentation poor, sometimes absent ranging to 5 where complete segmentation present.� i;;· 5 JO 149 12 Locality 1 l 1I 13 5 3 8 No. 2 46 7 "' ~ 215-240 165-199atemala: 145-162 I I 07-128 343 26-42 6-1241-47 145-162 13-3123-34 12-18 15-37 f.:'-boken 4 150-190 atemala: 166-183 3132-1661-3 95-112 31-37 4128-145 24-55 2 12-269-22 1-5-13 21-35 ;,: w Orleans OQ 186-228 169-2071-3 4 145-153 atemala:107-132 45-9 36-44 2 139-145 26-49 24-3114-24 14-18 18-33 s Angeles �I 211-248 136-166nduras: 2 5 5 161-186140-91-120 153 9-18 3-836-42 17-28 2711-15-21 5-10gami 835-236 248-270 xico: Oaxaca 157-173 120-132 1-2 3 43 7-10145-150 52-55 37-1446-53 15-33 30-42 160-178 xico: Miami 3 108 161-166 3-52 74-83 16-21 2 33-42 134-143 12-30 11-164-1015-18 l xico: 3 256-273 221-250l 11-124 157-174541-47 44-59 434 8-1426-6-1711-18 144-161 30-50 relos 0 2-3 xico: 169-2034 186-228 132-145 86-116 36-43 5-8 16-31 18-27 124-133 411-20 2 13-24 21-37 (/) zumel ;l nge 108-2503 31-53 160-273 74-132 9-59 1-1411-43132-1903.5 124-162 4-209-3450 15- � arrfeha uo ue uo oi e e eo eo a Tpp+wt GH GN GL HM M M MM MC R ZOOSYSRTO.S SIVoClA1. 1 M.Willieaatml sA. n :e gwe noufCs o ccidae 11 7 C .•'.• ·, . ,•,. 0B 0 . d 0 K. . II @ N G H Fi2gP..r ionoacmoecrciucssap nn..au ,ds u felmta lfroem M oreMleoxsi,Nc oott.eh t ahtfoi rsmi asl msoysmtm et rical, apafrrotmt hpeo siotfti homeno uthpLaarbtaessli. snF igIe.,x cRe= pv te ntsre atla . 118 M. Wilelatil aA.nm :es gw e nofu Cso cci•daZeO OSYRSOTS.S IVoClA1. 1 iii L D �-,--B, \ 0--- ,' II ' ' \ I J 0 : 0 ' \f : § f , N G @ Fi3g.P. r ionoacmoecrciuscspan .n.a u,ds ufe lmta frloem C ozumMeelx,iN coott.eh t ahtfoi rsmi sst roansgylmy­ metrLiacbaaelsil. nFs i Ige,.x cJe pm=ta rgin. ZOOSYRSOTS.S IVCoAl . 1M1. Wileltai laA.mn :se gwe noufsC occidae I 19 interactMe ipatmfreiod m M exi(ciosi;ii oz) fe latmearraglei anc2,h3 -µ3m0l onsgu;p po rting thlee gasna dn tentnhaleeo :n gleesg(tsfr o mbapro odrelvye lAonparelid wn.ig t6 sh e taaned MexiOcaox:a Mcoar,e Cloozsu,m Geula;t ema9lp ao,ra ensta;ul bseh o rt. interacLteo pAstn egde wleersme)o rteh tawni ceM argimna:r gsientaoalfte w toy p(eifls)a: g ­ thlee ngotfth h seh orlteeg(ssMt e xiocff o,e llatee ascuehptt ao3e 0µ, m l onagn,(d i i) Philodetnhadenr toenns)nh;ao ewt ehdse a mes pinsoest(eae ea wcihts hm adlilm palneds/ or trenwdistth,hl eo ngmeosrtteh aIn/ 3lrodn gesrpisno unal peihcaalolf)f2 t ypreosu:n bdueld­, thtahnse h or(tievts)hpt er; e soefsn ecgem enbtoau­ssp i(ne easc8µ h m l onagne)dl onsgpaatteu ­ tiboent wteiebanin tada rtshuwisa:sws e ldle ­lastpei (ne easc1 h1 -µ1m3l ondgi)s;t r ibution: veloopnte hdle o ngleesbgtusi t n disotrai bn­catr ouhnedab de tweeyeens p4bo utlsb:so pui sn es seonntt hsheo rtaen(sdvt t);hs eh aopfte hc el aw+ 8fl agelsleatotaneee a;c shi bdeet weeyeen­ digitduilsetsid:ni cffteolrnyet nhlteo ngleesgtss poatnsad n tesrtiiogrma arte3iab c:u lbous bumto roerl essism iolnta hsreh orTtaebs1lt e.s pi+n efls2a gelsleattlaeae t;e breatlwleye n givseosmo eft hdea fotra8 l ootfts h miast erisatli.g mat3ib cu labrsoepuais+sn :efl sIa gellate Iitcs l etahrta hte irmseu cohv erilnma opso tf setaan;pd o stetropi oosrt esrtiiogrma arteiac: thedsaebt uant os tructduirffaelrc eonucble eds wit2hb ulbsopuisan netse rtihoebrselecy o,m ­ founwdh iccohn stantltyh esImena.p d adriai­tnsegpd a tpuolsatteewr iit5ooh rr6 lo yfl, a t+t e7r tionnod, i ffere(nocttehhsea rin ns izwee)r efl agesleltaSatteei. g msaptiuinnced si fferentiated founidnt hoet hleirsfe t asgteusd aineddd­e fromm arginal spicnleesaft;bs s esntti.gmatic scribbeelIdoht wa .ts h erbeefoeranec cetphtaetdE yesopnmo atr gwiind;1t 1h- µ1m3. thmiussr te preass iennhgtgil helv ya riable Vesnptede­err:mme mbraPnroeuamsnu.al lt ilocu­ cietshd,ei fferences bbyhe oisntg- pcllaaadurnis tse cd- apbosreSenpsti .r adciuslca-erpa ocrhe s ore colofagcitcoarls . wit2h-( 5m ai3n)ll oyc iunalb ia n1pd o rwei de Pa.meriicesaa nssuieslp ya rfraobmlP.ae g avbeetwsepeinr aancmdla er gwiint3,dh i sc- pores in( il)a ckaid nigs tgirnocoutfsp p iinnee asc hi ne acahn tebrainaodnr 3d o r4 i ne acpho ste­ stigcmlaet(fiihtcia; )v idniffge renmta-rs­hraipboeardn V den.t rmailc ropdouscstriseb sltyr icted ginsapli annetse rainopdro lsyt erain(odir iliyt);o1 p ostteoer aicsohcr a 1pb ee,t weeaecpnhr o­ inha vitnhcgel adwi gimtuuclmheo srs ei m ilaanrmde socaonx4da- s,5u bmariganib ndaolmliy­ thoannP. aga ve. nasle gmeInIVt-IsP. r eantpeonranebassl e nt. Ventrtaulb udluacartb ss eVenntt.rs aelta anea:l First-innysmtpa(hrF i4g). lobsee tnaoedt i fferefrnotmis autbemda rginal setwaieat;p h a oiflro nsegt oanae b domsienga­l Descrfriobm3e s dp eciimnge onoscd o ndim­enVtI Ia,ns dh opratim resd ioanlV lIay n d tiaonnfrd o mafe wo thienpr oso croenrd iti(ouns.u Va;al lleysa)oc h absdeogmmiweninatatlh Unmounmtaetdel :nr oistae en. smamleld iolsaettoeaner aaclsh i dweih;to ut Mounmtaetdle: or viaaalb,oe uqtu arloluyn desde tnaeeae ra ccho xwai;ta sh i npgaloiefir n ter­ abto etnhd asn;ca llev fertsy h alLleonwg4.t3 h0 -antesnentaealae c2,h3 -µ3m3l onsgu;b marginal 91µ4m w;i d2t2h0 -µ4m1.2 set1ap ea:ai nrt eroinho eralIdby ,e tweeaecnh Dorsudme:r mme mbraDnoorusssa.el t1 a e:l ateral arseatani Idgo mnea atciahcb dom inal paoifmr i nusteet(e a aec2 µh m l,o nagb)od uotr ­segmAenntte.n enaac6ehs egmeannt1de3 d9 - satdoe ach (sacbaspoeenon ntse p eci me1n6)µ6.m l onsge;td ails trisbcuat3pi,peo e:nd :i ­ Dorpsoarlo efts h rteyep (eism):i nudtoer sacle 2l :+ c aam panpioforrseme, g mIeIn2It( : 1 microduwchtiaucplhpe esa,r b(iirliao)t c huevlrearry; IlVo1:nfl ge)s;h yo nsleVyt:1,a fl es hy fainmti nustiem pploere easc,sh l iglhatr lgyes re+t afl 1a gelsleatatane,Vd I w:i t3hfl es hy thaam ni crodwuicttohuno lefee a;c pho rtey pes etaane5do r6 fl agellatsee/tsaappeii;nc oasle inp aaiw ri;tpa hi orfps o rients w sou bma rgsienltaoaln 6g6,- µ9m6l onagn lda tfleargael l late antdw sou bmerdoiweasal,c l hiw nieta ph a oi fr seltoan g,8µ3m-C.1l 0y5p eoslhaib7er8la-dl8 5 poroenes a cabhd omianntadhl o rsaecgimc e nµtm,l onlga;b iwuimtfo hus re tSapei.r samcallels, buwti tohn loynp ea oifsr u bmargoinhn eaaldl;ey a cphe rit8r-eµ9mm we i dLee.g wse ldle vel­ anwdi t(hia ip ia)oi fsr m atlrli lpoocrunelesaa rro pelde;n gotfmh est athloer(gaµ scm i)cc:o xa antemrairogorinh n e aAdn.ap ll aetaeceshl on4­55-0t;r och+afe nmtue7rr2 -8t1i;b4 i3a- 52, gaatnedd i verlgeinn4gg6t,-h µ5 m5w, i dotfh tar3s1u-sc4 l0a1;w6 -s2e0td;ai ls tricbouxtaion: eapclha 2t5e-µ 2m8e; a cphl awtietav h e ryl ongw it7sh e tlaoen,g3 e0s-µt3m 3t; r ochwaintther aipcsaelpt ear h1a6p0s- µ2m7l 0o nagl;os noe 1l onsge tlae,n 3g0t-hµ3 m3fe; m uwri t2sh e ­ seatbao huatl fa-lwoaenyag ci hn nmearr g1i0n-,t atei,b2 ia ant da r4ss uest daies;mt aarlgo ifn 14µm l onagn,ads eotnae acshi odfel onagp i­each timbiicar owcittteahnr idsdiaiglai ;t ules casle teaa,c1 h8 -µ3m2l onAgn.o gefonli dtadli ssiwmiitolhnan era ,r rtohwoaetnrh beubrto th withp aaoi fsr e toanee a cahn terocloarnt eerrm,aulcl ho ntgheacrnl aw; clavwe rddyii sg­itules eac1h0- µ1m7l onagn,ad n otsheeotrnae achs imairol,n me ucbhr oatdheaornt hbeort,h 12 0 M.Willeiatal mA.sn : e gwe nofuCso ccidaeZOOSYSRTO.S SIVCoA1l 1. M ·�� ----� j�) r' -- o p Fi4gP..r ionoacmoecrciucssap nn..lu ,ss t -niynmsptfrhao rmC ozumMeelx,iL caob.ae sli snF iIge,.x cEe pd=to rsal trilopcourRle a,a=r n railn g.

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A new genus and new species of Coccidae from Central America (Homoptera: Coccinea) PDF

16 Pages·2002·1.5 MB·
by  Williams, M
#additional_collections #taxonomyarchive
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Preview A new genus and new species of Coccidae from Central America (Homoptera: Coccinea)

© ZoolIongsitciSattlu. tPee,t2 e0r0s2b urg, A ngeewnu ansd n eswp eocfCi oecsc friodmCa een Amtrearilc a (HomoCpotcecr ian:e a) M.WillCihaH.mo sd,g &s oEn. DMa.n zig Wi�lmlsiM,. H,o dgsCoh_&n. D,a nzEi.gM2,.0 0A2n .e gwe naunsnd e swp ecoifes Coccfr1odmCa een tArmaeln( cHao mopCtoecrcaiZo:no esay)s.t eRmoasts, i 1ic1ca(a 1): 111-126. An egwe noufss o sftc ale, PrWiiolnloiHcaoomdcsgc&,su D osan n ziisg ,i nttrooduced tatkweno e swp ecPia egasWv,ie l liams&,D aHnoazdnigPdga s moenr iWcainlulsi ams, Hodg&sD oann zfriogmC enAtmrearlTi hcaead .ufe lmta loefbs o atrhde e scraisab reed thles t�t-aairnn2 dn d-imnaslatenfa der m aanlpdeu poatf h lea tstpeercP ia emse;.i canus habse einn tpetrescdee vetriamilent s hU en itedo fAS mteartoienics ma p orted plants. T?en egwe niucsso nsitdobe erletodotn hgCe o cctirdiCaboeec cainnadki e,iy ps r o­ vidtesode pairfrtao tmoe t hgeern ientr hati rs i be. MikWiel liaDmesp,a rtofmE enntto moalnPodlgy a Pnatt hoAluobgyu,Ur nni ve3r0Is ity, FunchHeaslAsll ,a ba3m6a8 49-USA5.4 13, ChrHiosdgs onD,e partomfBe inotd ivaenrBdsi iotyl oSgyisctaelm ThaetNa itciso,n al Museuofm WalCeasrd,if Cf,F /3 NP,Wa leU.sK,. EvelMy.Dn aan zZiogo,l oIngsitciaRtluu stseAi,ca and eofm Syc ienUncievse,r sitetskaya nab1.S, t .Pet1e9r9s0bR3uu4rs,gs ia. Introduction matcilces fth alalnoacwll, ed fti stmianrcg ti,n s adprebsunstoe futds eDdo.r sduemrm:me mbra­ Anu ndescsrpiebcoeifsde o ssf cta (lHeo mon­ouosny ouandgu blutwt,i tanh a rrsocwl ero­ pteCroac:c iCnoecac:ifr doamCe e)n tArmaelr ictaisaatrioouannn dt emrairogorif an n apll ates; hsab eeinn terocnes petveerda l boytc hcea bseicoonmssi cnlge rmoetdiisoaeuldtl wyaa rtmd as­ quarantiinnteh Uesn eirtveidcT ehSsi(t sP.a tteusr.wi ittayhr; e olaltairongneesasm,rta rgbien­, ameriacnaadnn ao)tr heelras tpeedc( iP.ae gsa ­comisnmga lolrae brs meendti aDlolryss.ea tl a e vea)l,sfr oo mC entArmaelr iacrhaee, r e dset­oustpliyna onstdea pesrpiantrghs;re o ughout. scriabsne edaw n pdl aicnaen de gwe nPursi,o D­orpsoarloe fts h rteyep( eils)a: rp gree ope rcular nocoWciclulsi Haomdsg,s& o Dna nzig . poTrheeeascp,ho rheea vsicllye rwoittaibh sr eo da,d formsepre caipepset aobr ehs i gvhalryi bau btlbea sdailsi cand: i stlionncgti tbuadneidxn taeln d­ noc onstant cdoiuffbleefodr u entndocs eesp ai­nagn terfrioomar nlpayll a ttaoeb sop urto thorax; rattheve a riloouotsfms a te rial. (idio)rmsiaclr odeuacctwhui ltaefih ,n ien ner DepositBoMNrHiT:eh seN: a tuHriaslt odruyctule:t hfrreoquugaehnno(dtui atis ,im )a ll MuseLuomn,d UoKnU;,S NMU:SN atiMoun­alo pesni mplep-rpeosrteehn:rt o ugDhoorusta.l seuomfN atuHriaslt Woarsyh,i nDg.tC,o. nt,u budluacartn sdd o rtsuable racblseeAnsnt a.l U�A;Z IAZSo:o loIgnisctaAilct audteeom, fy p laatpepsr oxiqmuaatderilanytn meea,rr gins ScienSctPe.es t,e rsRbuusrsCgiS,aD ;AC :a lim­oroerl epsasr allel, possltiegrhtiloyr margins forniSat aDteep artomfAe gnrti cuSlatcurrace­o,n veeaxcp;hl awtiet3 sh t rsoentgda oer sally mentCoa,l ifoUrnSiAa., neaarp e(x n 1e aera cihn nmearr gnienaa,rp ex annde paors tmearrigoairnn wd)i ,ta sh h osrett a Prionogceonnc ..c us ventnreaaalrpl eyAx n.o -gefonliwdti ataplh a ir ofl onsge otnaa ntemrairogarin n3d p aiorfs Type species: Pargiaosvnpeno. .c occus shorteorn l asteetmaraera glpi lnutssh at near DiagnAodsuilst: Bfeomdaoylv aetlo.a l mosatp esxu;p pobratdrii nsgte ixntcetna,dn itnegr ior­ rounsdy,mm etorrai scyammle trpircoab la,bl lyty oa ntemrairogorif an n apll aAtneasrl.i ng ratflhaeitrnl iwfei,d eacsrtao bsdso msetni;gw -itfihv pea iorfss e taanewd i tahni ncomplete 11 2 M.Willeitaa lmA.sn : e gwe noufCs o ccidaZeOOSYSRTO.S SIVCoA1l 1. doubrlooewf p orMeasr.g mianr:g sientaaalle l i nH odgs(1o 9n)9 ,P4 r ionockoecyocsuua tst sligdhitslpyl oancteod doofrt swumoma ,i nM esolecMaonriruitmsow o hni,ic ithsc learly typ(eiass) :t osupti,ns oestleao ,n gpeosstt ericolrolsryee llyaL tiekMedes. o lecaPnriiuomn,o coc­ bustl igvhatrliiyans b hlaeap pei;hc aaollfef a chcuasp petaobr ers e strtiotc hNteee wdWo rl d. spiwnietm hi nudtiem pelaecswh;i tabh ro adP,rionoccaobnces c eupsa rfraotmoe tdhC eorc ­ we-ldleelvopbeads- aslocukseuta:rl alty h ecribnyir eplaccoiunp1gl2i e ntt h Ceo cckienyi abundaanndt a;s( eitiso)es teeaa ,cw hi tawh e llofH odg(s1o9nw9 i4t)th hfo el lowcionugp lets: develboapsseaodlc ksepta:rd siesltyr bieb­uted twesepni nsocsteSa tei.g msaptiiuncne dsi ffe1r2P­.reopeprocrupelrsae rsi enan b tr olaodn gitudinal entifraotmme adr gsipnianEley ess.mp aortg inalba.n adn tetroai noparll a tweistfe;hw s etparee sent aloanngo -gfoelndid;to artlsu able arbcsleeensat c;h Vendteerrmm:e mbraPnroeugse.dn iistca-lp ores postbearniodofd r i sc-npoeotxr teesn frdoimen agc h eacwhi t5h-l 7o cpurleis:oe npn rte gesneigt­alp osteriotrom esspoaifder aacmchel seo c.o.1x.3a menotn lmya,i nllayt etroaa ndt eerniodofa r n al Preopeprocraueblssa ewrni tnt;uh m ersoeutasal eo ng cleafnatdn palla Stpeisr.a dciuslca-wrpi o trhesa no-gfoelndid;to artlsu able prrcelseeesan cptho; s ­ main5ll oyc iunaln ia:r rboawne dx tenfrdoimn g terbiaonordfs piradciuslca-prpa oesratec psho ste- risopri rtaomc elseoa fed a cmhe soc.o.x...a.. .. eacshp irtaomc alregV ienn.tm riaclr ofrdeu­cts ..............S.u.a.r.e.zM.ia.em.le.lt.a . ... queinnat b romaadr gibnaanoldn ,h eaadn d 13M.argsientaoalf2e o r3t ypoensfie,n esleyt aonsde aroulnadb ibuupmto ssaibbslmeyen dti oanl ly thoet hsetrosus tpliyns otsieg;mc alteeiftacscw hi th abdomen. Pproeraaebnsst eeVnnetnn.att lru a­l spinnoectsl edairffleyr efrnotmmi aartgesidpn ianle s buldaurca tbss eVnetn.ts reatalan ela:ol bs eet ae ................P.r.i.o.n.og.ce.onnc...c. u s - Margsientaoalfoe nt ey psep,i nboeswneit,at b hl unt undifferefrnotmsi uabtmeadrs geitnaael ; with apesxt;i gcmlaeteftiasccw hi tthh rsetei gsmpaitniecs lopnrge gesneitotan3aes l e gmoennltbysue, ta ch cleadrilffye refrnotmmi aartgesidpn ianelea smc,he ­ segmweintatsh p abrasneod fs hosretta acer ossd iasnp idnies tilnocnttghleaylrn a tseprianl.e . s. . eacahb domsiengamlue snuta;l l2oy rm owriet h ............M.e.s.o.l.e.cM.ao.nr.ir.ui.ms on setnaeeaa rrt icoufel aaccthoi xoiann; t era ntennal settawepo:a iwrists;he veral small PsruibomnaorcsgohicanwcraiueltssMe h s olecatnh ieum setbaeet wseteing maarteiwacis t;1h p ai r foolflowcihnagr a(cipt)re erosp:e procrqueulsia tre hypopysgeitaalc ; othearn qdus ie ttleaaer igsnaemb ,ar lolla odn,g itbuadnaidnn taeltr oi or sparAsnet.e n6n-asee gmsecnatwpeiedt3 ;sh e ­anpalla (tiediso);r sseatslal ei gshptiln(yoi siei;) tapee;d i2cs eelt+:ac ea mpanpioforrseme; g ­dorswuimta hr eola(tiipvor)ne sg;ed niistca­l menItI 3Is :e tIaVe1:;fl essheyt oanelV y:I, porreess ttrmiooc stpteo ds taebrdioomrsi enga­l flesshey+t as1 e tsoestaean V,dI 3:fl essheyt amee n(tvp;)r egedniistca-wlpi omtrahei ssn elvy en + 5o r6 s teose/sspeitnMaooesu.et hpuasrut­lso c(uvlaiib);s eonfvc een ttruablud luacartn sd, ally apslyamcmeed trliacbaigluelmny e;r a(lvlityii)bios-etgamresnuatsba stoeirnpo ton o rly twiswtiet4dph ,a isrmsas lelt Sapei.r naocrl­edse veloped. maLle.g uss uaqluliwyte edl elv elbouhptie gdh lyEtymolPorgyi:o noicfosor cmcfreuodsm prionos varisaobmleet,wi imtehsdo iusttt irnoccth an(tGmor)­.m ean'isnagrw e'fe,rr tiotn hgme a rngail femuarn tdi biosteagrmseanlts aettciaooexn:a; s etael wohorkia ctlhhi etkrhet e e eotnah c hna­i 5-t6r;o ch1al notnsegeor tn av entmraarlga innd saawn,-d c ocacftuetsrhfa emi lnya m e. onseh osretot nad ormsaarlg feimnu;2r ;t i b2i-a Prionoccuorcrcceuonsnt tl2ays i pnescP. i es, 4a ntda r2s-u4ts;a rcsaamlp anpioforraemb ­agaavneP.d a mericbaontduhes s,c raisnb eewd senctl;aw wi tadh i stinctta rddseianglti itcublleeel;. so w extenwdeiblnelgy otniodpfc lacwl;ad wi gitules subeiqlnue anltg tota hr dsiagli atnuedlx etse nPdr­ionoacgoacsvcpenu.. s inwge lble yotniodpf c laown;ce l adwi gitule generbarlolatydh eaornt hcelrad wi gitule.Holotyp<;>e,,M exiScaoLn,u iPso teoxsl ieo,aAf fg a ve CommePnrti.o noicosec acsruieslc yo gnifalscea(dtA ag avaceae)R,. HL9o..d VgId.o1n9, 8 0(,U SNM: Lare1d3o9 6y1o)u:un ngs cleardoutfelimtsa elodena by( it)hl ei onfel a rpgree opeprocruaelnsat re ­ sliwdiet5a h <;>d ,h olofetmyapclele e amralryko enmd a p ritootr h aen apll a(tietiseh,)r estroiftc htei oonrn i ghtl-ahbaenld. pregedniistca-ltp oto hrmeeo ssp to stperrei­orP artay pMeesx.i acsfoo r,h olot4sy cplee,r aod9t ised gensietgaml(e intithi;ae) b seonftc ueb udluacrto sn s liwdieth ho lot<;>y p(eU SNSMl)id.de ep osi inted onb otthhd eo rsaunmvd e nt(eirtv;h)r e o b usUtSNM. Othmeart erMieaxlii.cn ot,e racteB prtoewdn sville, spinmoasreg sineatelaa eco,hf w hiacphp ears offl eoaffA gasvter iBcutragR,ee sis(s UtS,N 6M8 97), toh avase l igdhitmlpayln esddp insuulrafatce1e /,a1 d <;> i ng oocdo ndiStliidodenep. o sZtIeAdS .i n an(dv t)h peo sseosfts witooy np oefms a rg inal setsapei:ns oestaean efld a gelsleattaee . Aduflte ma(lFei1 g().D escfrroim6bs epde ci­ DiscusTshigeoe nnP.ur si onoacpopcecaumrses n 2si ,ne xcelcloenndti4 tm ioorhnee, a vily tob elotnotg ht er iCboec cFianlial sed ne fin esdcleraontddio sressdup mbl uisttt qiuliglto eo d). byH odg(s1o9n9 I4nt) h.ke e gyi vteogn e neraU nmoumnatteednr oistae le:n . ZOOSYRSOTS.S IVoClA1. 1 M.Willeitaa lmA.sn : e gwe noufCs o ccidae 11 3 K m .\ [ f>-..; '. �-,- p··. ._, ·\�V�,�- C - �u '/ ;/ � . .. . ' --<' , .J ,•-•.�...- •----.... -.,, ....,........•,_ 1......1,1,.--. ·'l·1·• ··.A .-,.-1 I I N ® G Fi1gP..r ionoacgoacsvcpeun. s.a ,d ufelmta lIent. ha insid nF i2g-sA6 = :d orsseatlBa= ed ;o rssiamlpp olreCe ;= dormsiaclr odDu =c tpurleeo;pp eorrGce =u;a l naparll a Ht= ea sn;o -gfoelndKi; =t m aalr gsienta(alibe :u lb ous spisneoitsiale:;a ncesoplianstoeesti eaifl ;ia :g esleltaLat a)e=n; t eMn =ns ap;i radciuslca-Nrp p o=rr eeg;ed niistca­l poPr ev=;e nmtircarlo Qd um=ce tt;a thloeorgrca lc aiwc. 114 M.Willeitaa lmA.sn : e gwe noufsC occidZaOeOSYSRT.O SSIVoClA.1 1 Mounmtaetde rbioadslyh: o wvienrsgyl i ghltenogft"hl ofinnsgee" t soesotenta e rmsienga­l sigonfas s ymm(emtorosybt v iionpu oss itimoenna tb o3u8µt m M.o uthupsauratslshl oyw ing ofm outhplaerntg1st.)h5; 7 -m2m.,w6 i3d tho nlsyl isgihgotnfa s s ymmeltenrogyfct; lh y pe­ 1.3-m2m. .0 olabsrhail1e 5l3d- µ1m6l;0a bisuomm etimes Dorsduomr:ss eatlsa pei neoisteth,ae pre ritnwgi sStpeidr.na ocrmlaeelsa: ca hn tepreiroirt reme toas hapropi( nBtr ownmsavtielorlrire aa lt )h e3r3-µ4m2e, a cpho stpeerriiotr3r 8e-mµ4em3 . bluanntpd a rall(eSla-Lnsu iiPdsoe tdmo astie ­Legpse rhraaptssh mearwl ilt,h aonudyti stinct riaela)c7;h- µ9m l onDgo.r spaolr( eisp):r e­segmenbteattwiteorenon c haannfedtm euror r opercpuolrearea scb,hr obaads dailss ucp porbte­twteiebanin tada rlsounsg;ce osxstae let aac h inagr atflhaectrl opsoerdle a;r( geea5 c-hµ9 m 33-µ4m3l onlgo;nt gr ochsaenettaaec arhb out widien)a d: i stinct blaonondfg3 i1t-u6d32i 6n-µa3ml4l onlge;n g(tµhmos)fm etathloerga:cic por(eisdi;o) r sal miacnr(doi daisu imc)at lucllo ex,8a 3 -96; t+rf oecm1hu0ar7n -tt1ei1rb6 i;a + opesni mples-ipmoiirsnlei a,tzr moe i crod ucttaurl1se3u.2s- c1l4a02w;1 -24. Anapll aetaec1sh3 4-µ1m4c,5o mbiwnieddt hE tymolTohgysi.ps e cinisae msae fdtt ehhreo st 15158-µ0m e;a cphl awtiet3 sh e teaaec2 h1 -32plagnetnA ugsa (vAeg ava ceae). µml onAgn.o -gefonlidlt:oa nslge o tnaa nteriCoormmeP.n atg.a ivsve e rsyi mitlota hforel ­ margeianca hb o4u0t- µ4m5A. n arli nwgi:t3 h lowisnpge cbiudetis ff (ec rhsa ractoenPr -states paiorfls o nsge taanetd w roa ther shoarmteerri.icnab nraa ckienht asv)i( nidg)i s tinct Margimna:r gsienta(alise )p: i nsoestleaa en ­grooufsp psi intne hsse t igcmlaset( fingtcor oups ceollaotnegp,eo sstt ebruistol rilgvyha trliya bolfse t igmati(cio ins)lop ynit eny eopsfe)s ,p inose isnh aipsnei :n rgolawer omuanrdg hiingv,ha lryi m­argisneatla an(ttehsrepi ionrso estaear ee ablfree qinu elnecny1g;1t -hµ2 m0l onagn;(d i im)a ibnullyb woiurtsoh u nadpeidwc heesr,te haes setsoesteea aec1 h0 -µ1m7l ondgi;s troi fbumtoispoton s taerrleia onrc eaonl(dai tieic))l, a twhe setoanBe r ownsmvaitlelr2ei0s apli:n+ o s8e d igivteurldyei sff eirnes nit(zo eft ewni tohn e setboestew eeyeens 1p5o-ts1sp8,i n+o s2e s ebtroosaetdheatrnho et hbeunrte vaesmr a rkseod ly betweeyeensa pnoadtn tesrtiiogrm at1i6c- acsol nP.e a ftg,a vOet)h.e rtwhietssweeso p eacriee s 17s pin+o ss1ee tboestew seteing mcalte ifcct lseacrlloysr eellya ted. late5r7a-ls5lp9yi ,n+ o ss6ee tboestew peoe­sn tersitoirg mcalteaifnctad sn acll eafnt7d,- 9 Prionocaomcecruiscs apnn. .u s spinoonslaeyl omnagr goifan n calle Sfta;Ln u is Potomsait er1i2a-ls2:p0 i n+o 4s-es8 te ose Holotayd9p ,Me e:x iCcooz,u meexCl o,c coslpo.b a anterbieotrweleyyee ns p7o-t1ss0p, i n+o1 s- e2 (Polygon2a0c.VeIaIeIM)..,1L W9.i8 l8l,(i UaSm NsM , setose betwaenaednn t eesrytieiosgrp m oatt ic AL-028S-l8id8de)ep. o siinUt SeNdM . cle5ft-,1s 3p in+o sslee tboestew seteing matPiacratMyepxeisac.sofo ,rh olot2y9sp lei,wd ie8ts h clelfattse 2r3a-ls4lp5yi ,n+ o ss6ee t boestew eeynou andg9 ( 2o fd orsounmls1y 3)o ,la dd 9 ( 3p oor), postsetriigomcral teaiftncsad n calle aftn,1d - 62n9d, 22 nc!d, 1 4c raw(l8pe oros+r )eIm br(yDoe.­ posit2oa rd9i B eMsN:H 2,a d9 Z IAISa ,d9 NMW; spin+o0 s-se1e taolsoemn agr goifan n calle ft. reisnUt S NM.) Stigmsaptiiuncne dsi fferfreonmtm iaartgeidn al Othmeart eMreixailCc.uo a:m aMvoarceael xYo usc,c a lancesopliabntueefots r miand gi stgirnocutps ,p3 . .,lI II.E1.S9 h9u6v,a (cZhIi7An3Sa- 926/a)5d9, i n with9 -1p0ec rl (eftB rownsvilaln4ed- 9m agotoecdor nidaialtn)isd oe nv eurnamlo uantd9e dl+ a rvae; (SaLnu is PotoAsniac llm aeabftto e1uri/ta1 l0)tC.hh iav,Oe alxaocnaC ,o ccoslpo".bC aa me(rPoo"l ygo­ bodlye ngEtyhe.s epaoct1hs2 -µ1m6w id e. nac)e1,a9 e2 G6F,F. e r(rCiSsD- AX,- 64-remount-.X.1964 RJW3)/,a3 d9 ( +embirngyo ooc)do ndiitnitoenr;c epted Venteprr:e genitalw idt1ih1s -co1-n 4p ores: Browns(vUiSlAel)xAe, n thurium (cAlraarcienaeer)i,u m eitheSrp isriaddceiu.sl ca-rwp iot1rh8e -si2: n5 9.VIIIB.u1r9g(7eU7sS,sN 7 M5 915/)a4,d9 e+m bryos; eacahn tedriisocr-b paonardne 1d 8 -i3ne7 a chi nterBcreopwtnesdv ilelxcA nt(hUuShrAoi)ou,km e ri, postbearniVedon.rt rmailc roadsiu ngc etnse r2i6c.V.G.1 B9u7r8g,(e UsSsN8 M3 613/)a2,d9 ( poor)+2 diagnVeonstirssae.lt a aneal:lo bsee tgaeen er2­nd9 ;i ntercBerpotwends( vUiSlAel)xeA, n thurium robus\t0u.mV,I IBIu.r1g9Vae7ns7vs,a lke(nUbSuNrMg ally undifrffoemsr uebnmtairasgteietbndaua etl 75981)/a,2d 9 ,2 2 nd9 ;V eracrduizvi. n,t ercepted occasiuoptn o2a 2lµ lmly o nlgo:n gpersetg eni­ Browns(vUiSlAel)xAe, n thusrp2i.3u,.m lll.1973, Stevens .asle teaaeca hb o7u4t- 1µ0ml1 o nogt;hs eert aeeta l(.U SN3M5 725/)a3,d9 ,22 nc!d, 2 2 n9d p+u pa; presmeendti aalclryao sbsd omsiengamle nitnst:e rcaetMp itaemodinP , h ilodesnp(d.Ar roanc eae), VI1I:5o r1 6V,I8 -1V1:9, - 11,e aIcV6h-- I8I;2:7 .11A..1SM9.i5 l1(l,Us S N2M0,0 561,. 812/4a3)d9 , + 2nc!d. G uatemianltae:r LcoeAspn tgeed(l UeSsoA n) , setaases ocwiiatethae cdch o xmae:t ac2o,x a leoaffu nknopwlna2 n0t.,l .K1.S9 h8i6m,(a UdSaN M, mesoc2oa xnpadr oc4oa,x lsalh oirntt;e ranten­ 05858764--5624/a14d9) ;,i nteracNte epOwtr elde ans, nasle ttawepo:a ilrosn,ga ebso2tu5 µt m l on go;nC occoulnoifbe1ar7 Va.I, II9 I 8.H2 .,F inge(rmUaSnNM wit6h-s 8m aslulb marsgeit(naeaeal3 c -hµ4 m 014852/0a9)d9 : ;i nteracHteo pbtoekoden An n,t hurium lonbge)t wseteing maarteoiatcsh s;ee rts amea lslal1vl i.iV,I IL..S1 i9r4(o9Ut,So N 1M3 6520;- 217/13), and qsupiatrAesn et.e ntnoatea:l 1l7e5n-agd9t ,5h2 n9d.H onduirnatse:r Mcieapmetixee, pd i phytic cact2u5s.,V IlR.E1.C9 o7t5(t,Ue SrN 1M1 7430/ad)5 , 20µ8m t;e rmsienqtauali l toenl gne,g t3h3µ m ; 9,22 nc!d 2+n 9d. ZOOSYRSOTS.S IVCoA1l 1. M.Willieatam lsA. n :e gwe nofu Cso ccidae 11 5 Descrfriobm3e 0sd p ecimmoesnitsnlfa, yi rOaxaicnMa e xihcaomd o satn)(d;i ais )e t ose tgoo ocdo ndiNtoittoehtn ah.tii assh igvhalryi s­e1t6a-,µ4 m0l o nwgi:4t -hb9 e tweeyeens pots; abslpee c-iicetos u rledp raec soemnptol fe x1 2-b etweeaecenhy esapnaodnt t esrtiiogrm atic spebcuinteoc s o nscthaanrtca ocutbleefodr u n dcle1ftl ;a tebreatlwsletyei ng cmlaetafint6cd-s tsoe patrhaeatmnei d it hs e arses utmhetadht i s7a loenagcs hio dfea bdomaenwndi, 0t h-o 2n variraetpiroehnso esntt-sio nerdn uvcierdo nemaelcnah­t mearraoglfia n nc alle Sftti.g cmlaetfitcs tally-(iobnrod tudhci)eff de r ences.sligshctlleyr aoltoimnsager dgs itni:g matic spiunnedsi fferfreonmmt airagtbieundla blo us Adufelmta le( Fi2g,3s ;Ta bl e). spiEnyeess.ap obou1tt1 -µ1m3w ide. VenteDre.rm me mbraPnroeugse.dn iistca­l Unmoumnatteedrd iraislep:de ciomve anlsp, orweis4t:-h 2 o0ne itshie(dr1se p echiam den ratflhatetsor l igchotnlvdyeo xr,ss ucml er oats iisnedgdilsec -opnso ergem VeIn)St.p ir acular anbdr owwinat t hh ciont tcoonveyex rc meep­t disc-wpiot9rh-e 3is4ne: a cahn tebrainaodn r d diallelcayof;v eirnfle udffw yh ictoet twoan xy,1 33-9i ena pcohs tbearnVideo.nr t mriaclr oducts whiicashs sutmohe advb ee esne crbeytt h eeda si ng enedriiacg nVeonstirssae.lt l aoen:g est adufelmta l es. prgee nsietteaaaleac bho 7u9t-3 µ1 m3l onlgo,n g Mountmeadt ebroidsayol m:e toivm(aeFls i gs.eatlars aor perleyso ensn etg mIeVon;tt h seer­ 2b)ugte nesrhaolwlsiyln igtg och otn sidetraapebr leesmeendtia aclrlaoybs dso msiengamle nts asymmweittorhnys e,i sdieg nifilcoanngte(lrtyo tVaI:l1I s1)-V:1 I98 ,- 1V5:4, - 1I8V5,,- 14, thoatnha enwrdi atnhla olob noe n sei sdter onIgIl5Iy-:1a n4Id I2 :-O 1s; e taases ocwiiaettahec dh con(vFei3xg) a.;n callae bfot1u /t9 -1b/oldOyctohx mae:t a1c-omx5ea,s oc2o-ax6na pd r ocoxa lenlgetnh1g;.t 3h-m 3m.,w1 i d0t.h8 5m-m2..51 7-,u suaallsllhy o orctc,a siloonnaglelsyt Dorsudmo:r sseattlaa ep etraosi hnagpr opi ntpr:ocsoexutapta 1o 8 µ ml onega;cp hr othoracic perhmaopscsto mmmoend ioanla lbyd o mesne;gmweinsttoh m see tmaeed iianltleyr;a ntennal ea7c-hµ1 m0l onDgo.rp soarl(e ips)r: e op ersceutl2apa ear:i( rrsa -r5eG,lu ya teamLtao lAsan ­ porfaeisrl layr egaec5,h- 1µ0mw idwei,ta h gelleosn)g,2e 5s-µt6m 5l onwgi;t6 h- 1s3m all brobaads als-udpipsocarc toinnvcgel xo sesdubmarsgeit(nalaeel n3 g-µt8mh l) a tebrea­lly poriaend : i sltoinngcitbt aunod9dfi- np5ao9lr est;wesetni gamraetoaitschs; ee rts ameab lufrlte ­ (iasi m)amlilc rodpuecrthmuaolpscesto: mm onquepnatr,t iicanub lraomraaldry g biannAadnl.­ nemaarr gainundp m argoifan nsca lle afntd,t ennIaIeIs:re dg mreanrtwe iltayph s eudo-ar­ (iiais )maolples ni mpleA-npapollr aept.oe ss­:t iculeaatc1ih6o 0n-µ;2m 7l 4o nlge;no gftt ehr ­ termiaorrgo iftnewsni tadh i stiinndcetn tamtiinsoaenl2t; 8a- µ5m2l; e nogflt ohns ge tsoestea len1g1t1h- µ1m6c,2o mbiwnied1dt4 h0 -2o1n0t ermsiengam3le3 n-µt9m 1M. o uthupsaur­ts µma;n apll asteeta alselt rosnegtlaoyns dea ldliys taisnycmtmleyrt artichcleaortlso o,en re subeiqnlu eanleg atch2h,8 -µ5m0l ondgo;r saplrocloexnaog;fct lhy peoslhai1be2rl5ad-l 1 90 surfafrceeq uewnittplhaya l ree (aasr eolaµtmiS.op nisr)eaacaclnhet se:pr eiroir2t 8r-eµ4mm7e, neaanrt emrairogArin no.- gfoelndil:to anslge ­eacpho stpeerriiot3rr3 e-µm6me5L. e gpsr:e s­ taoena ntemrairoegrai 2cn2h - µ5ml8 o nAgn.a lencoeft ibios-etgamresnavtlaa rtiiuaosbnul ­e, riwnig2t o hr3 p aiorfls o ntgh,is cektp ale2u s alwliyst lhi sgihgotnfa ss m aplsle udo-segmen­ o3rp aiortfsh insnheosrre,tt eaarne ta;ula b beo uttatoinso mna llleegbssutw,t i tadh i stsiengc­t 1.5-l2e.n0ogxfat nhpa lla tes. mentaotnli aorng leesgltse ;n g(tµhmos)f MargiMna.r gsienta(alise )p: i nsoestoea f em etathloercgao:cx7 ia4c - 1t3r3o;c +h afen­ter twtoy paenst:e rbiulobrwoliuyatms ho roerl e ssmu9r7 -2t2i8+b; it aar1s0u7s- c2l5a12w;2 - rounadpeedax n wdi tahd impslufaercd e 35µ ml;o ngceosxstae let aac5 h1- µ1m1l 2o ng; apiceaalcalhby o,8u -t1µ 7ml onagtp; o st erlioonrgtersotc hsaenettaaec arhb o4u2t- µ9m5 enodaf b dommaerng,si entmaaolerl ea nce olloantcgel;,ad wi giatlumlsoeissmt io lns amra llest butthv iasr isaobmslepe e,c ihmaevnsisln ig g hltelgdysi,s tdiinscstiolmnlyi a lrlageresg slsti, g htly poinstpeiednv eeosnn h eaeda;ca hb o1u3t- 2b5roatdhetaran r dsiaglia tnuedlx etse nwdeilnlg µml onagl oanngca lle iftna;s intgodl oeu blbeeyotniodpfc law. roawr omuanrdgw iin1t;5h - b5e0t weeyeens poDtisscutshsmiiasot ne:wr aiesax lt revmaerliy­ onh ea6d-;4b 3e tweeaecenhy esapnoadtn tea­blTehm.ea ivna riwaatisin (o:ibn )o dsyh a pe: risotri gmcalte1ift1c;- l4a3t ebreatlwl eyte hnfraotmM oreMleoxsi,(c Foi2 g)w.,a bsa r ely stigcmlaet(ftirscom wosro erl edsosu wbhle en asymmewthreirmceoaaslost t hmeart e(er .i ga.l spianbeusn d3a6n-t1a)3l,8oe nagcs hio dfae b ­fromG uateimnatlear,ic nMe ipat(meFidi3 g) .) dom(eo nfw hipcohs tIe0 r-i2ao0rlr ea nce oslhaotwece)odn sidaesryambml(eietni ru)ym ;bo efr anwdi 1t-h1o 2ne itshieodrfae n calle aftl,w aymsargsipnianlso estteah ee:s em owrteehr ae n lonagnemdso tsp to in(tmeadtfr eormHi oanld ut­wiacaseb unodnma antte frroimMa olr ealn ods raansdo fPhfi lodefrnodroMmne xihcafeodw e stOaxaMceax,i acsoo nt hmea tewriittahhl e spinmoasregs ientatalhe ofr,so emM orealn odsfe wemsatr gsientafraloe mH onduarntadhs a t r e : t y n - r ;: P. ble. Ranges in data for 8 collections of adult female americanus from various localities. Where: No. = number of specimens studied; column 1 = length of anal plates (µm); 2 = no. of eopercular pores; 3 = no. of marginal spines along anal cleft; 4 = no. of marginal spines laterally between stigmatic clefts; 5 = no. of marginal spines anteriorly between eyespots; 6 = no. of length of coxa (µm); 10 = length oftrochaegenital disc-pores; 7 = no. of spiracular disc-pores between anterior spiracles and margin; 8 = width of anterior spiracular peritreme (µm); 9 = mur (µm); 11 = length of tibia+ tarsus (µm); 12 � claw digitules: where O indicates digitules similar ranging to 5 very dissimilar; 13 = length of antennae (µm); 14 = degree of asymmetere i indicates barely asymmetrical, only mouthparts displaced; 2: intermediate; 3: margin of body showing high degree of asymmetry, particularly posteriorly (as in Fig. 3); 15 = tibio� rsal articulation: where I indicates pseudo-segmentation poor, sometimes absent ranging to 5 where complete segmentation present.� i;;· 5 JO 149 12 Locality 1 l 1I 13 5 3 8 No. 2 46 7 "' ~ 215-240 165-199atemala: 145-162 I I 07-128 343 26-42 6-1241-47 145-162 13-3123-34 12-18 15-37 f.:'-boken 4 150-190 atemala: 166-183 3132-1661-3 95-112 31-37 4128-145 24-55 2 12-269-22 1-5-13 21-35 ;,: w Orleans OQ 186-228 169-2071-3 4 145-153 atemala:107-132 45-9 36-44 2 139-145 26-49 24-3114-24 14-18 18-33 s Angeles �I 211-248 136-166nduras: 2 5 5 161-186140-91-120 153 9-18 3-836-42 17-28 2711-15-21 5-10gami 835-236 248-270 xico: Oaxaca 157-173 120-132 1-2 3 43 7-10145-150 52-55 37-1446-53 15-33 30-42 160-178 xico: Miami 3 108 161-166 3-52 74-83 16-21 2 33-42 134-143 12-30 11-164-1015-18 l xico: 3 256-273 221-250l 11-124 157-174541-47 44-59 434 8-1426-6-1711-18 144-161 30-50 relos 0 2-3 xico: 169-2034 186-228 132-145 86-116 36-43 5-8 16-31 18-27 124-133 411-20 2 13-24 21-37 (/) zumel ;l nge 108-2503 31-53 160-273 74-132 9-59 1-1411-43132-1903.5 124-162 4-209-3450 15- � arrfeha uo ue uo oi e e eo eo a Tpp+wt GH GN GL HM M M MM MC R ZOOSYSRTO.S SIVoClA1. 1 M.Willieaatml sA. n :e gwe noufCs o ccidae 11 7 C .•'.• ·, . ,•,. 0B 0 . d 0 K. . II @ N G H Fi2gP..r ionoacmoecrciucssap nn..au ,ds u felmta lfroem M oreMleoxsi,Nc oott.eh t ahtfoi rsmi asl msoysmtm et rical, apafrrotmt hpeo siotfti homeno uthpLaarbtaessli. snF igIe.,x cRe= pv te ntsre atla . 118 M. Wilelatil aA.nm :es gw e nofu Cso cci•daZeO OSYRSOTS.S IVoClA1. 1 iii L D �-,--B, \ 0--- ,' II ' ' \ I J 0 : 0 ' \f : § f , N G @ Fi3g.P. r ionoacmoecrciuscspan .n.a u,ds ufe lmta frloem C ozumMeelx,iN coott.eh t ahtfoi rsmi sst roansgylmy­ metrLiacbaaelsil. nFs i Ige,.x cJe pm=ta rgin. ZOOSYRSOTS.S IVCoAl . 1M1. Wileltai laA.mn :se gwe noufsC occidae I 19 interactMe ipatmfreiod m M exi(ciosi;ii oz) fe latmearraglei anc2,h3 -µ3m0l onsgu;p po rting thlee gasna dn tentnhaleeo :n gleesg(tsfr o mbapro odrelvye lAonparelid wn.ig t6 sh e taaned MexiOcaox:a Mcoar,e Cloozsu,m Geula;t ema9lp ao,ra ensta;ul bseh o rt. interacLteo pAstn egde wleersme)o rteh tawni ceM argimna:r gsientaoalfte w toy p(eifls)a: g ­ thlee ngotfth h seh orlteeg(ssMt e xiocff o,e llatee ascuehptt ao3e 0µ, m l onagn,(d i i) Philodetnhadenr toenns)nh;ao ewt ehdse a mes pinsoest(eae ea wcihts hm adlilm palneds/ or trenwdistth,hl eo ngmeosrtteh aIn/ 3lrodn gesrpisno unal peihcaalolf)f2 t ypreosu:n bdueld­, thtahnse h or(tievts)hpt er; e soefsn ecgem enbtoau­ssp i(ne easc8µ h m l onagne)dl onsgpaatteu ­ tiboent wteiebanin tada rtshuwisa:sws e ldle ­lastpei (ne easc1 h1 -µ1m3l ondgi)s;t r ibution: veloopnte hdle o ngleesbgtusi t n disotrai bn­catr ouhnedab de tweeyeens p4bo utlsb:so pui sn es seonntt hsheo rtaen(sdvt t);hs eh aopfte hc el aw+ 8fl agelsleatotaneee a;c shi bdeet weeyeen­ digitduilsetsid:ni cffteolrnyet nhlteo ngleesgtss poatnsad n tesrtiiogrma arte3iab c:u lbous bumto roerl essism iolnta hsreh orTtaebs1lt e.s pi+n efls2a gelsleattlaeae t;e breatlwleye n givseosmo eft hdea fotra8 l ootfts h miast erisatli.g mat3ib cu labrsoepuais+sn :efl sIa gellate Iitcs l etahrta hte irmseu cohv erilnma opso tf setaan;pd o stetropi oosrt esrtiiogrma arteiac: thedsaebt uant os tructduirffaelrc eonucble eds wit2hb ulbsopuisan netse rtihoebrselecy o,m ­ founwdh iccohn stantltyh esImena.p d adriai­tnsegpd a tpuolsatteewr iit5ooh rr6 lo yfl, a t+t e7r tionnod, i ffere(nocttehhsea rin ns izwee)r efl agesleltaSatteei. g msaptiuinnced si fferentiated founidnt hoet hleirsfe t asgteusd aineddd­e fromm arginal spicnleesaft;bs s esntti.gmatic scribbeelIdoht wa .ts h erbeefoeranec cetphtaetdE yesopnmo atr gwiind;1t 1h- µ1m3. thmiussr te preass iennhgtgil helv ya riable Vesnptede­err:mme mbraPnroeuamsnu.al lt ilocu­ cietshd,ei fferences bbyhe oisntg- pcllaaadurnis tse cd- apbosreSenpsti .r adciuslca-erpa ocrhe s ore colofagcitcoarls . wit2h-( 5m ai3n)ll oyc iunalb ia n1pd o rwei de Pa.meriicesaa nssuieslp ya rfraobmlP.ae g avbeetwsepeinr aancmdla er gwiint3,dh i sc- pores in( il)a ckaid nigs tgirnocoutfsp p iinnee asc hi ne acahn tebrainaodnr 3d o r4 i ne acpho ste­ stigcmlaet(fiihtcia; )v idniffge renmta-rs­hraipboeardn V den.t rmailc ropdouscstriseb sltyr icted ginsapli annetse rainopdro lsyt erain(odir iliyt);o1 p ostteoer aicsohcr a 1pb ee,t weeaecpnhr o­ inha vitnhcgel adwi gimtuuclmheo srs ei m ilaanrmde socaonx4da- s,5u bmariganib ndaolmliy­ thoannP. aga ve. nasle gmeInIVt-IsP. r eantpeonranebassl e nt. Ventrtaulb udluacartb ss eVenntt.rs aelta anea:l First-innysmtpa(hrF i4g). lobsee tnaoedt i fferefrnotmis autbemda rginal setwaieat;p h a oiflro nsegt oanae b domsienga­l Descrfriobm3e s dp eciimnge onoscd o ndim­enVtI Ia,ns dh opratim resd ioanlV lIay n d tiaonnfrd o mafe wo thienpr oso croenrd iti(ouns.u Va;al lleysa)oc h absdeogmmiweninatatlh Unmounmtaetdel :nr oistae en. smamleld iolsaettoeaner aaclsh i dweih;to ut Mounmtaetdle: or viaaalb,oe uqtu arloluyn desde tnaeeae ra ccho xwai;ta sh i npgaloiefir n ter­ abto etnhd asn;ca llev fertsy h alLleonwg4.t3 h0 -antesnentaealae c2,h3 -µ3m3l onsgu;b marginal 91µ4m w;i d2t2h0 -µ4m1.2 set1ap ea:ai nrt eroinho eralIdby ,e tweeaecnh Dorsudme:r mme mbraDnoorusssa.el t1 a e:l ateral arseatani Idgo mnea atciahcb dom inal paoifmr i nusteet(e a aec2 µh m l,o nagb)od uotr ­segmAenntte.n enaac6ehs egmeannt1de3 d9 - satdoe ach (sacbaspoeenon ntse p eci me1n6)µ6.m l onsge;td ails trisbcuat3pi,peo e:nd :i ­ Dorpsoarlo efts h rteyep (eism):i nudtoer sacle 2l :+ c aam panpioforrseme, g mIeIn2It( : 1 microduwchtiaucplhpe esa,r b(iirliao)t c huevlrearry; IlVo1:nfl ge)s;h yo nsleVyt:1,a fl es hy fainmti nustiem pploere easc,sh l iglhatr lgyes re+t afl 1a gelsleatatane,Vd I w:i t3hfl es hy thaam ni crodwuicttohuno lefee a;c pho rtey pes etaane5do r6 fl agellatsee/tsaappeii;nc oasle inp aaiw ri;tpa hi orfps o rients w sou bma rgsienltaoaln 6g6,- µ9m6l onagn lda tfleargael l late antdw sou bmerdoiweasal,c l hiw nieta ph a oi fr seltoan g,8µ3m-C.1l 0y5p eoslhaib7er8la-dl8 5 poroenes a cabhd omianntadhl o rsaecgimc e nµtm,l onlga;b iwuimtfo hus re tSapei.r samcallels, buwti tohn loynp ea oifsr u bmargoinhn eaaldl;ey a cphe rit8r-eµ9mm we i dLee.g wse ldle vel­ anwdi t(hia ip ia)oi fsr m atlrli lpoocrunelesaa rro pelde;n gotfmh est athloer(gaµ scm i)cc:o xa antemrairogorinh n e aAdn.ap ll aetaeceshl on4­55-0t;r och+afe nmtue7rr2 -8t1i;b4 i3a- 52, gaatnedd i verlgeinn4gg6t,-h µ5 m5w, i dotfh tar3s1u-sc4 l0a1;w6 -s2e0td;ai ls tricbouxtaion: eapclha 2t5e-µ 2m8e; a cphl awtietav h e ryl ongw it7sh e tlaoen,g3 e0s-µt3m 3t; r ochwaintther aipcsaelpt ear h1a6p0s- µ2m7l 0o nagl;os noe 1l onsge tlae,n 3g0t-hµ3 m3fe; m uwri t2sh e ­ seatbao huatl fa-lwoaenyag ci hn nmearr g1i0n-,t atei,b2 ia ant da r4ss uest daies;mt aarlgo ifn 14µm l onagn,ads eotnae acshi odfel onagp i­each timbiicar owcittteahnr idsdiaiglai ;t ules casle teaa,c1 h8 -µ3m2l onAgn.o gefonli dtadli ssiwmiitolhnan era ,r rtohwoaetnrh beubrto th withp aaoi fsr e toanee a cahn terocloarnt eerrm,aulcl ho ntgheacrnl aw; clavwe rddyii sg­itules eac1h0- µ1m7l onagn,ad n otsheeotrnae achs imairol,n me ucbhr oatdheaornt hbeort,h 12 0 M.Willeiatal mA.sn : e gwe nofuCso ccidaeZOOSYSRTO.S SIVCoA1l 1. M ·�� ----� j�) r' -- o p Fi4gP..r ionoacmoecrciucssap nn..lu ,ss t -niynmsptfrhao rmC ozumMeelx,iL caob.ae sli snF iIge,.x cEe pd=to rsal trilopcourRle a,a=r n railn g.

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